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Luis R. Padilla,1,2 Noah K. Whiteman,3 Jane Merkel,1,3 Kathryn P. Huyvaert,3 Patricia G. Parker1,3
1Saint Louis Zoo, 1 Government Drive, St. Louis, Missouri 63110, USA 2College of Veterinary Medicine, University of Missouri-Columbia, 203 Veterinary Medicine Building, Columbia, Missouri 65211, USA 3Department of Biology, University of Missouri-St. Louis, 8001 Natural Bridge Road, St. Louis, Missouri 63121, USA
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