Price, M. J. (2014). Hedwig-Schwägrichen types catalogue update (1): new typifications and typification corrections. Candollea 69: 119–133. In English, English and French abstracts.
An update of the “Catalogue of the Hedwig-Schwägrichen Herbarium (Part 1)” of 2005 has been compiled herein. It collates the 38 recently published typifications for the Hedwig moss names and includes updated information on 12 typifications that were unintentionally omitted or incorrectly cited in the original catalogue. Information on the place and page of publication of each typification as well as the details of the types themselves is given for each name concerned. Given the complexities of the typification process of the Hedwig material (for example, the historical nature of the material, presence of different entities with different temporal or geographic origins on the type sheet, two or more species attached to the same sheet, the presence of Schwägrichen's annotations, labels that do not correspond exactly with the protologue or validating description) the correct citation of the G-Hedwig herbarium sheet and of the elements on it being typified are of crucial importance. To facilitate the unequivocal identification of the Hedwig specimens in G a list of the barcodes assigned to each of the Hedwig herbarium sheets figured in the original catalogue is provided.
Introduction
The importance of the Hedwig-Schwägrichen collection to moss taxonomy has been outlined in previous works (see Florschütz, 1960; Geissler, 2000; Price, 2005). The Hedwig part of the Hedwig-Schwägrichen herbarium is intrinsically linked with the publication “Species muscorum frondosorum” (Hedwig, 1801): the designated starting point for the nomenclature of mosses (excluding the Sphagnaceae). The goal of the ongoing work on the Hedwig-Schwägrichen collection is to create a set of reference sources that contain more accessible information on the types and typifications of the Hedwig and Schwägrichen moss names. The collation of the published typifications forms a precursor to the creation of a typification registration database by the botanical community.
Since the publication of the “Catalogue of the Hedwig-Schwägrichen herbarium (Part I)” by Price (2005) 38 new typifications for the Hedwig moss names have been published and 12 entries to the original catalogue that needed correcting were discovered (see the full list below). With the aim of providing a full record of the Hedwig types, and the typifications of the Hedwig moss names, an update to the original catalogue has been compiled. Publication information on the new typifications, including information on the designation of the material (if given), and the typifications that were unintentionally omitted or incorrectly cited in Price (2005) are indicated.
To maintain coherence between the original catalogue of Price (2005) and current list the format adopted herein is as follows:
BN: basionym.
CN/SY: current use or synonymy for the basionym.
LI: label information copied from the herbarium sheet indicated.
PL: type citation from the protologue.
LT: information on the place of publication of the type and the type designation. If given, information on the specific type material designated is included. The G barcode for the Hedwig herbarium sheet is given in square brackets.
Figure: reference to the scan of the herbarium sheet in Price (2005), with a reference to any subsequent works that contain an image of the herbarium sheet with the type indicated.
Notes: additional information, when present, for each entry.
The barcodes for the Hedwig herbarium sheets are given in Appendix 1 for each specimen treated in Price (2005) so that any typified specimens can be unequivocally associated with identifiable herbarium sheets within the Hedwig collection in G.
Newly published typifications and corrections to typifications of the Hedwig moss names
BN: Anictangium aquaticum Hedw., Sp. Musc. Frond.: 41. 1801.
CN / SY: Cinclidotus aquaticus (Hedw.) Bruch & Schimp., Bryol. Eur. 3: 170. 1842.
PL: Alpestrium aquarum incola, Scotiae, Helvetiae, Austriae, Galliae.
LI: [Anoectangium aquaticum Hedw. Stirp. III.t. a. von Hedwig b. (four illegible words) 1800 c. von Host 1799 (three illegible words)].
LT: Ederra (2006: 792), lectotype in G, “specimens marked ‘a’”. [G00040078].
Figure: Price (2005: Fig. 6).
BN: Anictangium cirrosum Hedw., Sp. Musc. Frond.: 42, t. V, f. 1–2. 1801.
CN / SY: Macromitrium cirrosum (Hedw.) Brid., Bryol. Univ. 1: 316. 1826.
PL: Jamaica, Montserrat.
LI: Anictangium cirrhosum Spec. Musc. p. 42. t. 5. f. 1–3. Hypnum cirrhosum Swartz. Prodr. p. 142. [b. St. Vincent Menzies c. var. longifolia Menzies] Jamaica, Montserrat.
LT: Wilbraham & Price (2013: 122), lectotype in G, “Jamaica, sin. coll., s.n., G (G00040187, Hb. Hedwig-Schwägrichen). First stem on the left of the upper row (specimen 1, Figure 1) on herbarium sheet G00040187 labelled ‘Anictangium cirrhosum Spec. Musc. p. 42. t. 5. f. 1–3. Hypnum cirrhosum Swartz. Prodr. p. 142. [b. St. Vincent Menzies c. var. longifolia Menzies] Jamaica, Montserrat.’ The handwriting of Schwägrichen is indicated using square brackets”. [G00040187].
Figure: Price (2005: Fig. 8), Wilbraham & Price (2013: Fig. 1, with lectotype indicated).
BN: Anictangium lapponicum Hedw., Sp. Musc. Frond.: 40. 1801.
CN / SY: Amphidium lapponicum (Hedw.) Schimp., Coroll. Bryol. Eur. 39. 1856.
PL: In alpibus Lapponiae. (In Hercynia Ehrh. Schrad.).
LI: Hedwigia lapponica Hedw. St. Cr. v.3. p. t. nomine Gymnost. Exempla Cl. Swartz.
LT: See Frahm & al. (2000: 175), for discussion of the type material. Ochyra & al. (2008: 174), lectotype in G, “The first specimen from the right on the sheet labelled ‘Hedwigia lapponica Hedw. St. Cr. v.3. p. t. nomina Gymnost. Exempla Cl. Swartz.’ — G-Hedwig-Schwägrichen”. [G00040003].
Figure: Price (2005: Fig. 9).
BN: Barbula fallax Hedw., Sp. Musc. Frond.: 120. 1801.
CN / SY: Didymodon fallax (Hedw.) R.H. Zander, Phytologia 41: 28. 1978.
PL: In interstitiis muscorum, ambulacris Germaniae, Galliae, in Africa secundum Bridelium, ad Lancaster Pensilvaniae Hedwig.
LI: Barbula fallax Hedw. St. Cr. Vol. 1. p. 62. t. 24. Chemnitii lecta. [G00040367].
LT: Zander (1978: 28), lectotype in G, “Barbula fallase [sic] Hedw. St. Cr. Vol. 1. P. 62. T. 24. Cheminitis [sic] lecta”. [G00040367].
Figure: Price (2005: Fig. 19).
Notes: The typification of Zander (1978) was inadvertently omitted in Price (2005).
BN: Bartramia halleriana Hedw., Sp. Musc. Frond.: 164. 1801.
CN / SY: Bartramia halleriana Hedw., Sp. Musc. Frond.: 164. 1801.
PL: Habitat in rupibus silvarum montosarum granite constantibus in Saxonia, Hercynia, monte pinifero Franconiae, in alpibus Judenburgensibus Styriae, Carinthia, nec non in Helvetia.
LI: Bartramia halleriana Hedw. St. Crypt Vol. II p.III. t. 40. Bryum laterale S.V* Planta delineata.
LT: Ederra (2006: 791), lectotype in G, “the third specimen on the left on the sheet — (Herbarium Hedwig-Schwägrichen in G)”. [G00040042].
Figure: Price (2005: Fig. 27).
BN: Bartramia pomiformis Hedw., Sp. Musc. Frond.: 164. 1801.
CN / SY: Bartramia pomiformis Hedw., Sp. Musc. Frond.: 164. 1801.
PL: In umbrosis, fissuris rupium per omnem Europam.
LI: Bartramia pomiformis Hedw. St. Cr. Vol. II. p. Bryum pomiforme S. V. [b. exempla picta a Bock].
LT: Ederra (2006: 792), lectotype in G, “first specimen to the left in the upper row — (Herbarium Hedwig-Schwägrichen in G)”. [G00040043].
Figure: Price (2005: Fig. 28).
BN: Bryum argenteum Hedw., Sp. Musc. Frond.: 181. 1801.
CN / SY: Bryum argenteum Hedw., Sp. Musc. Frond.: 181. 1801.
PL: Ad muros, tecta, rupes Europae.
LI: Bryum argenteum antheris pendulis surculis cylindricis imbricatis lævibus Linn. Spec. pl. 2. p. 1586. n. 27. Bryum pendulum iulaceum argenteum et sericeum Dill H. Musc. 392 t. 50 f. 62. [a. Brasil Beyrich ad muros Rio Janeiro, b. lanatum Palisot c. argyrioides Brid. Bryo U 1. 846.].
LT: Ochyra & al. (2008: 453), lectotype in G, “the central specimen glued to the sheet between the specimens marked ‘b’ and ‘a’ on the sheet bearing the label ‘Bryum argenteum antheris pendulis surculis cylindricis imbricatis lævibus Linn. Spec. pl. 2 p. 1586. n. 27. Bryum pendulum iulaceum argenteum et sericeum Dill H. Musc. 392 t. 50 f. 62.’ G-Hedwig/ Schwägrichen”. [G00040370], syntype in G [G00040059].
Figure: Price (2005: f. 31 & 30, respectively).
BN: Bryum dichotomum Hedw., Sp. Musc. Frond.: 183, t. XLII, f. 8–12. 1801.
CN / SY: Bryum dichotomum Hedw., Sp. Musc. Frond.: 183. 1801.
PL: Nova Seelandia.
LI: Bryum dichotomum Spec. Musc. p. 183. t. 42, f. 8–12. aus Neu Seeland.
LT: Ochyra & al. (2008: 462), lectotype in G, “the third shoot from the left on the sheet labelled ‘Bryum dichotomum Spec. Musc. p. 183. t. 42, f. 8–12. aus Neu Seeland’ — G-Hedwig/ Schwägrichen”. [G00040365].
Figure: Price (2005: Fig. 35).
BN: Bryum patens Hedw., Sp. Musc. Frond.: 86. 1801.
CN / SY: Grimmia ramondii (DC) Margad. Lindbergia 1: 128. 1972.
PL: Ad rupes in rivulis alpinis Scotiae. Ben Nevis.
LT: Geissler & Maier (1995: 499), specimen not in G. Lectotype in BM [herb. Dickson, BM000966028], isotype in BM [BM001058734].
BN: Cynontodium capillaceum Hedw., Sp. Musc. Frond.: 57. 1801.
CN / SY: Distichium capillaceum (Hedw.) Bruch & Schimp., Bryol. Eur. 2: 156 (fasc. 29–30. Monogr. 4). 1846.
PL: Saxonia inferior, Niedersachswerfen et Neuhof, in montibus gypsaceis prope Steigerthal, ad ripas fluvii Bode, (in calcareis Franconiae, alpestribus Austriae) Suecia, Islandia in fissuris rupium.
LI: Swartzia capillacea Hedw. St. Cr. Vol.II. p. 72. t. 26. [b. ex ins Melville (illegible word)].
LT: Ochyra & al. (2008: 131), lectotype in G, “The right specimen in the lower row, on the sheet labelled ‘Swartzia capillacea Hedw. St. Cr. Vol.II. p. 72. t. 26.’ — G-Hedwig/ Schwägrichen”. [G00040111].
Figure: Price (2005: Fig. 42).
BN: Cynontodium inclinatum Hedw., Sp. Musc. Frond.: 58. 1801.
CN / SY: Distichium inclinatum (Hedw.) Bruch & Schimp., Bryol. Eur. 2: 157, 194. 1846.
PL: none.
LI: Swartzia inclinata Hedw. St. Cr. Vol.II. p. 74. t. 27.
LT: Ochyra & al. (2008: 135), lectotype in G, “The middle specimen with the distinctly inclined capsule, on the sheet labelled ‘Swartzia inclinata Hedw. St. Cr. Vol.II. p. 74. t. 27.’ G-Hedwig-Schwägrichen” [G00040112].
Figure: Price (2005: Fig. 44).
BN: Dicranum cerviculatum Hedw., Sp. Musc. Frond.: 149. 1801.
CN / SY: Dicranella cerviculata (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: In Suecia et in turfosis aridis ericetis Hamsteadiani prope Londinum. (nec non in Sudetibus Germaniae lectum a Ludwigio. S.).
LI: Dicranum cerviculatum Hedw. St. Cr. III. p. 89. t. 27. A. Suec. D. Swartz.
LT: Ellis & Price (2013a: 277), lectotype in G, “Sweden, Swartz s.n. [‘Dicranum cerviculatum Hedw. St. Cr. III. p. 89. t. 27. A. Suec. D. Swartz’] Hb. Hedwig-Schwägrichen (G000 40097), the three fertile plants on herbarium sheet G0004 0097”. [G00040097].
Figure: Price (2005: Fig. 49).
BN: Dicranum crispum Hedw., Sp. Musc. Frond.: 132. 1801.
CN / SY: Dicranella crispa (Hedw.) Schimp., Coroll. Bryol. Europ. 13. 1856.
PL: In Suecia prope Upsalam, Scotiae paludosis et alpinum Sanensium humidis.
LI: Dicranum crispum Hedw. St. Cr. Vol. II. p. 91. t. 33. Exemplaria in Suecia ab Ehrhardo lecta.
LT: Ellis & Price (2013a: 278), lectotype in G, “Sweden, Ehrhart s.n. ‘Dicranum crispum Hedw. St. Cr. Vo. II. P. 91. T. 33. Exemplaria in Suecia ab Ehrhardo lecta” (Hb. Hedwig-Schwägrichen — G00040017), the eight plants on herbarium sheet G00040017”. [G00040017].
Figure: Price (2005: Fig. 51).
BN: Dicranum curvatum Hedw., Sp. Musc. Frond.: 132, t. XXXI, f. 7–12. 1801.
CN / SY: Dicranella subulata (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: Locus ad declivitates viarum cavarum montanarum, inter Didymodon homomallum.
LI: Dicranum curvatum Sp. Musc. 132. t. 31, f. 7–12. ad declivia viarus cavaras Ludwig.
LT: Ellis & Price (2013a: 279), lectotype in G, “[Germany/ Poland, Lausitz?] Ludwig s.n. ‘Dicranum curvatum Sp. Musc. 132. t. 31, f. 7.–12. Ad declivia viarus cavaras Ludwig.’ (Hb. Hedwig-Schwägrichen – G00040103), plants in “b”–“d” on herbarium sheet G00040103”. [G00040103].
Figure: Price (2005: Fig. 52).
BN: Dicranum interruptum Hedw., Sp. Musc. Frond.: 129, t. XXIX, f. 9–12. 1801.
CN / SY: Dicranella heteromalla (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: In montosis uliginosis silvatici ad arbores, praesertim putrescentes, in Saxonia Chemnizii, Zeisigwald (in Anglia, Germania prope Halam Saxonum Bridel.).
LI: Dicranum interruptum Hedw. Spec. Musc. p. 129. t. 29, f. 8. 9.–12. Specimina patiora Chemnitzii lecta.
LT: Ellis & Price (2013a: 282), lectotype in G, “[Germany/ Saxony] Chemnitz, [Hedwig?] s.n. ‘Dicranum interruptum Hedw. Spec. Musc. p. 129. t. 29, f. 8. 9.–12. Specimina patiora Chemnitzii lecta.’ (Hb. Hedwig-Schwägrichen — G00040100), all plants on sheet G00040100”. [G00040100].
Figure: Price (2005: Fig. 60).
BN: Dicranum longifolium Hedw., Sp. Musc. Frond.: 130. 1801.
CN / SY: Paraleucobryum longifolium (Hedw.) Loeske, Hedwigia 47: 171. 1908.
PL: In Suecia Upsaliae et in Scoticis alpibus ad rivulorum ripas (in Hercynia et in Sudetibus Ludwig, in Austria Host).
LI: [Dicranum longifolium Hedw. St. Cr. III. p. 341. t. 9. Upsalia Ehrh ad J. Hedwig].
LT: Frahm & Isoviita (1988: 95), holotype in G, corrected in Price (2012: 302), lectoype in G, “the two extant stems on herbarium sheet G00040217, labelled as above, and as illustrated in Hedwig (1791–1792)” [G00040217].
Figure: Price (2005: Fig. 63).
Notes: The original typification by Frahm & Isoviita (1988) was not listed in Price (2005).
BN: Dicranum orthocarpum Hedw., Sp. Musc. Frond.: 131, t. XXX, f. 1–5. 1801.
CN / SY: Dicranella heteromalla (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: Locus terrestris. Lancaster in Pensilvania.
LI: Dicranum orthocarpum Sp. Musc. p. 131. tab 30, f. 1–5. ad Lancaster Pensylvaniae lect.
LT: Ellis & Price (2013a: 283), holotype in G, “[North America, USA], Pennsylvania, Lancaster, [Mühlenberg] s.n. ‘Dicranum orthocarpum Sp. Musc. p. 131. t. 30, f. 1–5. ad Lancaster Pensylvaniae lect.’ (Hb. Hedwig-Schwägrichen — G00040101), all plants on sheet G00040101” [G00040101].
Figure: Price (2005: Fig. 65).
BN: Dicranum purpurascens Hedw., Sp. Musc. Frond.: 137, t. XXXV, f. 1–7. 1801.
CN / SY: Ceratodon purpureus (Hedw.) Brid., Bryol. Univ. 1: 480. 1826.
PL: Lancaster Pensilvaniae in terra humoso-turfosa.
LI: Dicranum
purpurascens Spec. Musc. p. 137. t. 35, f. 1–7. Circa Lancaster Pensylvan [b a Pens, f. nom.].LT: Burley & Pritchard (1990: 37–38), holotype in PH.
Figure: Price (2005: Fig. 70).
Notes: the original typification by Burley & Pritchard (1990) was not listed in Price (2005). Original Hedwig material is present in G [G00040077], shown in Price (2005: Fig. 70).
BN: Dicranum purpureum Hedw., Sp. Musc. Frond.: 136, t. XXXVI, f. 1–11. 1801.
CN / SY: Ceratodon purpureus (Hedw.) Brid., Bryol. Univ. 1: 480. 1826.
PL: Locus terrestris, ad ligna putrescentia, per omnem Europam vulgatissima species, inque America septentrionali.
LI: Bryum tenue stellatum setis purpureis. Dill. Musc. 386. t. 49, f. 51. Mnium purpureum caule dichotomo, axillis pedunculiferis, anthera exerta, foliis carinatis. Linn. Spec. ed. II. 1575.6.
LT: Burley & Pritchard (1990: 37), lectotype in G [G0004 0077].
Figure: Price (2005: Fig. 71).
Notes: The original typification by Burley & Pritchard (1990) was inadvertently omitted in Price (2005).
BN: Dicranum pusillum Hedw., Sp. Musc. Frond.: 139. 1801.
CN / SY: Dicranella cerviculata (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: Prope Berolinum, Vindobonam, in Hercyniae umbrosis et Scotiae pascuis ad terram densis caespitibus.
LI: Dicranum pusillum Hedw. St. Cr. Vol. II. p. 78. t. 29.13.
LT: Ellis & Price (2013a: 285), lectotype in G, “[Germany/Berlin?] [Willdenow?] s.n. ‘Dicranum pusillum Hedw. St. Cr. Vol. II. p. 78. t. 29.B’ (Hb. Hedwig-Schwägrichen — G00040098), all plants on sheet G00040098” [G00040098].
Figure: Price (2005: Fig. 72).
BN: Dicranum schreberianum Hedw., Sp. Musc. Frond.: 144, t. XXXIII, f. 6–10. 1801.
CN / SY: Dicranella schreberiana (Hedw.) H.A. Crum & L.E.Anderson, Mosses E. N. Amer. 1: 169. 1981.
PL: Locus ad fossas argillosas Lipsiae.
LI: Dicranum schreberianum Hedw. Sp. M. p. 144. t. 33. fig. 6–10. Bryum crispum Schreb. Spicil.
LT: Ellis & Price (2013a: 285), lectotype in G, “[Germany, Saxony, Leipzig], sin. coll. s.n. ‘Dicranum schreberianum Hedw. Sp. M. p. 144. t. 33. fig. 6–10. Bryum crispum Schreb. Spicil.’ (Hb. Hedwig-Schwägrichen – G00040018), all plants on sheet G00040018” [G00040018].
Figure: Price (2005: Fig. 73).
BN: Dicranum subulatum Hedw., Sp. Musc. Frond.: 128, t. XXXIV, f. 1–5. 1801.
CN / SY: Dicranella subulata (Hedw.) Schimp., Coroll. Bryol. Eur. 13. 1856.
PL: Sueciae loco humido mense Septembri lectum misit D. Swartz.
LI: Dicranum subulatum Sp. M. p. 128. t. 34, f. 1–5. in humidis Swec. Swartz.
LT: Ellis & Price (2013a: 287), lectotype in G, “Sweden, Swartz s.n. ‘Dicranum subulatum Sp. M. p. 128. t. 34, f. 1–5. in humidis Swec. Swartz’ (Hb. Hedwig-Schwägrichen — G00040102), 3 shoots at extreme right of specimen, as indicated in Figure 4” [G00040102].
Figure: Price (2005: Fig. 76), Ellis & Price (2013: Fig. 4, with lectotype indicated).
BN: Funaria hygrometrica Hedw., Sp. Musc. Frond.: 172. 1801.
CN / SY: Funaria hygrometrica Hedw., Sp. Musc. Frond.: 172. 1801.
PL: Locis subhumidis umbrosis aut apertis, in rupibus terra tenui tectis per omnem Germaniam.
LI: Bryum bulbiforme aureum, calyptra quadrangulari, capsulis pyriformibus nutantibus Dill. H. Musc. 407. t. 52, f. 75. Mnium hygrometricum acaule anthera nutante calyptra reflexa tetragona Linn. Spec. pl. 2. p. 1575. n. 5.
LT: Ochyra & al. (2008: 118), lectotype in G, “the second specimen from the left in the upper row forming a tuft with many mature capsules on the sheet labelled ‘Bryum bulbiforme aureum, calyptra quadrangulari, capsulis pyriformibus nutantibus Dill. H. Musc. 407. t. 52, f. 75. Mnium hygrometricum acaule anthera nutante calyptra reflexa tetragona Linn. Spec. pl. 2. p. 1575. n. 5’ — G-Hedwig-Schwägrichen” [G00040143].
Figure: Price (2005: Fig. 101).
BN: Grimmia alpicola Hedw., Sp. Musc. Frond.: 77, t. XV, f. 1–5. 1801. nom. rej.
CN / SY: Schistidium alpicola (Hedw.) Limpr., Die Laubm. Deutschl. 1: 707. 1889.
PL: Suecia. O. Swartz. In saxis, ad rivulos alpinos.
LI: Grimmia alpicola Spec Musc. p. 77. t. 15, f. 1–5. accepta ab O. Swartz.
LT: Bremer (1980: 2), “type in G (information communicated by Deguchi)”. See also Price & Maier (2011a: 288). Original material: s. loc., s.d. [Swartz] s.n. (G00040295), all plants on herbarium sheet G00040295.
Figure: Price (2005: Fig. 103).
BN: Grimmia cribrosa Hedw., Sp. Musc. Frond.: 76. 1801.
CN / SY: Coscinodon cribrosus (Hedw.) Spruce, Ann. Mag. Nat. Hist., ser. 2, 3: 491. 1849.
PL: In petris Goslariae, nec non Hassiae montis Meisner elegantem speciem primi dexerunt Persoon et Schrader.
LI: Grimmia cribrosa Hedw. St. Cr. Vol.III.p. 73.t. 31. A [a J Hedw. b. a Goslar c. Gogendbrug am Ochsenkopf c. An der Mühle unter der Salbergstra e Schwarzenbg. (illegible word) e Scotia Arnott 1839].
LT: Deguchi (1978: 244), holotype in G [G00040082]; Price & Maier (2011b: 795), lectotype in G, “the element annotated “b” (Fig. 2, element 2) on the original Hedwig herbarium sheet [G00040083] in G. This element corresponds to fig. 3 of t. 31 in Hedwig (1791–1792)” [G00040083].
Figure: Price (2005: Fig. 105), Price & Maier (2011b: Fig. 2, with lectotype indicated).
Notes: The specimen identified as the holotype by Deguchi (1978: 244) was not original Hedwig material (see discussion in Price & Maier 2011a).
BN: Grimmia plagiopodia Hedw., Sp. Musc. Frond.: 78, t. XV, f. 6–13. 1801.
CN / SY: Grimmia plagiopodia Hedw., Sp. Musc. Frond.: 78. 1801.
PL: (In Saxoniae rupibus detexit Cl. Flügge.).
LI: [Grimmia plagiopus a Hg l. n. b a Flörkio].
LT: Degughi (1984: 22), holotype in G. Ochyra & al. (2008: 271), lectotype in G, “The right specimen in the upper row on the sheet labelled ‘Grimmia plagiopus a Hg l. n. b a Flörkio’ — G-Hedwig-Schwägrichen” [G00000040348].
Figure: Price (2005: Fig. 106).
Notes: The typification of Ochyra & al. (2008) was missed in Maier (2010: 282) and Price & Maier (2011a: 295).
BN: Gymnostomum heimii Hedw., Sp. Musc. Frond.: 32. 1801.
CN / SY: Desmatodon heimii (Hedw.) Mitt., J. Proc. Linn. Soc., Bot. 8: 28. 1864.
PL: Prope Spandau primus invenit Heim, consiliarius aulicus, Berilonensis. Circa Malchin ducatus Mecklenburgici clariss Timm; Upsaliae ad palos vetustos Ol. Swartz, (in Austria D. Host in Sudetibus Ludwig.).
LI: Gymnostomum Heimii Hedw. St. Cr. I.80.t. 30. [b en putis fontis in arce Upsal. c. Yarmouth] ab ipso inventore Cons. Aulico Heim acceptus.
LT: Ochyra & al. (2008: 359), lectotype in G: “The specimen marked “a” on the sheet labelled ‘Gymnostomum Heimii Hedw. St. Cr. I.80.t. 30. a ab ipso inventore Cons. Aulico Heim acceptus (manu Hedwig) b. ex. putis fontis in arce Upsal. c. Yarmouth, (manu Schwägrichen)’ — G-Hedwig-Schwägrichen” [G00040352].
Figure: Price (2005: Fig. 108.).
BN: Gymnostomum ovatum Hedw., Sp. Musc. Frond.: 31. 1801.
CN / SY: Pterygoneurum ovatum (Hedw.) Dixon, Rev. Bryol. Lichénol. 6: 96. 1934.
PL: In muris argillaceis circa Lipsiam aliisque regionibus frequens.
LI: Gymnostomum ovatum Hedw. Crypt. 1, 15, 6.
LT: Guerra & al. (1995: 166–167), lectotype in G, “In muris argillaceis circa Lipsiam. — En el herbario Hedwig-Schwaegrichen en G existe una muestra de P. ovatum (sub Gymnostomum ovatum Hedw.) con una etiqueta manuscrita por Hedwig en los términos anteriores que podría corresponder con parte del material original que se utilizó para la descripción (Lectótipo: G!)”. [G00040267].
Figure: Price (2005: Fig. 111).
Notes: The typification of Guerra & al. (1995) was inadvertently omitted in Price (2005).
BN: Gymnostomum pennatum Hedw., Sp. Musc. Frond.: 31. 1801.
CN / SY: Schistostega pennata (Hedw.) F.Weber & D.Mohr, Index Mus. Pl. Crypt. 2. 1803.
PL: Ad muros septentrionales civitatis Mariaemontani in Saxonia: (in Sudetibus invenit Ludwig, Erlangae in saxo arenario S. Etiam in Hercynia, Pyrmont occurit teste Hoffm. flor. G. II. p. 28.) in Anglia primus invenit D. Newberg in cavitatibus aggerum sub sepibus antiquis solo pingui comitatus Devoniae. Vere sporangia matura.
LI: Gymnostomum pennatum Hedw. St. Cr. 1. 77. t. 29. [sub Dicksonia pusilla ab Erhard ad Schrebr e m. Bruch. Granite e Euchsburg (illegible symbol) d (illegible word)] Mariaemontanii in Saxonia lect. [in Gneisso].
LT: Ellis & Price (2012: 20), lectotype in G, “[Germany], Saxony, Marienburg, Hedwig s.n. ‘Gymnostomum pennatum Hedw. St. Cr. 1. 77. t. 29. Mariaemontanii in Saxonia lect. [in Gneisso]’ (G [G00040297] [a])” [G00040297].
Figure: Price (2005: Fig. 112), Ellis & Price (2012: Fig. 2, with lectotype indicated).
BN: Gymnostomum pulvinatum Hedw., Sp. Musc. Frond.: 36, t. III, f. 1–7. 1801.
CN / SY: Gymnostomum pulvinatum Hedw., Sp. Musc. Frond.: 36. t. III, f. 1–7. 1801.
PL: Ad pagum Reinhausen, prope Goettingam, legit misitque Hoffmann et Persoon.
LT: Ochyra (1989a: 91), lectotype in LE-Mertens. See Ochyra (1989a, b) for discussion on this name and associated specimens. The details on the location of the lectotype material were inadvertently omitted in Price (2005).
BN: Leskea caespitosa Hedw., Sp. Musc. Frond.: 233, t. XLIX, f. 1–5. 1801.
CN / SY: Acroporium caespitosum (Hedw.) Buck, Brittonia 23: 310. 1983.
PL: Hispaniola, in ligno putrefacto.
LI: Leskea cespitosa Hedw. Sp. Musc. 233 t. 49, f. 1–5. Hypnum cespitosum Swartz Prodr. p. 142. [b. ad tr. arb. (illegible word) antiqua pr Mandioia Brasil Beyrich] Hispanolae lecta et accepta ab ipso O. Swartz.
LT: Buck (1983: 309), lectotype in G. [G00040303].
Figure: Price (2005: Fig. 198).
Note: The typification of Buck (1983) was inadvertently omitted in Price (2005).
BN: Leskea mollis Hedw., Sp. Musc. Frond.: 234. 1801.
CN / SY: Weymouthia mollis (Hedw.) Broth., Nat. Pflanzenfam. 1(3): 812. 1906.
PL: Habitat in nova Seelandia.
LI: Leskea mollis St. Cr. IV. P. 103. t. 40. Aus Neu Holland, im.
LT: Tangney (2010: 147), lectotype in G, “Herb. Hedwig, New Zealand, St. Cr. IV. p. 103. t. 40 (G)”. [G00040342].
Figure: Price (2005: Fig. 210).
BN: Meesia uliginosa Hedw., Sp. Musc. Frond.: 173. 1801.
CN / SY: Meesia uliginosa Hedw., Sp. Musc. Frond.: 173. 1801.
PL: In pratis turfosis paludosis Hassiae, Saxoniae, aliisque Germaniae locis, in rupibus alpestribus calcareis Austriae vulgaris.
LI: Meesia uliginosa Hedw. Cryptog. p. 1. t. 1, f. 1–2. Bryum trichodes Linn. [b. angustifolia alp Helvet. c. longirostris Brid. ms d. stricto a. Bridel alp Carinth.].
LT: Ochyra & Lewis-Smith (1999: 6), syntypes in G. Ochyra & al. (2008: 492), lectotype in G, “the specimen glued to the sheet and marked “a” bearing the label ‘Meesia uliginosa Hedw. Cryptog. p. 1. t. 1, f. 1–2. Bryum trichodes Linn. {manu Hedwig} b. angustifolia alp Helvet. c. longirostris Brid. Mt. d. stricta a Bridel alp Carinth {manu Schwägrichen} — G-Hedwig-Schwägrichen”. [G00040192].
Figure: Price (2005: Fig. 231).
BN: Mnium chrysocomum Hedw., Sp. Musc. Frond.: 74. 1801.
CN / SY: Breutelia chrysocoma (Hedw.) Lindb., Öfvers. Förh. Kongl. Svenska Vetensk.-Akad. 20: 389. 1863.
PL: Locus in paludosis Scotiae, Cambriae, Angliae.
LI: [Bartramia arcuata a. Turn b. ins Bourbon Taylor].
LT: Price & Ellis (2011: 314), lectotype in G, “[British Isles] sin. coll., s.n. (Hb. Hedwig-Schwägrichen ex Hb. Turner, G [G00040056]), the first specimen on the left of the herbarium sheet (Specimen 1 [‘a’])”. [G00040056].
Figure: Price (2005: Fig. 233), Price & Ellis (2011: Fig. 1, with lectotype indicated).
BN: Mnium crudum Hedw., Sp. Musc. Frond.: 189. 1801.
CN / SY: Pohlia cruda (Hedw.) Lindb., Musci Scand. 18. 1879.
PL: Locis udis, montosis ad saxa in Germania passim.
LI: Bryum pendulum hornum molle foliis et lanceolatis et gramineis Dill. H. M. 401. t. 51, f. 70 Mnium crudum antheris pendulis calyptra recurvata foliis pellucidis Linn. Spec. pl. 2. p. 1576. n-ii.
LT: Ochyra & al. (2008: 440), lectotype in G, “the first right specimen on the sheet bearing the label ‘Bryum pendulum hornum molle foliis et lanceolatis et gramineis Dill. H. M. 401. t. 51, f. 70 Mnium crudum antheris pendulis calyptra recurvata foliis pellucidis Linn. Spec. pl. 2. p. 1576. n-ii’ G Hedwig/ Schwägrichen” [G00040361].
Figure: Price (2005: Fig. 234).
BN: Mnium pseudotriquetrum Hedw., Sp. Musc. Frond.: 190. 1801.
CN / SY: Bryum pseudotriquetrum (Hedw.) P. Gaertn., B. Mey. & Scherb., Oekon. Fl. Wetterau 3: 102. 1802.
PL: Locis apricis udis paludosis Europae.
LI: Duby -Boissier Herbarium: “Meesia longiseta Hostio Austr 94. Br. pstriq. Simile Timmiae austriacae, f differt. Foliio supremis ovatis, 1 ovatolanciolatus margine revolutis, quae integra videntur, (illegible word) conduplicatus, caulinis per id, quod margine valde revolutus lanceolatis (illegible word) deest (illegible word) basis. Habet aliquum similitudineum cum br. pedunci (illegible word, illegible symbol) rigidi distinguent”.
LT: Demaret & Empain (1985: 276), lectotype in G “échantillon envoyé par Host (Autriche) en 1799 à Schwägrichen ou à Hedwig…”, clarified in Ochyra & al. (2008: 472), lectotype in G, “Herb. Duby. junct. Herb. Boissier 1886. Br. ptrq. Meeisa longiseta Hostio ovalotanceolatis maegine revolutis, quae… Sub nomine Meesiae longisetae misit Host (austriacus) AD 1799 ad Schwaegrichen vel ad Hedwigium. Bryum pseudotriquetrum Hedw. Paris Index bryologicus, p. 201 — G — Hedwig/Schwägrichen”. [G00040366],
Figure: Price (2005: Fig. 241).
Note: The original typification of Demaret & Empain (1985) was inadvertently omitted in Price (2005).
BN: Mnium tetragonum Hedw., Sp. Musc. Frond.: 73. 1801.
CN / SY: Conostomum tetragonum (Hedw.) Lindb., Öfvers. Förh. Kongl. Svenska Vetensk.-Akad. 20: 392. 1863.
PL: none.
LI: [Conostomum boreale a pita e Suec. Sw. b. e Groenland c. ex Angl. Turner.]
LT: Frahm & al. (1996: 106), “in alpibus scoticis, Ben Lomond, Dickson s.n. (holotypus BM [BM000079531])”.
Note: The typification of Frahm & al. (1996) was inadvertently omitted in Price (2005). The Hedwig herbarium sheet present in G [G00040081] and depicted in Price (2005: Fig. 246), annotated “Conostomum boreale” contains potential type material for Swartz's Conostomum boreale Sw. that was described in 1806.
BN: Polytrichum alpinum Hedw., Sp. Musc. Frond.: 92, t. XIX, f. 2, b. 1801.
CN / SY: Polytrichastrum alpinum (Hedw.) G.L.Sm., Mem. New York Bot. Gard. 21: 37. 1971.
PL: Habitat in alpibus Scoticis, Scandinavicis, Tyrolensibus, Helveticis, in Sudetibus, Bructero Hercyniae etc.
LI: [Polytrichum alpinum e. Bruct. b. ambiguum a Rich.].
LT: Ochyra & al. (2008: 94), lectotype in G, “The central specimen in the upper row with four mature capsules on the sheet labelled ‘Polytrichum alpinum e. Bruct.’ — G-Hedwig-Schwägrichen”. [G00040254].
Figure: Price (2005: Fig. 283).
BN: Polytrichum juniperinum Hedw., Sp. Musc. Frond.: 89, t. XVIII, f. 6–10. 1801.
CN / SY: Polytrichum juniperinum Hedw., Sp. Musc. Frond.: 89. 1801.
PL: Locus montosus, subalpestris, delineata exempla ex Helvetia.
LI: [Polytrichum iuniperinum b. canescens Rich. (illegible word) c. in muris Lantz.].
LT: Ochyra & al. (2008: 105), lectotype [icon in] Hedwig (1801: Pl. 18. figs. 6–10), epitype “E. Bauer: Musci Europaei Exsiccati No. 1549b: Polytrichum juniperinum Willd., c.fr. Et. Masc. Voralberg: Bregenz, linker oberer Rand des Schliefertobers auf trockenen Waldboden, 550 m s.m. 6. Mai 1916 legit Joseph Blumrich — KRAM”.
BN: Polytrichum piliferum Hedw., Sp. Musc. Frond.: 90. 1801.
CN / SY: Polytrichum piliferum Hedw., Sp. Musc. Frond.: 90. 1801.
PL: Locis editioribus siccioribus, in ericetis, muris argillaceis. Lipsiae ad Bieniz, Sct. Theclae, in muris pagorum, Schoenfeld. etc.
LI: [Polytrichum piliferum d. a Richard b. Schleich. c. Duval c alp Brid. c. Mandon R. subrigidi fol].
LT: Ochyra & al. (2008: 101), lectotype [icon in] Dillenius (1741, Pl. 54, Fig. 3), epitype “E. Bauer: Musci Europaei Exsiccati No. 1199: Polytrichum piliferum Schreb., fr. Frankreich: Le Harve, Wald von Montgeon, April 1905 legit I. Theriot — KRAM”.
BN: Tetraphis pellucida Hedw., Sp. Musc. Frond.: 45, t. VII, f. 1, 1 a-f. 1801.
CN / SY: Tetraphis pellucida Hedw., Sp. Musc. Frond.: 45. 1801.
PL: Locis umbrosis udis, truncos deiectos putrescentes, praecipue montosarum regionum.
LI: Mnium pellucidum caule simplici foliis ovatis Linn. Spec, plant. 2. p. 1574. n.l. Mnium serpilli foliis tenuibus pellucidis Dill. Hist. Musc 232. t. 31, f. 2. Tetraphis pellucida Spec. Musc.p. t, f.
LT: Price (2010: 16), lectotype in G, “s.loc., s.d., s.coll. (G! [G00040314])”, “The second stem from the left in the group of specimens located at the top left (upper row) of the herbarium sheet [G00040314]”. [G00040314], syntype in G [G0004315].
Figure: Price (2005 : Fig. 320), Price (2010: Fig. 2), with lectotype indicated.
BN: Tortula tortuosa Hedw., Sp. Musc. Frond.: 124. 1801.
CN / SY: Tortella tortuosa (Hedw.) Limpr., Die Laubm. Deutschi. 1: 604. 1888.
PL : In montibus Pedemontii, Helvetiae, Galliae, Scotiae, Sueciae, Thuringiae, Hercyniae, in silvis arenosis Palatinatus, in rupibus calcareis Franconiae, Austriae, nec non in Virginia.
LI : [Barbula tortuosa a. J. Hedwig, b. alpes Carinth Horns. c. Ol. Swartz, c. George's Sound Menzies. d. Cap b. Sp. Menz. e. Dicranum flexifol. Hook a Menz. Cap. b-Sp, f. Cap. b. Sp. Mz.].
LT: Ellis & Price (2013b: 219), lectotype in G, “Hedwig s.n. G [G00040331], top row of specimens on sheet (plants 1-5). [G00040331],
Figure: Price (2005: Fig. 326), Ellis & Price (2013b: Fig. 1, with lectotype indicated).
BN: Trichostomum fasciculare Hedw., Sp. Musc. Frond.: 110. 1801.
CN / SY: Racomitrium fasciculare (Hedw.) Brid., Muscol. Recent. Suppl. 4: 80. 1819 [1818].
PL : Habitat ad saxa in Bructero, Achtermannshoehe aliisque Hercyniae montibus.
LI: Duby-Boissier herbarium: “Trichost. fasciculare e rupibus Bruct Schw” In another hand ‘Ex. Herb. Schwaegrichen’.
LT: Bednarek-Ochyra & Ochyra (2005: 1070), lectotype “[icon in] Dillenius (1741) f. 28 on pl. 47”; epitype, the second specimen from the left on the sheet labelled “28. Rhacomitrium fasciculare 1892. N. 28. Bryum hypnoides, hirsutie virescens, fasciculare, alpinum Hist. Musc. p. 370. N. 28” (OXF-Dillenius).
BN: Trichostomum lanuginosum Hedw., Sp. Musc. Frond.: 109. 1801.
CN / SY: Racomitrium lanuginosum (Hedw.) Brid., Muscol. Recent. Suppl. 4: 79. 1819 [1818],
PL: Montana, arida, arenosa, lapidosa loca; (maxime in frustis granitis montium subalpinorum, Bructeri, Sudetum.).
LI : Trichostomum lanuginosum Hedw. St. Cr. Vol. III. p. 3. t. 2.
LT: Frisvoll (1986: 83), lectotype in G, clarified in Ochyra & al. (2008: 289), lectotype in G, “the left specimen on the sheet labelled ‘Trichostomum lanuginosum Hedw. St. Cr. Vol. III. p. 3. t. 2.’ corresponding to fig. 2 on table 2 in Hedwig (1792) — G-Hedwig/Schwägichen”. [G00040358].
Figure: Fig. 333.
Note : see Price & Maier (2011a) under this taxon for information on the original typification.
BN: Webera nutans Hedw., Sp. Musc. Frond.: 168. 1801.
CN / SY: Pohlia nutans (Hedw.) Lindb., Musci Scand. 18. 1879.
PL : Locis siccioribus, sterilibus, arenosis, apertis et umbrosis gregatim, nec non in uliginosis turfosis sparsim in omni Germania frequens; solo pingui rarior.
LI: Webera nutans Hedw. St. Cr. 1. 9. t. 4. [b a Richardson (two illegible words). Pleurique caulis simplices quidam (illegible word) alterne rammo versus apices contin.] Specimina Lipsiae im Universitaetsholz inter Sphagn. palustre lecta.
LT: Ochyra & al. (2008: 444), lectotype [icon] Dillenius (1741 : PI. 50, Fig. 61, of Bryum trichodes laete virens, capitulis cernis oblongis), epitype “Böhmen: isergebirge, Raspenau, an einen Ealdwege beim Eisenbahndurchlasse, 360 m s.m., 18. Juli 1911 legit Jos. Blumrich' (E. Bauer, Musci europaei exsiccati No. 909) — KRAM, isoepitypes: BM, FH, G, NY, PC and apparently in many other herbaria to which these exsiccati were distributed (fide Sayre, 1971)”.
BN: Webera pyriformis Hedw., Sp. Musc. Frond.: 169. 1801.
CN / SY: Leptobryum pyriforme (Hedw.) Wilson, Bryol. Brit. 219. 1855.
PL: Locis lapidosis, sabulosis, subhumidis, in Germania.
LI: Webera pyriformis Hedw. St. Cr. 1. p. 5. t. 3. Bryum Dill. H. M. 60. Mnium pyriforme Linn. Chemnizii partim, partim Lipsiae lecta.
LT: Ochyra & al. (2008: 419), lectotype in G, “the upper left specimen on the sheet bearing the label, ‘Webera pyriformis Hedw. St. Cr. 1. p. 5. t. 3. Bryum Dill. H. M. 60. Mnium pyriforme Linn. Chemnizii partim, partim Lipsiae lecta.’ — G-Hedwig/ Schwägrichen” [G00040363].
Figure: Price (2005: Fig. 339).
BN: Weissia crispula Hedw., Sp. Musc. Frond.: 68, t. XII, f. 1–6. 1801.
CN / SY: Dicranoweisia crispula (Hedw.) Milde, Bryol. Siles. 49. 1869.
PL: Fagimonti (Buchberg) in Bohemia legit Ludwig.
LI: Weissia crispula Sp. Musc. p. 58. t. 12, f. 1. 2.–6. Fagimonti (Buchberg) in Bohemia Ludw.
LT: Ochyra & al. (2008: 210), lectotype in G, “the second specimen from the left on a sheet labelled ‘Weissia crispula Sp. Musc. p. 58. t. 12, f. 1. 2.–6. Fagimonti (Buchberg) in Bohemia Ludw.’ — G-Hedwig-Schwägrichen”. [G00040106].
Figure: Price (2005: Fig. 345).
BN: Weissia heteromalla Hedw., Sp. Musc. Frond.: 71. 1801.
CN / SY: Ditrichum heteromallum (Hedw.) E. Britton, N. Amer. Fl. 15: 64. 1913.
PL: In montanis Saxoniae silvis, ad vias cavas, udas et earum orbitas pfofundiores [sic]. (In Hercynia, Sudetibus, Austria.).
LT: Ochyra & Lewis-Smith (1998: 35), no material suitable for typification present in G. Ochyra & al. (2008: 139), lectotype [icon] Hedwig (1787: t. VIII), epitype, “a. Kopsch: Bryotheca Saxonica. No. 13. Ditrichum homomallum (Hedw.) Hpe. Erzgebirge: an Waldrändern bei Oberwiesenthal. 2.II.1919. leg. Alfr. Kopsch — KRAM”.
BN: Weissia recurvirostris Hedw., Sp. Musc. Frond.: 71. 1801.
CN / SY: Bryoerythrophyllum recurvirostrum (Hedw.). P. C. Chen, Hedwigia 80: 5. 1941.
PL: Locis lapidosis sabulosis.
LI: Duby-Boissier herbarium: “Grimmia recurvirostra Gottingae Herb. Schwaegrichen”.
LT: Ochyra & al. (2008: 329), lectotype [icon in] Hedwig (1785–1787: t. 7), epitype “[Deutschland] Kgr. Saschen: Am Elbufer bei Königstein, etwa 120 m.s.m., 26 Sept. 1905 legit W. Kreiger’ (E. Bauer, Musci europaei exsiccate Mo. 273) — KRAM; isoepitypes: BM, FH, G, NY, PC”.
Acknowledgments
This work was supported by the Conservatoire et Jardin botaniques de la Ville de Genève (CJBG). The staff of the CJBG library are acknowledged for their help in facilitating access to the literature necessary for this work. Patrick Perret, Head Librarian, provided help with nomenclatural issues. I would also like to thank Camille Agier for translating the French résumé, Eva Maier for her help in interpreting the labels in the Hedwig-Schwägrichen collection and for her comments on an earlier version of this manuscript. I acknowledge the fruitful collaboration with Mr. Len Ellis (Natural History Museum, London) on typifications of certain groups of Hedwig mosses and thank him for corrections to an earlier version of this article. I thank also Drs. María J. Cano and Paulo E. A. S. Câmara for their work on the Pottiaceae and Sematophyllaceae types, respectively, in the Hedwig-Schwägrichen collection in G.
References
Appendices
Appendix 1. —
The G barcodes for the Hedwig type specimens in G that are presented in the “Catalogue of the Hedwig-Schwägrichen Herbarium (Part 1 )” (PRICE, 2005). The records are listed alphabetically by basionym with the corresponding G barcode and figure number from PRICE (2005).
Appendix 1. — Cont.
Appendix 1. — Cont.
Appendix 1. — Cont.
