Faqi Zhang,^1,* Yinhu Li,^1,2,* Qingbo Gao,¹ Shuyun Lei,^1,2 Gulzar Khan,¹ Huiling Yang,¹ Shilong Chen<sup>1,**

1</sup>2Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology
²3University of the Chinese Academy of Sciences, Beijing 100039, People's Republic of China

^* These authors contributed equally to this work.
^** Author for correspondence: slchen@nwipb.cas.cn