Spiders and beetles were pitfall-trapped in the foreland of the receding Hardangerjøkulen glacier in central south Norway. At each of six sampling sites, ages 3 to 205 years, twenty traps covered the local variation in moisture and plant communities. Thirty-three spider species and forty beetle species were collected. The species composition was correlated to time since glaciation and vegetation cover. A characteristic pioneer community of spiders and mainly predatory beetles had several open-ground species, and some species or genera were common to forelands in Svalbard or the Alps. While the number of spider species increased relatively constant with age, the number of beetle species seemed to level off after about 80 years. Half of the beetle species were Staphylinidae, and contrary to Carabidae, most of these were rather late colonizers. Most herbivore beetles colonized after more than 40 years, but the moss-eating Byrrhidae species Simplocaria metallica and also certain Chironomidae larvae developed in pioneer moss colonies after 4 years. The large Collembola Bourletiella hortensis, a potential prey, fed on in-blown moss fragments after 3 years. In the present foreland, chlorophyll-based food chains may start very early. Two pioneer Amara species (Carabidae) could probably feed partly on seeds, either in-blown or produced by scattered pioneer grasses.

Our knowledge of stem secondary growth of arctic shrubs (a key component of tundra net primary production, NPP) is very limited. Here, we investigated the impact of the physical elements of the environment on shrub secondary growth by comparing annual growth rates of model species from similar habitats at contrasting altitude, microtopography, latitude, geographical location, and soil type, in both the sub- and High Arctic. We found that secondary growth has a modest sensitivity to the environment but with large differences among species. For example, the evergreen Cassiope tetragona is affected by altitude, microtopography, and latitude, whereas the evergreen Empetrum hermaphroditum has rather constant secondary growth in all environments. Deciduous species seem to be most affected by microtopography. Furthermore, the impact of the environment on secondary growth differed from the impact on primary growth (stem apical growth, stem length, and apical growth of stem plus leaves), in some cases even with opposite responses. Thus caution should be taken when estimating the impact of the environment on shrub growth from apical growth only. Integration of our data set with the (very limited) previously published information on secondary growth provides an overview of its contribution to NPP and annual growth rates for 9 arctic species at 18 sites in Sweden, Greenland, Svalbard, Alaska, and the Alps.

Six open-canopy high-elevation northern red oak (Quercus rubra L.) ring-width records were evaluated along the Southern Appalachian mountain range for a common climate-driven growth signal. Ring-width records show significant correlations over the past two centuries with principal component one (PC-1) accounting for 50% of the common variance. Spectral analysis reveals that the relative variance in ring-width is concentrated at decadal time scales. Ring-width records show positive correlations (p < 0.05) with prior fall–early winter and current summer mean temperature, and negative correlations (p < 0.05) with prior late summer–winter total precipitation. Yet, temperature-precipitation covariance along the Southern Appalachian mountain range during winter and summer seasons undergoes a significant reversal with intervening spring and fall seasons showing weak association. As a result, ring-width-climate signal strength exhibits time-dependence over the instrumental period, although the temporal change is not statistically significant. This temporal change suggests that both temperature and precipitation can have a marked influence on ring-width variability depending on the degree of seasonal covariance. Finally, obtaining a stable interannual climate-growth calibration for these particular ring-width records remains a foremost challenge, and is a primary consequence of mixed temperature-precipitation signal strength and little power at higher frequencies.

Instrumentation to study snowpack in situ was deployed in Lassen Volcanic National Park (LVNP), California, in an area of deep seasonal snow accumulation and known snow algal bloom recurrence. Included in the instrumentation were 11 temperature sensors, evenly spaced up to 2 m above the ground, which provided (1) temperature data within the snowpack when buried, and (2) estimates of snowpack height during accumulation and ablation periods. Beginning in April, moisture sensors measured a strong increase of snowpack liquid water content to greater than 15% by volume; this high melt content is usually coincident with the start of runoff from the snowpack. Snow depth profiles showed a rapid ablation of the final 2 m of the snowpack over about 23 days beginning in late June. SNTHERM numerical modeling confirmed that solar radiation was the dominant energy term throughout the melt season. By modeling a variety of snowpack parameters, such as albedo and initial snow density, we determined that the date of snow loss is the most sensitive observable that can be used to constrain the modeled parameters. These data sets from LVNP can also be applied to knowledge of snow algae lifecycles in deep snow to help understand whether the availability of light, water, or both controls the onset of snow algae germination at the base of a thick snowpack. Data and modeling indicate that meltwater was present throughout the snowpack beginning in March and runoff is initiated in April, when the snowpack was still several meters deep. However, significant levels of light did not penetrate to the soil until June, when the snow was less than 2 m deep.

Natural water level fluctuations (WLFs) are an inherent characteristic of Mediterranean inland waters, which are projected to be amplified by global climate change. La Caldera (Sierra Nevada National Park, Spain) is an oligotrophic high mountain lake (3050 m a.s.l.) that has experienced large fluctuations in water volume (13–100%) during the past 20 years due to irregular annual precipitation patterns (371–1816 mm). Because of the lake's cold and dilute abiotic environment, it is likely susceptible to projected increases in global temperature and represents an ideal sentinel of global change. We analyze the effect of WLFs on water quality and on plankton community in La Caldera to better understand the potential effects of recurrent droughts (3 droughts in a 20-year period) on lake ecology. We have found significantly positive effects of WLFs on total phosphorus (TP) concentrations. There was extreme variability in TP concentrations during three recurrent droughts (1995, 1999, and 2005) reflecting sediment resuspension. However, the data also suggest that this was not the only source of phosphorus. Extremely high P-enriched atmospheric dust inputs could have maintained the abnormally high TP in-lake concentrations measured during 2005. The data indicate that recurrent droughts have reduced lake resistance to TP changes but have increased lake resistance to total nitrogen (TN) changes, which supports the idea that a P-enriched atmospheric dust inputs during 2005. An increase in dissolved inorganic nitrogen (DIN)∶TP mass ratio after 2005 was observed, revealing a higher ecosystem homeostasis of this ratio.

Low temperature is considered the main limiting factor for plant growth and nutrient supply at high elevations. It has been repeatedly reported that an increase in foliar nutrient contents occurs with elevation which is interpreted as the plants' inability to use the absorbed resources for growth. However, although large data sets from various mountainous regions are available, data from elevations exceeding 5000 m elevation are rare, leaving uncertainties on the relevance of these patterns under extreme alpine conditions. To fill this gap, we examined foliar macronutrients (N, P, K, Ca, Mg) content in Poa attenuata and Waldheimia tridactylites along an elevation gradient of 1600 m in Ladakh, northwestern Himalaya. Species showed rather similar response: N, Ca, and Mg concentration decreased with elevation. However, P concentration decreased with elevation in Poa but slightly increased in Waldheimia; K concentration was related to elevation in Poa only (positively). The surprising decreases of N, and/or N/P and N/K ratios towards higher elevation suggest that nitrogen uptake decreased with elevation and it may limit plant growth. Our results suggest that plants growing at very high elevations tend to be limited by a combination of lower nutrient uptake, possibly because of poorly developed soils, and scarcity of water.

Alpine ecosystems are generally nitrogen (N) limited with low rates of N mineralization. Herbivory may affect N cycling and N losses and thus long-term productivity of ecosystems.

Using a controlled grazing experiment in a low-alpine region at Hol, southern Norway, with three density levels of sheep, we determined effects of grazing on in situ availability of inorganic N, potential N mineralization, and mobility of dissolved inorganic N (DIN) and dissolved organic N (DON) in soil water of O-horizons in grazing-preferred grassland habitats. In addition, we studied the within-season and spatial variation of these processes.

The low alpine grasslands at Hol were characterized by small rates of N mineralization and relatively large plant demands for N. Significantly greater rates of potential N mineralization were found at sites with high sheep density compared to those with low density or no grazing. Effects of grazing on bioavailable N (as determined by buried PRS™ exchange resins) were greater at low as compared to high altitudes. At low altitudes, low sheep density reduced amounts of bioavailable N. Nitrogen concentration of plants as a proxy of N availability in soils revealed, however, no significant effects of grazing. There was a strong seasonal effect on inorganic N and DIN∶DON ratios of the soil water, with decreasing values in the course of the growing season, probably due to increasing nutrient demand of plants and/or microbes.

We conclude that grazing may significantly stimulate N-cycling, but not sufficiently to release the system from its strong N deficiency, as we found no evidence for short-term increased risk in N loss via soil water due to herbivore activity. Nitrogen removal through grazing is small compared to the total soil N pool and at high sheep density is about half of the N deposition. This suggests that grazing in grassland habitats in this low alpine ecosystem is sustainable from a nutrient point of view.

This study focused on simulated glacier surface conditions (simulated Surface Melt and liquid Precipitation available for supra-, en-, sub-, and proglacial flow processes [after vertical percolation and potential storage within the snowpack] [henceforth SMP]), internal water storage and release, and runoff from the Kangerlussuaq drainage area of the Greenland Ice Sheet (GrIS), West Greenland, for the period 2006/2007 to 2007/2008. GrIS winter accumulation and summer ablation processes, including SMP, was simulated on both daily and hourly time steps. Using hourly meteorological driving data produced more realistic meteorological conditions instead of daily-averaged data, in relation to snow and melt threshold surface processes, and produced 9–17% higher annual cumulative SMP. The difference between simulated SMP and observed catchment runoff showed a decreasing lag time through the summer, and a drainage system storage buildup through approximately June and early July of up to 0.29 × 10⁹ m³, and a storage release through approximately late July and August of up to 0.25 × 10⁹ m³. The simulated total Kangerlussuaq SMP for 2006/2007 and 2007/2008, indicated a reduction of 30%. This reduction in SMP occurred simultaneously with the reduction in the overall pattern of satellite-derived GrIS surface melt from 2007 to 2008.

Physical, chemical and thermal properties of five pedons in the Uinta Mountains of northeastern Utah were studied to determine how soils and microclimates vary across the ecotone between subalpine forest and dry meadow. All five profiles have developed in the same parent materials, from base upward: basal till, glaciofluvial sediment, and coarse supraglacial debris with infilled eolian silt. Temperatures were measured at the soil surface and at depths of 10 and 50 cm for 420 days. Meadow soils (Typic Humicryepts) are warmer, have Bw horizons, and contain more Ca, organic carbon, and clay in the A horizon. Forest soils (Inceptic Haplocryalfs) have Bt horizons, higher B/A horizon clay ratios, are more acidic, and contain more exchangeable Mg and total exchangeable cations than meadow soils. Although subzero temperatures were never recorded at the surface of the forest soil in the summers of 1998 and 1999, those at the surface of the meadow soil fell below 0 °C on ∼35% of the nights. Mean 4:00 a.m. temperatures in the meadow during the summer are significantly colder than those in the forest (2.2 °C vs. 5.2 °C). The meadow formed following an initial forest clearing event, possibly fire or an insect infestation. Since that time, forest encroachment has been slow because seedling establishment in the meadow is inhibited by a combination of frequent summer freezing events, moisture stresses resulting from textural discontinuities between soil parent materials, and competition between tree seedlings and meadow vegetation.

This study reports changes of small glaciers in the Trans-Himalayan Kang Yatze Massif, Ladakh, northwest India, between 1969 and 2010. The region covers an area of about 1000 km² and is located in a transitional position between predominantly receding glaciers of the Central Himalaya and some advancing ice masses of the Karakorum. A multi-temporal remote sensing approach based on satellite images (Corona, SPOT, Landsat) was used to detect and analyze area changes of 121 small glaciers and to measure the retreat of 60 cirque and valley glaciers between 1969 and 2010. Over the last four decades, the glaciated area decreased by about 14% (0.3% yr⁻¹) from 96.4 to 82.6 km² and the average ice front retreat amounts to 125 m (3 m yr⁻¹). The ice cover loss shows a high decadal variability with the maximum shrinkage between 1991 and 2002 (0.6% yr⁻¹), followed by a lower decrease rate since then (0.2% yr⁻¹). Due to the high variability of glacier change with a generally decreasing trend and a few stable glaciers, it becomes obvious that an extrapolation even on a regional scale is problematic. Therefore, a consideration of differing responses of various glacier types and glacier sizes is of utmost importance.

Since 26 September 2005 an Automatic Weather Station (AWS1 Forni) has been running on the ablation area of the largest Italian valley glacier, Forni, in the Ortles–Cevedale Group. A 4-year record (from 1 October 2005 to 30 September 2009) of air temperature, relative humidity, wind speed and direction, incoming and outgoing radiative fluxes, air pressure, liquid precipitation, and snow depth is considered. The meteorological data are analyzed to describe glacier surface conditions, to calculate the energy balance, and to evaluate the ice ablation amount. Snow accumulation was measured, thus permitting the estimation of the glacier point mass balance.

An annual average amount of melt of −5.4 ± 0.021 m w.e. was calculated and an annual average amount of accumulation of 0.7 ± 0.006 m w.e. was measured at the AWS site. The annual average amount of mass balance was −4.7 ± 0.023 m w.e.

Our analyses show that surface conditions during summer and fall seasons are important in regulating glacier albedo and then mass balance. In particular, snow cover presence, due to a longer persistence of spring snow, summer snowfalls and earlier fall solid precipitation, drives the duration of the ice melt period.

Modified environmental conditions are driving phenological changes in ecosystems around the world. Many plants have already responded to warmer temperatures by flowering earlier and sustaining longer periods of growth. Changes in other environmental factors, like precipitation and atmospheric nitrogen (N) deposition, may also influence phenology but have been less studied. Alpine plants may be good predictors of phenological response patterns because environmental changes are amplified in mountain ecosystems and extreme conditions may make alpine plants particularly sensitive to changes in limiting factors like precipitation, temperature, and N. We tested the effects of increased snowpack, temperature, and N on alpine tundra plant phenology, using snow fence, open-top warming chamber, and N fertilization treatments at the Niwot Ridge Long Term Ecological Research (LTER) site. Flowering phenology of three abundant species was recorded during two growing seasons. Treatment responses varied among species and functional types. Forbs responded to warming by flowering earlier and responded to snowpack and N by flowering later; however, when both snow and N were increased simultaneously, phenology was unchanged. Graminoids flowered earlier in response to N addition. Our results demonstrate that changing environmental conditions influence plant phenology, and specifically highlight that N and multiple factor interactions can yield stronger responses than warming alone.

The importance of fog precipitation in the alpine hydrological processes of Pinus pumila canopy was evaluated on Mt. Tateyama, central Japan. We observed rain and fog precipitation, throughfall, and wind direction and velocity at Jodo-daira (36.566°N, 137.606°E, 2840 m a.s.l.) for 3 years. During the snow-free period (August and September), mean monthly rain and fog precipitation was 0.45 mm h⁻¹ and 0.14 mm h⁻¹, respectively. The mean rainfall interception by P. pumila canopy was about 48%, which is higher than that of other forest canopies at lower altitudes. During rainfall, the dense canopy intercepts rain and the water evaporates from the needle surfaces. On the other hand, the canopy captured fog precipitation even in the absence of rainfall. The amount of throughfall increased with increasing fog deposition. Using δ¹⁸O and δD analysis, the mean contribution of fog water to the throughfall was estimated at approximately 35%, consistent with the result from direct measurement. These results indicate that P. pumila should have a significant influence on the local hydrological processes of the high mountain ecosystem. The large contribution of fog precipitation can be attributed to the high wind velocity and humidity of the Japan Alps.