Kevin Weitemier,^1,* Shannon C. K. Straub,^1,* Richard C. Cronn,² Mark Fishbein,³ Roswitha Schmickl,⁴ Angela McDonnell,³ Aaron Liston<sup>1,**

1</sup>2Department of Botany and Plant Pathology, Oregon State University, 2082 Cordley Hall, Corvallis, Or
²3Pacific Northwest Research Station, USDA Forest Service, 3200 SW Jefferson Way, Corvallis, Oregon 9
³4Department of Botany, Oklahoma State University, 301 Physical Sciences, Stillwater, Oklahoma 74078
⁴5Institute of Botany, Academy of Sciences of the Czech Republic, CZ-25243 Průhonice, Czech Republic

^*These authors contributed equally to this work.
^**Author for correspondence: listona@science.oregonstate.edu