This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies.
EUAESTHETINAE
[26 genera; 724 species; All Regions except Antarctic]
Euaesthetinae Thomson, 1859.
— Eichelbaum, 1909: 130 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).
— Blatchley, 1910: 458 (characters; key to Indiana genera).
— Bernhauer and Schubert, 1911: 186 (world catalog).
— Cameron, 1921a (characters; key to Singapore species and genera).
— Porta, 1926: 64 (characters; species of Italy).
— Portevin, 1929: 389 (characters; key to French genera).
— Cameron, 1930: 444 (characters; key to genera of British India).
— Scheerpeltz, 1933: 1202 (world catalog supplement).
— C. Koch, 1934: 66 (key to tribes).
— Scheerpeltz, 1940: 31 (key to Palaearctic genera).
— Blackwelder, 1943: 226 (characters; notes).
— Blackwelder, 1944: 115 (checklist of species from Mexico, Central and South America, and the West Indies).
— Tottenham, 1954: 74 (characters; British species).
— Hatch, 1957: 48, 240 (characters).
— Palm, 1961a: 117 (characters; species of Sweden).
— Arnett, 1963: 243 (key to North American genera).
— Arnett, 1963: 265 (characters; list of North American genera).
— Horion, 1963: 400 (list of central European species).
— Székessy, 1963: 37 (characters; key to genera of Hungary).
— Lohse, 1964: 130 (characters; key to central European genera and species).
— Moore, 1964a: 88 (characters; notes).
— Kasule, 1966: 266 (larval characters; relationship to Steninae and Pselaphidae).
— Moore, 1967 (characters).
— Coiffait and Saiz, 1968: 395 (list of Chilean species).
— Scheerpeltz, 1968a: 46 (catalog of Austrian species).
— Saiz, 1972: 173 (key to genera of Chile).
— Moore and Legner, 1974g: 555 (key to genera of America north of Mexico).
— Scheerpeltz, 1974: 102 (key to genera).
— Moore and Legner, 1975: 261 (catalog; North America north of Mexico).
— Pope, 1977: 26 (list of British species).
— Puthz, 1978c: 127 (key to species of Australia).
— Topp, 1978: 307 (larval characters).
— Moore and Legner, 1979: 242 (characters; tabular key to genera of North America north of Mexico).
— Tóth, 1981: 99 (list of species of Hortobágy National Park, Hungary).
— Watt and McColl, 1982: 13, 20 (characters; notes on New Zealand taxa).
— Coiffait, 1984f: 353 (characters; key to tribes; keys to genera and species of western Palaearctic region).
— Newton, 1985: 205 (notes on austral distribution).
— Outerelo and Gamarra, 1985: 19 (key to genera of Iberian Peninsula).
— Orousset, 1988 (characters; key to tribe, genera and species of Madagascar; discussions of taxonomic position, wing reduction, depigmentation, eye reduction, endemism, and speciation in Madagascar; collecting techniques).
— Frank, 1991: 345 (larval characters).
— Newton and Thayer, 1992: 13, 56 (summary of current classification; discussion).
— Lawrence and Newton, 1995: 829 (notes).
— Terlutter, 1995: 54 (list of species collected in Westphalia).
— Sparacio, 1995: 149 (characters; species of Sicily).
— Klimaszewski, Newton, and Thayer, 1996: 150 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New Zealand species).
— M. Hansen, 1996: 37, 102 (listed in Staphylinine group of subfamilies; list of Danish species).
— Downie and Arnett, 1996: 459 (characters; key to genera of Northeastern North America).
— M. Hansen, 1997: 128, 175, 177, 179 (autapomorphic features; key adult and larval characters; notes).
— Anderson, Nash, and O’Connor, 1997: 17 (annotated checklist; Ireland).
— Newton, Thayer, Ashe, and Chandler, 2000: 323, 382 (key to Nearctic tribes and genera).
— Klimaszewski, 2000: 22, 61 (characters; notes on classification, natural history, distribution, and key to tribes in Canada).
Euaesthetina Thomson, 1859: 42 (genera included: Euaesthetus). Type genus: Euaesthetus (see Newton and Thayer, 1992: 56).
Tribe Alzadaesthetini
Alzadaesthetini Scheerpeltz, 1974: 102 (genera included: Alzadaesthetus). Type genus: Alzadaesthetus (see Newton and Thayer, 1992: 56).
Alzadaesthetus
[2 species; Neotropical Region]
Alzadaesthetus Kistner, 1961: 217 (species included: chilensis). Type species: Alzadaesthetus chilensis Kistner, fixed by original designation and monotypy.
— Coiffait and Saiz, 1968: 395 (Alzadaesthetus; list of Chilean species).
— Saiz, 1970a: 309 (characters).
— Saiz, 1972: 174 (characters; key to species).
— Scheerpeltz, 1974: 102 (characters).
— Newton, 1985: 205 (characters).
chilensis Kistner, 1961: 217 (Alzadaesthetus; Type locality: Chile: Puyehue, Rio Gol-Gol, 350 m).
furcillatus Saiz, 1972: 175 (Alzadaesthetus; Type locality: Chile: Cordillera Pelada, Valdivia).
Tribe Austroesthetini
Austroesthetini Cameron, 1944.
Austraesthetini Cameron, 1944b: 69 (spelling based on unjustified emendation according to Newton and Thayer, 1992: 56; [Note: I found no evidence in Cameron, 1944b to suggest that Cameron's alteration is an intended emendation rather than a misspelling]; genera included: Austroesthetus, cited as Austroaesthetus). Type genus: Austroesthetus (see Newton and Thayer, 1992: 56).
Austroesthethus
[4 species; Australian Region]
Austroesthethus Oke, 1933: 112 (species included: passerculus, gippsensis, punctatus). Type species: Austroesthethus passerculus Oke, by original designation.
— Blackwelder, 1952: 70 (type species: passerculus).
— Scheerpeltz, 1974: 103 (characters).
— Newton, 1985: 205 (characters).
gippsensis Oke, 1933: 114 (Austroesthethus; Type locality: Australia: Victoria: Pakenham; Traralgon).
passerculus Oke, 1933: 112 (Austroesthethus; Type locality: Australia: Victoria: Fern Tree Gull, Belgrave, Warburton: Mt. Donna Buang at 4000 feet; New South Wales: Mt Kiosciusko at 6500 feet).
punctatus Oke, 1933: 114 (Austroesthethus; Type locality: Australia: Victoria: Grampians).
tasmanicus Puthz, 1978c: 120 (Austroesthethus; Type locality: Tasmania: Parawee [Parrawe]).
Chilioesthetus
[3 species; Australian and Neotropical Regions]
Chilioesthetus Saiz, 1968: 73 (species included: lorenae, marcelae). Type species: Chilioesthetus lorenae Saiz, fixed by original designation.
ernstjuengeri Puthz, 1995: 111 (Chilioesthetus; Type locality: Australien: Queensland, Mt. Lewis, 1060 m, 16°34′S, 145°17′E).
lorenae Saiz, 1968: 74 (Chilioesthetus; Type locality: Chile: Cordillera Pelada, Valdivia).
marcelae Saiz, 1968: 76 (Chilioesthetus; Type locality: Chile: Cordillera Pelada, Valdivia).
Mesoaesthetus
[3 species; Australian Region]
Mesoaesthetus Cameron, 1944b: 68 (species included: wilsoni). Type species: Mesoaesthetus wilsoni Cameron, fixed by monotypy.
tasmanicus Puthz, 1978c: 118 (Mesoaesthetus; Type locality: Tasmania: Parawee [Parrawe]).
whitehorni Broun, 1912: 400 (Dimerus; Type locality: Retaruke, near Erua).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Bernhauer, 1939f: 214 (Agnosthaetus; characters in key).
— Newton, 1996: 159 (Mesoaesthetus; characters; notes).
▴ Distribution: New Zealand.
wilsoni Cameron, 1944b: 69 (Mesoaesthetus; Type locality: Australia: Warburton).
Nothoesthetus
[8 species; Neotropical Region]
Nothoesthetus Saiz, 1970a: 295 (cited as Notoesthetus at the head of the description and at the type species designation but as Nothoesthetus elsewhere; species included: coiffaiti, scitulus, australis, obesus, montanus). Type species: Nothoesthetus coiffaiti Saiz, fixed by original designation.
affinis Saiz, 1973: 178 (Nothoesthetus; Type locality: Chile: Bahía Mansa, Osorno).
australis Saiz, 1970a: 304 (Nothoesthetus; Type locality: Bahia Orange, Magallanes).
coiffaiti Saiz, 1970a: 299 (Nothoesthetus; Type locality: Chile: Palmas de Cocalan).
montanus Saiz, 1970a: 307 (Nothoesthetus; Type locality: Chile: Cordillera Pelada, Valdivia).
obesus Saiz, 1970a: 305 (Nothoesthetus; Type locality: Chile: Cordillera Pelada, Valdivia).
saizi Puthz, 1988g: 149 (Nothoesthetus; Type locality: Chile: Malleco Province: 15 km W. Victoria, 200 m).
scitulus Saiz, 1970a: 302 (Nothoesthetus; Type locality: Chile: Agua de la Gloria, Bulnes, Concepcion).
singularis Saiz, 1973: 179 (Nothoesthetus; Type locality: Chile: Camino Viejo a La Unión, 25 kms. al S. de Valdivia).
Tasmanosthetus
[1 species; Australian Region]
Tasmanosthetus Puthz, 1978c: 119 (species included: okei). Type species: Tasmanosthetus okei Puthz, fixed by original designation.
okei Puthz, 1978c: 119 (Tasmanosthetus; Type locality: Tasmania: Mt. Wellington).
Tribe Euaesthetini
Euaesthetini Thomson, 1859.
—Jakobson, 1908: 446, 478 (characters; list of species of Russia and western Europe).
— Bernhauer and Schubert, 1911: 187 (world catalog; Turellus, Octavius, Edaphus, Edaphellus, Tamotus, Euaesthetus, Ctenomastax).
— Johansen, 1914: 475 (characters).
— Scheerpeltz, 1933: 1202 (world catalog supplement).
— C. Koch, 1934: 67, 68 (characters in key; key to genera).
— Scheerpeltz, 1940: 31 (key to Palaearctic genera).
— Cameron, 1944b: 69 (characters).
— Arnett, 1963: 266 (characters; list of North American genera).
— Scheerpeltz, 1974: 104 (characters).
— Coiffait, 1984f: 353, 354 (characters; key to genera).
— Newton and Thayer, 1992: 56 (tribe of Euaesthetinae).
— M. Hansen, 1996: 102 (list of Danish species).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (characters; notes).
Tamotini Coiffait, 1984f: 353 (genera included: Tamotus). Type genus: Tamotus (see Newton and Thayer, 1992: 56).
Coiffaitia
[6 species; Madagascan Region]
Coiffaitia Kistner and Shower, 1965: 317 (species included: minor). Type species: Coiffaitia minor Kistner and Shower, fixed by original designation and monotypy.
—Scheerpeltz, 1974: 107 (characters).
— Orousset, 1988: 56 (characters; key to species of Madagascar).
manjarivolo Orousset, 1988: 67 (Coiffaitia; Type locality: Madagascar Centre: massif de l’Andringitra, base de la forêt du cirque de Manjarivolo, 1200 m).
marojejy Orousset, 1988: 67 (Coiffaitia; Type locality: Madagascar Est: massif du Marojejy, 600 m).
minor Kistner and Shower, 1965: 319 (Coiffaitia; Type locality: East Madagascar, Baie d’Antongil, Varingohitra).
peyrierasi Orousset, 1988: 65 (Coiffaitia; Type locality: Madagascar Centre: Est du lac de Mantasoa, forêt d’Ambohiboatavo, 1340 m).
spectabilis Orousset, 1988: 68 (Coiffaitia; Type locality: Madagascar Est: environs de Maroantsetra, Fampanambo).
vadoni Orousset, 1988: 68 (Coiffaitia; Type locality: Madagascar Est: presqu’île de Masoala, Ambohitsitondroina, 1100 m).
Ctenomastax
[7 species; Palaearctic and Ethiopian Regions]
Ctenomastax Kraatz, 1870b: 84 (species included: kiesenwetteri). Type species: Ctenomastax kiesenwetteri Kraatz, fixed by monotypy.
—Fauvel, 1873: 133 [= 1873a: 291] (characters).
— Duvivier, 1883: 177 (catalog; cited as Ctenotomax).
— Heyden, Reitter, and Weise, 1906: 151 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1911: 190 (world catalog; 3 species).
— Winkler, 1925: 358 (catalog for Palaearctic region).
— Scheerpeltz, 1933: 1206 (world catalog supplement).
— Normand, 1935: 364 (list of species of Tunisia).
— Blackwelder, 1952: 114 (type species: kiesenwetteri).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Scheerpeltz, 1974: 106 (characters).
— Coiffait, 1984f: 363 (characters; key to species of western Palaearctic region).
— Puthz, 1988e: 51 (characters; notes; species complexes; key to species).
— Puthz, 1989f: 70 (map showing distribution for genus).
— Orousset, 1990b: 131 (characters; notes).
asiaticus Puthz, 1988e: 57, 59 (Ctenomastax; Type locality: Pakistan: Umg. Rawalpindi, Dhok Pathan, Sohan R., bei Pindi Gheb).
hummleri Bernhauer, 1913a: 221 (Ctenomastax; Type locality: Kephallenia).
— Scheerpeltz, 1933: 1206 (Ctenomastax; catalog).
— Coiffait, 1984f: 366 (Ctenomastax; characters; Ionian Islands).
— Puthz, 1988e: 57 (Ctenomastax; cited as though synonym of kiesenwetteri but not formally synonymized).
▴ Distribution: Ionian Islands.
kiesenwetteri Kraatz, 1870b: 85 (Ctenomastax; Type locality: Umgebungen von Sevilla).
— Fauvel, 1873: 133 [= 1873a: 291] (Ctenomastax; characters; Spain).
— Fauvel, 1900: 58 (Ctenomastax; characters; Algeria; Tunisia; Andalousie).
— Bernhauer and Schubert, 1911: 190 (Ctenomastax; catalog).
— Scheerpeltz, 1933: 1206 (Ctenomastax; catalog).
— Coiffait, 1984f: 365 (Ctenomastax; characters; Spain; Algeria; Tunisia).
— Puthz, 1988e: 57 (Ctenomastax; characters; Spain; Algeria; Cephallonia; Tunisia; Cyprus).
— Orousset, 1990b: 133 (Ctenomastax; Portugal; Algeria; Tunisia).
▴ Distribution: Portugal, Spain, Algeria, Tunisia, Ionian Islands, Cyprus.
nesterovi Petrenko, 1984: 46 (Euaesthetus; Type locality: Tadzhikistan: okr. Dushanbe).
mirei Orousset, 1990b: 132 (Ctenomastax; Type locality: Maroc: Zagora. Also cited from Niger).
microthorax C. Koch, 1936, see: pharaonum J. Sahlberg, 1908.
nesterovi Petrenko, 1984, see: kiesenwetteri Kraatz, 1870.
pharaonum J. Sahlberg, 1908: 32 (Ctenomastax; Type locality: Ghizeenses prope urbem Cairo).
— Bernhauer and Schubert, 1911: 190 (Ctenomastax; catalog).
— Coiffait, 1984f: 365 (Ctenomastax; characters; Egypt).
— Puthz, 1988e: 57, 58 (Ctenomastax; characters; Egypt).
▴ Distribution: Egypt.
microthorax C. Koch, 1936b: 149 (Ctenomastax; subspecies of pharaonum; Type locality: Nord Sinai: El Kontella).
—Puthz, 1988e: 58 (Ctenomastax; synonym of pharaonum).
— Orousset, 1990b: 131 (Ctenomastax; synonym of pharaonum).
stocki Bernhauer, 1940, see: varicolor Fauvel, 1900.
stussineri Bernhauer, 1913s: 221 (Ctenomastax; Type locality: Syrien: Haifa).
—Scheerpeltz, 1933: 1206 (Ctenomastax; catalog).
— Coiffait, 1984f: 366 (Ctenomastax; characters; Israel).
— Puthz, 1988e: 57 (Ctenomastax; cited as though synonym of kiesenwetteri but not formally synonymized).
— Orousset, 1990b: 131 (Ctenomastax; possible synonym of kiesenwetteri).
▴ Distribution: Israel.
varicolor Fauvel, 1900: 57 (Ctenomastax; Type locality: Andalousie: Archena près Murcie; Tunisie: environs de Mactar).
— Bernhauer and Schubert, 1911: 190 (Ctenomastax; catalog).
— Coiffait, 1984f: 365 (Ctenomastax; characters; Spain; Tunisia).
— Puthz, 1988e: 57, 58 (Ctenomastax; characters; Spain; Tunisia).
— Orousset, 1990b: 131, 133 (Ctenomastax; possible synonym of kiesenwetteri; Tunisia).
▴ Distribution: Spain, Tunisia.
stocki Bernhauer, 1940b: 633 (Ctenomastax; Type locality: Tunis: Le Kef).
Edaphus
[345 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]
Edaphus Motschulsky, 1857: 7 (species included: nitidus, politus). Type species: Edaphus nitidus Motschulsky, fixed by subsequent designation by R. Lucas, 1920: 256, and cited as E. nitidus J. Lec.”
— Gemminger and Harold, 1868: 632 (catalog).
— Mulsant and Rey, 1878a: 301 (characters).
— Porta, 1926: 65 (characters; key to species of Italy).
— Blackwelder, 1952: 144 (type species).
— Székessy, 1963: 38 (characters).
— Puthz, 1973k: 57 (key to Neotropical species).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Scheerpeltz, 1974: 106 (characters).
— Puthz, 1974t: 911 (characters; fixation of type species: Edaphus nitidus and discussion of previous designations; notes; key to Nearctic species).
— Puthz, 1975c: 170 (key to species of Palaearctic region; notes).
— Moore and Legner, 1975: 261 (catalog; North America north of Mexico).
— Shibata, 1976: 210 (checklist of species of Japan).
— Puthz, 1979: 107, 142 (characters; anatomy; species groups classification; discussions; key to species of India, Sri Lanka, and Indochina).
— Moore and Legner, 1979: 243 (characters; notes).
— E. Matthews, 1982: 8 (notes; Australia).
— Hammond, 1984: 202 (checklist of species of Borneo).
— Frank, 1986a: 367 (checklist of species of Florida).
— Shibata, 1986a: 121 (supplemental list of species from Taiwan).
— Orousset, 1988: 28 (characters; key to species of Madagascar; notes on world distribution and habitat).
— Puthz, 1990c: 823 (key to species of Sumatra).
— Puthz, 1991: 292 (key to species of Cook and Fiji Islands).
— Silfverberg, 1992: 19 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Zanetti, 1995: 19 (list of species of Italy).
— Lundgren, 1998: 48 (list of species from Florida; records for other States listed for each species).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (4 Nearctic species; notes; characters in key).
Edaphus LeConte, 1861a: 67 (species included: nitidus; [Note: This name has been cited as having been originated by LeConte (Bernhauer and Schubert, 1911: 187) or as a subsequent use of Motschulsky's name (Puthz, 1974). LeConte (1861a) did not refer to Motschulsky's use of the name and later (1863) specifically referred the name to himself]). Type species: Edaphus nitidus LeConte, fixed by monotypy.
—LeConte, 1863: 25 (list of North American species).
— Fauvel, 1873: 60 [= 1873a: 218] (characters).
— Fauvel, 1875a: XIV [= 1875b: 216] (catalog).
— Duvivier, 1883: 179 (catalog).
— Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 543 (characters).
— Heyden, Reitter, and Weise, 1906: 151 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1911: 187 (world catalog; 15 species).
— Leng, 1920: 100 (catalog of North American species).
— Cameron, 1921a: 352, 401 (characters; key to Singapore species; catalog).
— Winkler, 1925: 357 (catalog for Palaearctic region).
— Cameron, 1925: 31 (catalog of Indian species).
— Portevin, 1929: 390 (characters).
— Cameron, 1930: 447 (characters; key to species of British India).
— Cameron, 1930d: 330 (key to species of Sumatra).
— Scheerpeltz, 1933: 1203 (world catalog supplement).
— Scheerpeltz, 1936 (distribution and key to western Palaearctic species).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 144 (synonym of Edaphus Motschulsky).
— Kistner, 1962a: 619 (characters; key to Ethiopia and Nearctic species).
— Horion, 1963: 400 (list of central European species).
— Lohse, 1964: 130 (characters; key to central European species).
— Scheerpeltz, 1968a: 46 (catalog of Austrian species).
— Puthz, 1974t: 911 (cited as subsequent reference of Motschulsky name).
— Coiffait, 1984f: 357 (characters; key to species of western Palaearctic region).
— Puthz, 1992b: 17, 30 (key to species of franzi group; catalog of African species).
— Downie and Arnett, 1996: 460 (key to species of Northeastern North America).
— Herman, 2001: 19 (junior synonymic homonym of Edaphus Motschulsky).
Tetratarsus Schaufuss, 1877: 24 (species included: plicatulus). Type species: Tetratarsus plicatulus Schaufuss, fixed by monotypy.
—Schaufuss, 1877a: 460 (characters).
— Ganglbauer, 1895: 543 (synonym of Edaphus).
— Bernhauer and Schubert, 1911: 187 (synonym of Edaphus).
— Blackwelder, 1952: 383 (type species: plicatulus).
— Kistner, 1962a: 619 (synonym of Edaphus).
— Scheerpeltz, 1974: 106 (synonym of Edaphus).
— Puthz, 1974t: 911 (synonym of Edaphus).
— Coiffait, 1984f: 356 (synonym of Edaphus).
— Orousset, 1988: 29 (synonym of Edaphus).
Tetrameres Schaufuss, 1877a: 460 (emendation of Tetratarsus). Type species: Tetratarsus plicatulus Schaufuss, fixed by objective synonymy with Tetratarsus.
—Blackwelder, 1952: 382 (type species).
— Puthz, 1974t: 911 (synonym of Edaphus).
— Orousset, 1988: 29 (synonym of Edaphus).
Edaphellus Fauvel, 1878f: 220 (species included: novaeguineae). Type species: Edaphellus novaeguineae Fauvel, fixed by monotypy.
—Duvivier, 1883: 179 (catalog).
— Bernhauer and Schubert, 1911: 188 (world catalog; 1 species).
— Skékessy, 1939: 98 (synonym of Edaphus by transfer of type species).
— Blackwelder, 1952: 144 (type species: novaeguineae).
— Scheerpeltz, 1974: 106 (synonym of Edaphus).
— Puthz, 1974t: 911 (synonym of Edaphus).
— Orousset, 1988: 29 (synonym of Edaphus).
Microphthartus Blattný, 1925: 185 (species included: luridus). Type species: Microphthartus luridus Blattný, fixed by monotypy.
Rhenanus Wüsthoff, 1935: 48 (species included: rosskotheni). Type species: Rhenanus rosskotheni Wüsthoff, fixed by monotypy.
—Holdhaus, 1936: 10 (synonym of Edaphus).
— Blackwelder, 1952: 339 (type species: rosskotheni).
— Puthz, 1974t: 912 (synonym of Edaphus).
— Coiffait, 1984f: 356 (synonym of Edaphus).
— Orousset, 1988: 29 (synonym of Edaphus).
abdominalis Puthz, 1979b: 171 (Edaphus; Type locality: W. Sarawak: Mt. Matang).
abessinicus Puthz, 1990d: 40 (Edaphus; Type locality: Äthiopien: Addis Abeba, Gullalè).
acuticollis Puthz, 1976f: 157 (Edaphus; Type locality: Ceylon: Uva, Monaragala).
adazla Kistner, 1961: 35 (Edaphus; Type locality: Congo: Parc National de la Garamaba).
—Kistner, 1962a: 621 (Edaphus; characters).
— Puthz, 1974s: 754 (Edaphus; characters; Ghana).
— Puthz, 1983f: 173 (Edaphus; Zaire).
— Puthz, 1984g: 196 (Edaphus; characters; Ghana).
— Puthz, 1990a: 203 (Edaphus; notes; Ivory Coast).
▴ Distribution: Congo, Ghana, Ivory Coast.
endrodyi Jacobson and Kistner, 1974: 87 (Edaphus; Type locality: Ghana: Ashanti Region, Kwadaso).
flavescens Jacobson and Kistner, 1974: 89 (Edaphus; Type locality: Ghana: Brong-Ahafo Reg., Kintampo Catarract, Fullner Falls).
adiposus Puthz, 1979: 127 (Edaphus; Type locality: Thailand: Sakaerat Experimental Station).
aequifrons Bernhauer, 1929c: 95, 103 (Edaphus; Type locality: Süd-Luzon: Mt. Isarog).
africanus Eppelsheim, 1885: 139 (Edaphus; Type locality: Goldküste: Adda).
—Bernhauer and Schubert, 1911: 187 (Edaphus; catalog).
— Bernhauer, 1922b: 169 (Edaphus; Seychelles).
— Scheerpeltz, 1933: 1203 (Edaphus; catalog).
— Kistner, 1960: 31 (Edaphus; Ethiopia).
— Kistner, 1962a: 632 (Edaphus; incertae sedis).
— Puthz, 1974s: 752 (Edaphus; characters).
— Puthz, 1983f: 172 (Edaphus; Zaire; Guinea; Togo; Cameroon).
— Puthz, 1984g: 196 (Edaphus; characters; Angola; South Africa; Ghana).
— Orousset, 1988: 37 (Edaphus; characters; Seychelles).
— Puthz, 1989a: 74 (Edaphus; South Africa).
▴ Distribution: Angola, Cameroon, Congo, Ethiopia, Ghana, Guinea, Seychelles, South Africa, Togo, Zimbabwe.
marshalli Bernhauer, 1934h: 501 (Edaphus; Type locality: Mashonaland: Salisbury; [Note: See lectotype designation by Orousset, 1986a]).
—Kistner, 1962a: 628 (Edaphus; characters; Southern Rhodesia).
— Puthz, 1984g: 196 (Edaphus; synonym of africanus).
— Orousset, 1986a: 170 (Edaphus; lectotype designation).
— Orousset, 1988: 37 (Edaphus; synonym of africanus).
— Puthz, 1992b: 30 (Edaphus; synonym of africanus).
marlieri Kistner, 1962a: 628 (Edaphus; Type locality: Congo Republic: Kivu, Uvira territory, Kavimvira).
—Kistner, 1963: 199 (Edaphus; Congo Republic).
— Puthz, 1974s: 760 (Edaphus; characters).
— Puthz, 1984g: 196 (Edaphus; synonym of africanus).
— Orousset, 1988: 37 (Edaphus; synonym of africanus).
— Puthz, 1992b: 30 (Edaphus; synonym of africanus).
sexpunctus Kistner, 1962: 106 (Edaphus; Type locality: Angola: Dundo).
—Kistner, 1962a: 629 (Edaphus; characters; Congo Republic).
— Kistner, 1963: 199 (Edaphus; Congo Republic).
— Puthz, 1974s: 761 (Edaphus; characters).
— Puthz, 1984g: 196 (Edaphus; synonym of africanus).
— Orousset, 1988: 37 (Edaphus; synonym of africanus).
— Puthz, 1992b: 30 (Edaphus; synonym of africanus).
alutipennis Bernhauer, 1929c: 96, 109 (Edaphus; Type locality: Luzon: Mt. Polis).
americanus Puthz, 1974: 915, 920 (Edaphus; Type locality: Kentucky: Edmonson Co.: Mammoth Cave Nt. Pk., Bruce Hollow. Also cited from Pennsylvania, Ohio, Indiana, Illinois, Missouri, Virginia, Michigan, North Carolina, South Carolina, Georgia, Florida, Alabama, Louisiana).
angolensis Cameron, 1950f: 119 (Edaphus; Type locality: Angola: Dundo: River Chiumbe; [Note: See lectotype designation by Puthz, 1984g]).
—Kistner, 1962: 104 (Edaphus; characters; Angola).
— Kistner, 1962a: 621 (Edaphus; characters; Angola).
— Puthz, 1984g: 200 (Edaphus; lectotype designation: Angola: Valley of R. Chiumbe, 45 km E Dundo; characters; Angola; Zaire).
▴ Distribution: Angola, Congo.
angulaticollis Puthz, 1986c: 312 (Edaphus; Type locality: Gabun: Belinga).
angulicollis Székessy, 1939: 103 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay).
angulipennis Puthz, 1973k: 56 (Edaphus; Type locality: Brazil: São Paulo, Alto de Serra).
angustulus Puthz, 1990a: 205 (Edaphus; Type locality: Côte d’Ivoire: Dropleu).
angustus Székessy, 1939: 101 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay).
annamensis Puthz, 1979: 130 (Edaphus; Type locality: Vietnam: Cuc phuong, Ninh binh).
antennatus Székessy, 1939: 98 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen und Stephansort, Astrolab-Bay).
armiger Orousset, 1988: 40 (Edaphus; Type locality: Madagascar Est: route de Tamatave, km 181, Beforona, 540 m).
aschnaae Makhan, 1995, see: humeralis Puthz, 1973.
australicus Puthz, 1978c: 126 (Edaphus; Type locality: Queensland: Winescrub forest near Maipoton).
badiorufus Orousset, 1988: 43 (Edaphus; Type locality: Madagascar Nord: Montagne d’Ambre, environs de la station forestière).
bakeri Bernhauer, 1929c: 96, 106 (Edaphus; Type locality: Luzon: Mt. Makiling).
bakerianus Bernhauer, 1929, see: princeps Bernhauer, 1929.
bambousi Kistner, 1962a: 622 (Edaphus; Type locality: Congo Republic: Kivu: Contref. S. of Kahuzi near Kalehe, 2200 m).
basilewskyi Kistner, 1960: 38 (Edaphus; Type locality: Kenya: Elgon, east slope, 2500 m).
— Kistner, 1962a: 622 (Edaphus; characters).
— Puthz, 1974s: 756 (Edaphus; characters).
▴ Distribution: Kenya.
batrisoides Comellini, 1977, see: remyi Jarrige, 1963.
benicki Puthz, 1975c: 178 (Edaphus; Type locality: China: West-Szechuan, Wassuland, Bezirk San-kiang-kou, Sankiang, 1600 m).
bernhaueri Székessy, 1939: 101 (Edaphus; Type locality: Neu-Guinea: Montes Hansemann).
besucheti Comellini, 1977: 258 (Edaphus; Type locality: Sri Lanka: province Northern, Mullaittivu).
beszedesi Reitter, 1914: 189 (Edaphus; Type locality: Umgebung von Abazia, Istrien).
— Scheerpeltz, 1933: 1203 (Edaphus; cited as beczedesi; catalog).
— Scheerpeltz, 1936: 202, 219 (Edaphus; characters; Istria).
— Horion, 1963: 401 (Edaphus; Istria; Yugoslavia; Hercegovina; Montenegro; Austria).
— Lohse, 1964: 130 (Edaphus; characters; central Europe).
— Puthz, 1974t: 913, 923 (Edaphus; characters; notes; Illinois; Kansas).
— Puthz, 1975c: 174, 176 (Edaphus; characters; France; Germany; Switzerland; Italy; Austria; Hungary; Czechoslovakia; Yugoslavia; Bulgaria; Asia Minor; Japan; North America).
— Puthz, 1983g: 163 (Edaphus; notes; China).
— Coiffait, 1984f: 358 (Edaphus; characters; France; central Europe; Balkans; Turkey; Japan; USA).
— Lohse and Lucht, 1989: 149 (Edaphus; synonym of beszedesi).
— Lohse and Lucht, 1989: 149 (Edaphus; Holarctic).
— Kuschel, 1990: 26 (Edaphus; cited as beczedesi; introduced, Palaearctic species; winged; habitat; New Zealand).
— J. Li, 1993: 27 (Edaphus; China).
— Terlutter, 1995: 54 (Edaphus; Germany).
— Zanetti, 1995: 19 (Edaphus; Italy).
— Downie and Arnett, 1996: 461 (Edaphus; characters; USA).
▴ Distribution: Europe, Turkey, China, Japan, New Zealand, USA.
piceus Cameron, 1933a: 169 (Edaphus; Type locality: Japan: Kobe).
—Puthz, 1975c: 176 (Edaphus; synonym of beszedesi).
— Coiffait, 1984f: 358 (Edaphus; synonym of beszedesi).
rosskotheni Wüsthoff, 1935: 48 (Rhenanus; Type locality: Nordabhang des Lousberges).
—Holdhaus, 1936: 10 (Edaphus; synonym of beszedesi).
— Scheerpeltz, 1936: 202, 215 (Edaphus; characters; Germany).
— Horion, 1963: 401 (Edaphus; Germany).
— Lohse, 1964: 130 (Edaphus; synonym of bluehweissi).
— Puthz, 1971p: 202 (Edaphus; cited as valid species; notes).
— Puthz, 1974t: 923 (Edaphus; synonym of beszedesi).
— Puthz, 1975c: 176 (Edaphus; synonym of beszedesi).
— Coiffait, 1984f: 358 (Edaphus; synonym of beszedesi).
bluehweissi Scheerpeltz, 1936: 201, 210 (Edaphus; Type locality: Gebiet des Kahlenberges bei Wien).
—Scheerpeltz, 1949: 29 (Edaphus; ecological and distributional notes; Austria).
— Lohse, 1963: 176 (Edaphus; valid species).
— Horion, 1963: 400 (Edaphus; Austria; Germany; Switzerland; Italy).
— Lohse, 1964: 130 (Edaphus; characters; central Europe).
— Jarrige, 1968a: 141 (Edaphus; characters; notes; France).
— Puthz, 1971p: 203 (Edaphus; synonym of rosskotheni).
— Puthz, 1974t: 923 (Edaphus; synonym of beszedesi).
— Puthz, 1975c: 176 (Edaphus; synonym of beszedesi).
— Coiffait, 1984f: 358 (Edaphus; synonym of beszedesi).
— Lucht, 1987: 98 (Edaphus; checklist; Germany; Czechoslovakia; Austria; Switzerland).
— Lohse and Lucht, 1989: 149 (Edaphus; synonym of beszedesi).
bicicatrix Comellini, 1977: 251 (Edaphus; Type locality: Sri Lanka: province Eastern, Kantalai).
— Puthz, 1979: 127 (Edaphus; characters; Nepal; Thailand).
▴ Distribution: Nepal, Thailand, Sri Lanka.
bidenticollis Puthz, 1990a: 217 (Edaphus; Type locality: Côte d’Ivoire: Agboville, Forêt de Yapo, près Yapo-Gare).
biforatus Puthz, 1985c: 356 (Edaphus; Type locality: Paraguay: Canendiyu prov., Estancia “La Cordillera”, env. 8 km sud de Ypé Jhu).
biformatus Puthz, 1992b: 26 (Edaphus; Type locality: Zaire: Mwenga, 1250 m).
biformis Puthz, 1992b: 25 (Edaphus; Type locality: Zaire: Bendera, terr. Albertville, 1000 m).
bifoveicollis Puthz, 1992b: 29 (Edaphus; Type locality: Angola: Saurimo, Foret-galerie chutes R. Chicapa).
bifurcus Puthz, 1990a: 196 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
biimpressus Puthz, 1981h: 403 (Edaphus; Type locality: Vietnam: Cuc phuongl, Ninh binh).
biliranensis Bernhauer, 1929c: 96, 110 (Edaphus; Type locality: Biliran).
binodulus Cameron, 1943: 5 (Edaphus; Type locality: Darjeeling district: Ghum, Tiger Hill, altitude 8500–10,000 feet).
bipunctatus Puthz, 1979: 126 (Edaphus; Type locality: Vietnam: Co xan).
bipunctipennis Puthz, 1987a: 452 (Edaphus; Type locality: Nepal: Gorkha Distr., Chuling Khola, Djinshi Kharka, 3400 m).
biroi Székessy, 1939: 103 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolabe-Bay).
bisulcatus Bernhauer, 1929c: 95, 102 (Edaphus; Type locality: Siargao).
bisulcicollis Puthz, 1990a: 209 (Edaphus; Type locality: Côte d’Ivoire: Man: Mt. Tonkoui).
bitalensis Kistner, 1962a: 622 (Edaphus; Type locality: Congo Republic: Kivu: Kalehe territory, Bitale, northwest of Kahuzi, 1800 m. Also cited from Ruanda).
bluehweissi Scheerpeltz, 1936, see: beszedesi reitter, 1914.
boettcheri Bernhauer, 1929c: 94, 97 (Edaphus; Type locality: Luzon: Mt. Polis).
bolamanus Puthz, 1990d: 39 (Edaphus; Type locality: [Guinea Portoghese] Guinea Bissau: Bolama).
borneensis Cameron, 1930d: 330, 331 (Edaphus; Type locality: Borneo: Mt. Matang. Also cited from Sumatra).
brachypterus Puthz, 1979: 123 (Edaphus; Type locality: Nepal: Barahbise gegen Ting Sang La [Wald oberhalb Durumtali am Aufstieg zum Ting Sang La], 2200–2300 m).
brasilianus Wendeler, 1956: 41 (Edaphus; Type locality: Brasilien: Nova Teutonia).
— Puthz, 1973k: 52 (Edaphus; characters; Brazil).
— Puthz, 1974q: 36 (Edaphus; aedeagus).
— Puthz, 1985c: 355 (Edaphus; Paraguay).
▴ Distribution: Brazil, Paraguay.
brevialatus Puthz, 1979: 123 (Edaphus; Type locality: Malaya, Kuala Buka bei Trengganu).
brevicornis Székessy, 1939: 103 (Edaphus; Type locality: Neu-Guinea: Erima, Astrolab-Bay).
brevipennis Cameron, 1943: 5 (Edaphus; Type locality: Darjeeling district: Ghum).
—Puthz, 1979: 120 (Edaphus; characters; India).
— Puthz, 1987a: 452 (Edaphus; Nepal).
▴ Distribution: India, Nepal.
brittoni Puthz, 1978c: 125 (Edaphus; Type locality: Queensland: Winescrub forest near Maipoton, 350 m).
brunneiventris Puthz, 1974r: 92 (Edaphus; Type locality: Nepal: Gurjakhan, 83°14′E, 28°37′N, 8500 feet).
—Puthz, 1979: 129 (Edaphus; characters; Nepal).
— Puthz, 1987a: 452 (Edaphus; Nepal).
▴ Distribution: Nepal. bryanti Puthz, 1973k: 58 (Edaphus; Type locality: Brazil: Rio de Janeiro, Corcovado).
▴ Distribution: Brazil.
bufo Puthz, 1985c: 352 (Edaphus; Type locality: Peru: Loreto: Samiria).
cambeforti Puthz, 1987: 33 (Edaphus; Type locality: Malawi: Forêt de Likhubula, 1100 m).
cameroni Comellini, 1977, see: duplex Puthz, 1979.
cameroni Puthz, 1974r: 91 (Edaphus; Type locality: India: Ghum District, Tiger Hill, 8500–10,000 feet).
camponoti Oke, 1933, see: termitophilus Bernhauer, 1916.
capitatus Bernhauer, 1929c: 95, 100 (Edaphus; Type locality: Luzon: Los Banos).
carinatus Casey, 1884, see: nitidus Motschulsky, 1857.
carinicollis Bernhauer, 1907: 380 (Edaphus; Type locality: Not cited).
—Bernhauer and Schubert, 1911: 187 (Edaphus; catalog).
— Cameron, 1943: 5 (Edaphus; characters [in footnote]).
— Puthz, 1975c: 172, 179 (Edaphus; characters; Japan).
— Ito, 1992: 63 (Edaphus; Yakushima Island).
— J. Li, 1993: 26 (Edaphus; China).
▴ Distribution: Japan, China.
carinipennis Bernhauer, 1929c: 96, 108 (Edaphus; Type locality: Luzon: Mt. Polis).
carinulifer Puthz, 1986c: 315 (Edaphus; Type locality: Gabun: Makokou).
chalcographus Puthz, 1979: 134 (Edaphus; Type locality: India: Maharashtra, Bhaja).
chinense J. Li, 1993: 155 (Edaphus; Type locality: Not cited but presumably China).
chinensis Bernhauer, 1934: 5 (Edaphus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m).
—Puthz, 1975c: 174 (Edaphus; characters; China).
— Puthz, 1979: 155 (Edaphus; characters).
— Puthz, 1983g: 163 (Edaphus; China).
▴ Distribution: China.
cinghalicus Comellini, 1977: 264 (Edaphus; Type locality: Sri Lanka: province Central, Mululla).
claripennis Puthz, 1990a: 219 (Edaphus; Type locality: Côte d’Ivoire: env. Gregbeu).
clavator Comellini, 1977: 265 (Edaphus; Type locality: Sri Lanka: province Central, Kandy).
clavicornis Puthz, 1989c: 57 (Edaphus; Type locality: Malaysia: Pahang: Bukit Rengit c/o Lanchang, 120 m).
clypeopunctus Puthz, 1990b: 277 (Edaphus; Type locality: Neu Guinea: Astrolabe Bay: Mt. Hansemann).
coiffaiti Puthz, 1986c: 314 (Edaphus; Type locality: Gabun: Makokou).
—Puthz, 1990a: 221 (Edaphus; characters).
— Puthz, 1992b: 18, 19 (Edaphus; characters).
▴ Distribution: Gabon.
colossus Comellini, 1977: 262 (Edaphus; Type locality: Sri Lanka: province Central, Horton Plains, 2100 m).
columbianus Puthz, 1973k: 54 (Edaphus; Type locality: Colombia: C. Amara, Tequendama, 7600 feet).
confusus Puthz, 1992b: 27 (Edaphus; Type locality: Zaire: Kibali-Ituri, terr. Epulu, Mambasa-Mungbere, 900 m).
congener Puthz, 1974q: 35 (Edaphus; Type locality: Brazil: Nova Teutonia).
congerminalis Puthz, 1986d: 620 (Edaphus; Type locality: Brasilien: Guanabara: Represa Rio Grande).
cooki Puthz, 1991: 278 (Edaphus; Type locality: Niue: Liku F.).
cordicollis Puthz, 1990c: 818 (Edaphus; Type locality: Sumatra: Aceh-Selatan, 100 m, Paya Laot, Meulaboh).
coriaceus Puthz, 1987: 29 (Edaphus; Type locality: Zaïre: Katanga: Kundelungu, 1750 m.
Also cited from Gabon, Angola, and N. Rhodesia).
corpulentus Puthz, 1979: 125 (Edaphus; Type locality: Malaya: Penang: Penang Hills, Tiger Hill, 2100 feet).
costaricensis Puthz, 1973k: 58 (Edaphus; Type locality: Costa Rica: Hamburgfarm, Reventazon, Ebene Limon).
crassus Puthz, 1973k: 55 (Edaphus; Type locality: Brazil: r. Santo, near São Paulo).
cribricollis Schaufuss, 1887: 109 (Edaphus; Type locality: Sumatra).
—Fauvel, 1905e: 81 (Edaphus; Java; Sumatra; Celebes).
— Bernhauer and Schubert, 1911: 187 (Edaphus; catalog).
— Bernhauer, 1928: 10 (Edaphus; Sumatra; Java).
— Cameron, 1930d: 330 (Edaphus; Sumatra).
— Scheerpeltz, 1933: 1203 (Edaphus; characters; catalog).
— Puthz, 1974r: 89 (Edaphus; characters).
— Puthz, 1979: 115 (Edaphus; Malaya).
— Puthz, 1990c: 823 (Edaphus; characters; Sumatra; Malaya).
▴ Distribution: Indonesia, Malaysia.
crinitus Puthz, 1979: 125 (Edaphus; Type locality: Thailand: Sakaerat Experimental Station, Wald oberhalb der Station. Also cited from Malaya and possibly Nepal).
cubanus Puthz, 1997d: 138 (Edaphus; Type locality: Cuba: Playa Marianao).
cuspidiventris Puthz, 1990c: 818 (Edaphus; Type locality: Sumatra: Aceh-Selatan, Babahrot, 100 m).
dapanus Bernhauer, 1929c: 96, 106 (Edaphus; Type locality: Siargao: Dapa; Luzon: Imugan).
densatus Fauvel, 1905e: 81 (Edaphus; Type locality: Tjibodas).
—Bernhauer and Schubert, 1911: 187 (Edaphus; catalog).
— Puthz, 1979b: 168 (Edaphus; characters; Java).
▴ Distribution: Indonesia.
densepunctatus Puthz, 1983f: 174 (Edaphus; Type locality: Zaïre: Kivu: Terr. Kalehe, Bitale, N.O. Kahusi, 1600 m).
densipennis Cameron, 1929b: 451 (Edaphus; Type locality: Selangor: The Gap).
—Scheerpeltz, 1933: 1203 (Edaphus; catalog).
— Puthz, 1979: 131 (Edaphus; characters; Malaysia).
▴ Distribution: Malaysia.
densus Bernhauer, 1934: 5 (Edaphus; Type locality: Mittel-Szechuan: Hwa-Yin-Shan, 1600 m).
dentiventris Fauvel, 1905a: 137 (Edaphus; Type locality: Singapore. Sumatra: Buitenzorg).
—Fauvel, 1905e: 81 (Edaphus; Java).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Cameron, 1921a: 369, 401 (Edaphus; characters; Singapore).
— Scheerpeltz, 1933: 1203 (Edaphus; catalog).
— Puthz, 1979: 157, 159 (Edaphus; characters).
— Puthz, 1979b: 169 (Edaphus; characters; Singapore; Sumatra).
— Puthz, 1990c: 824 (Edaphus; characters; Sumatra).
▴ Distribution: Singapore, Indonesia.
mandibularis Cameron, 1930d: 330 (Edaphus; Type locality: Fort de Kock).
depressus Puthz, 1973k: 56 (Edaphus; Type locality: Brazil: Alexandra, 20 km. W. Paranagua).
dilutus Schaufuss, 1887: 109 (Edaphus; Type locality: Tabac., Sumatra; [Note: See lectotype designation by Puthz, 1974r]).
—Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Cameron, 1930d: 330 (Edaphus; Sumatra).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1974r: 87 (Edaphus; lectotype designation: Tabac., Sumatra; characters; Vietnam; Java).
— Watanabe and Shibata, 1976: 330 (Edaphus; Malaysia).
— Puthz, 1979: 142 (Edaphus; characters).
— Puthz, 1990c: 824 (Edaphus; characters; Sumatra; Java; Vietnam; Luzon).
▴ Distribution: Myanmar, Indonesia, Malaysia, Philippines, Vietnam.
luridus Blattný, 1925: 185 (Microphthartus; Type locality: Birma).
magnipennis Bernhauer, 1929c: 96, 108 (Edaphus; Type locality: Luzon: Manila, Zambales).
—Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1979: 142 (Edaphus; synonym of dilutus).
— Puthz, 1990c: 824 (Edaphus; synonym of dilutus).
discipunctus Puthz, 1974, see: variabilis Puthz, 1974.
dissimilis Aubé, 1863: 37 (Euaesthetus; Type locality: Toulon).
—Fauvel, 1873: 61 [= 1873a: 219] (Euaesthetus; characters; France; Corsica).
— Mulsant and Rey, 1878a: 302 (Edaphus; characters; France).
— Ganglbauer, 1895: 543 (Edaphus; characters; France; Corsica; Italy; Dalmatia).
— Sainte-Claire Deville, 1906: 80 (Edaphus; Corsica).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Reitter, 1914: 188 (Edaphus; characters; Corfu).
— Portevin, 1929: 390 (Edaphus; characters; France).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Scheerpeltz, 1936: 200, 202 (Edaphus; characters; Pyrenees; Italy; France; Corsica; Corfu; Crete; Bosnia).
— Lohse, 1964: 130 (Edaphus; characters; central Europe).
— Jarrige, 1968a: 142 (Edaphus; characters).
— Puthz, 1975c: 172, 174 (Edaphus; characters; notes; France; Italy; Yugoslavia; Greece).
— Coiffait, 1984f: 357 (Edaphus; characters; Pyrenees; Crete; Corsica; Italy; Dalmatia).
— Orousset, 1986b: 20 (Edaphus; notes).
— Zanetti, 1995: 19 (Edaphus; Italy).
▴ Distribution: Europe.
kaufmanni Reitter, 1909b: 303 (Edaphus; Type locality: Fiume).
—Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Reitter, 1914: 188 (Edaphus; characters; Fiume).
— Scheerpeltz, 1936: 190 (Edaphus; republication of orginal description; characters; discussion).
— Scheerpeltz, 1938a (Edaphus; characters; notes).
— Puthz, 1975c: 174 (Edaphus; synonym of dissimilis).
— Coiffait, 1984f: 357 (Edaphus; synonym of dissimilis).
divisicollis Puthz, 1979: 128 (Edaphus; Type locality: India: W. Bengal, Darjeeling District, Ghum, 2200 m).
dollmani Puthz, 1990d: 37 (Edaphus; Type locality: [N. Rhodesia] Zambia: Namwala).
dubius Puthz, 1998b: 49 (Edaphus; Type locality: China: Hongkong: Tai Po).
duplex Puthz, 1979: 139 (Edaphus; replacement name for cameroni Comellini).
cameroni Comellini, 1977: 267 (Edaphus; [preoccupied]; Type locality: Sri Lanka: province North Western, Rajakadaluwa).
electrus Puthz, 1987a: 451 (Edaphus; Type locality: Nepal: südliches Annapurna-Massiv, Parbat Distr., zw. Chitre und Ghandrung, Chitre-Seite des Passes, 2950–3050 m).
elongatus Puthz, 1979: 132 (Edaphus; Type locality: Nepal: Sandakphu, 27°06′N, 38°01′E, 12,000 feet).
endrodyi Jacobson and Kistner, 1974, see: adazla Kistner 1961.
etiennei Orousset, 1988, see: remyi Jarrige, 1963.
euplectoides Sharp, 1886: 640 (Edaphus; Type locality: Guatemala, Capetillo).
—Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Puthz, 1973k: 54 (Edaphus; characters; Guatemala).
excellens Puthz, 1991: 275 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m).
excelsiventris Puthz, 1991: 272 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m).
excisicollis Puthz, 1991: 268 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m).
exemplatus Puthz, 1991: 277 (Edaphus; Type locality: Fiji: Lau Islands: Lakeba Island: Rasea forest).
exiguus Puthz, 1991: 282 (Edaphus; Type locality: Fiji: Viti Levu: Nandrau, 750 m).
exiliventris Puthz, 1991: 273 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu, 1000 m).
exorbitans Puthz, 1991: 274 (Edaphus; Type locality: Fiji Vanua Levu: Ndelaikoro).
exornaticollis Puthz, 1991: 269 (Edaphus; Type locality: Fiji: Viti Levu: Nandrau, 750 m).
exosus Puthz, 1991: 286 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 800 m).
exoticus Puthz, 1991: 266 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m.).
expeditus Puthz, 1991: 279 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 500 m).
explorator Puthz, 1991: 280 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatau ridge, 1000 m).
exquisitus Puthz, 1991: 284 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 450 m).
exscriptus Puthz, 1991: 281 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 500 m).
exsecticollis Puthz, 1991: 269 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 800 m).
exstans Puthz, 1991: 274 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu, 900 m).
exsulans Puthz, 1991: 287 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 450 m).
extennatus Puthz, 1991: 283 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 500 m).
externus Puthz, 1991: 286 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 800 m).
exterraneus Puthz, 1991: 276 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m).
fauveli Puthz, 1979b: 165 (Edaphus; Type locality: Singapore. Also cited from Sumatra).
—Puthz, 1979: 117 (Edaphus; characters; Malaya; Vietnam; Borneo; Malayasia).
— Puthz, 1990c: 823 (Edaphus; characters; Malaya; Sumatra; Singapore; Borneo; Vietnam).
▴ Distribution: Borneo, Indonesia, Malaysia, Singapore, Vietnam.
feai Puthz, 1981h: 401 (Edaphus; Type locality: Burma: Bhamo).
ferdinandae Puthz, 1997d: 145 (Edaphus; Type locality: Cuba: Sierra del Rosario, Rangel).
fijiensis Puthz, 1974r: 88 (Edaphus; Type locality: Fiji: Taveuni).
flavescens Jacobson and Kistner, 1974, see: adazla Kistner, 1961.
fossicollis Puthz, 1979: 128 (Edaphus; Type locality: India: W. Bengal: Darjeeling District, Kurseong).
foveicollis Bernhauer, 1929c: 95, 101 (Edaphus; Type locality: Luzon: Balbalan, Mt. Polis).
franzi Puthz, 1974: 763 (Edaphus; Type locality: Tchad: Polder von Guini bei Bol).
— Puthz, 1984g: 198 (Edaphus; Ghana).
— Puthz, 1990a: 221 (Edaphus; characters).
— Puthz, 1992b: 16, 19 (Edaphus; use of franzi by Puthz, 1983f: 173 was misidentification and is renamed simillimus; characters).
▴ Distribution: Chad, Ghana.
franzianus Puthz, 1974r: 84 (Edaphus; Type locality: Thailand: Sakaerat Exp. Stat.).
franziellus Puthz, 1979a: 75 (Edaphus; characters published under “Edaphus C” by Puthz, 1978c: 122, 128; Type locality: S. Australia: env. Adelaide).
furcaticollis Puthz, 1979: 136 (Edaphus; Type locality: India: W. Bengal, Darjeeling, North Point, 1300 m).
fuscescens Orousset, 1986e: 76 (Edaphus; Type locality: Venezuela: Cordillera de Mérida, Sierra de la Culata, Mucubaji, alt. 3600 m).
garambicus Kistner, 1961: 37 (Edaphus; Type locality: Congo: Parc National de la Garamba).
—Kistner, 1962a: 624 (Edaphus; characters).
— Puthz, 1974s: 757 (Edaphus; characters).
— Puthz, 1984g: 198 (Edaphus; characters; Ghana).
▴ Distribution: Congo, Ghana.
geginati Puthz, 1994c: 381 (Edaphus; Type locality: Zaire: Ruwenzori: Ostseite, Oberlauf des Cunsilubi, 3900 m).
globulicollis Székessy, 1939: 102 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay and Marega).
gouzei Orousset, 1988c: 186 (Edaphus; Type locality: Thaïlande: env. de Chiang Maï, Doï Inthanon, alt. 2500 m).
gracilis Puthz, 1979: 140 (Edaphus; Type locality: India: W. Bengal, Darjeeling District, Ghum, 2200 m).
grandis Bernhauer, 1929c: 94, 97 (Edaphus; Type locality: Luzon: Balbalan).
granulifer Puthz, 1990a: 200 (Edaphus; Type locality: Côte d’Ivoire: Agboville: Forêt de Yapo, près Yapo-Gare).
granulipennis Puthz, 1979: 139 (Edaphus; Type locality: Vietnam: Tuong linh near Phu ly).
hammondi Puthz, 1975c: 180 (Edaphus; Type locality: Nordchina: Heilungkiang, Harbin).
heikertingeri Bernhauer, 1929c: 95, 100 (Edaphus; Type locality: Siargao: Dapa).
longicollis Bernhauer, 1929c: 100 (Edaphus; Type locality: Siargao; Samar, Catbalogan).
hepaticola Puthz, 1974r: 89 (Edaphus; Type locality: Nepal: Gurjakhan, 83°14′E, 28°37′N, 8500 feet).
hirsutus Puthz, 1973l: 147 (Edaphus; Type locality: Mexico: Oaxaca, 15 miles South Valle Nacional, 4000 feet).
hirtus Bernhauer, 1929c: 96, 107 (Edaphus; Type locality: Luzon: Imugan).
holdhausi Székessy, 1939: 104 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay).
hova Jarrige, 1968: 877 (Edaphus; Type locality: Madagascar-Est, Périnet).
huggerti Orousset, 1987a: 19 (Edaphus; Type locality: Equateur: Pichin., Tinalandia).
humeralis Puthz, 1973k: 55 (Edaphus; Type locality: Brazil: Para, Faz. Pirelli Belem).
—Puthz, 1985c: 352 (Edaphus; characters; Paraguay; Costa Rica; Panama).
— Puthz, 1997d: 138 (Edaphus; notes).
▴ Distribution: Costa Rica, Panama, Surinam, Brazil, Paraguay.
aschnaae Makhan, 1995: 36 (Edaphus; Type locality: Suriname: District Commewijne).
rishwani Makhan, 1995: 35 (Edaphus; Type locality: Suriname: District Commewijne).
illotus Puthz, 1987: 32 (Edaphus; Type locality: Äthiopien: Gamo province: Mt. Gughé, ca. 10,500 feet).
imitator Puthz, 1979: 116 (Edaphus; Type locality: Thailand: Sakaerat Exp. Station).
impennatus Puthz, 1979: 122 (Edaphus; Type locality: India: W. Bengal, Darjeeling District, 3 km S of Ghum. Also cited from Nepal).
impenniger Puthz, 1979: 122 (Edaphus; Type locality: Nepal: Khumbu, Schluchtwald am Shutlje-Khola, oberhalb des Ortes Shutlje).
inaequalis Puthz, 1979: 115 (Edaphus; Type locality: S-Thailand: Kachong Experimental Station bei Trang, 80 m).
incertus Bernhauer, 1929c: 96, 105 (Edaphus; Type locality: Luzon: Mt. Banahao).
incisicollis Puthz, 1987a: 449 (Edaphus; Type locality: Nepal: Parbat Distr., zw. Chitre und Ghandrung, Chitre-Seite des Passes, 2950–3050 m).
inconspicuus Puthz, 1990a: 201 (Edaphus; Type locality: Côte d’Ivoire: Forêt de Languédédou).
infidus Puthz, 1997d: 142 (Edaphus; Type locality: Cuba: Playa Marianao).
ingloriosus Puthz, 1990a: 205 (Edaphus; Type locality: Côte d’Ivoire: Agboville: Forêt de Yapo, près Yapo-Gare).
inglorius Puthz, 1986c: 313 (Edaphus; Type locality: Gabun: Belinga).
ingratus Puthz, 1990a: 208 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
ingravescens Puthz, 1990a: 208 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
insignis Bernhauer, 1929c: 94, 99 (Edaphus; Type locality: Süd-Luzon: Mt. Isarog).
insigniventris Bernhauer, 1929c: 95, 104 (Edaphus; Type locality: Luzon: Mt. Makiling).
insperatus Puthz, 1979: 138 (Edaphus; Type locality: Malaya: Pahang, Temerloh).
invidiosus Puthz, 1978c: 124 (Edaphus; Type locality: Queensland: Dividing Range, highway to Warwick, 800 m).
invocatus Puthz, 1992b: 13 (Edaphus; Type locality: Zaire: Kivu: Mwenga, 1250 m).
jaccoudi Puthz, 1979: 118 (Edaphus; Type locality: Malaya: Pahang, Cameron Highlands, Berinchang).
japonicus Bernhauer, 1907, see: perversus Puthz, 1994.
japonicus Sharp, 1889: 325 (Edaphus; Type locality: Nagasaki; [Note: See lectotype designation by Puthz, 1975c]).
—Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Puthz, 1975c: 172, 180 (Edaphus; lectotype designation; characters; Japan).
jeanneli Puthz, 1987: 34 (Edaphus; Type locality: Tanzania: Kilimandjaro versant Sud-Est, zône inférieure, Neu-Moschi, 800 m. Also cited from Kenya).
jiloriensis Puthz, 1990d: 35 (Edaphus; Type locality: Kenya: Kilifi district, Jilore [ouest de Malindi]).
juanae Puthz, 1997d: 143 (Edaphus; Type locality: Cuba: Aspiro).
kabobomontis Puthz, 1992b: 21 (Edaphus; Type locality: Zaire: Mont Kabobo, terr. Albertville, Hte. Kiymbi, 1700 m).
kahuziensis Kistner, 1962a: 624 (Edaphus; Type locality: Congo Republic: Kivu: counterfort South of Kahuzi, 2300 m. Also cited from Ruanda).
kalehensis Kistner, 1962a: 626 (Edaphus; Type locality: Congo Republic, Kalehe territory, southwest of Kahuzi, Mukaba River, 2200 m. Also cited from Ruanda).
kashmiricus Puthz, 1979: 121 (Edaphus; Type locality: Kashmir: Daksum, 2400–2700 m).
kaszabi Jacobson and Kistner, 1974: 89 (Edaphus; Type locality: Ghana: Brong-Ahafo Region, Kintampo Catarract, Fullner Falls).
kaufmanni Reitter, 1909, see: dissimilis Aubé, 1863.
kenyanus Puthz, 1990d: 36 (Edaphus; Type locality: Kenya: Kilifi district, Jilore [ouest de Malindi).
kivuensis Puthz, 1974s: 758 (Edaphus; Type locality: Kivu: Kabare, Nyakasiba, 2350 m).
klapperichi Puthz, 1990a: 197 (Edaphus; Type locality: Sierra Leone: Guma mountain).
kuscheli Puthz, 1991: 271 (Edaphus; Type locality: Fiji: Viti Levu: Nandarivatu ridge, 1000 m).
kuschelianus Puthz, 1991: 270 (Edaphus; Type locality: Fiji: Viti Levu: Nandrau, 750 m).
laticeps Bernhauer, 1929c: 95, 103 (Edaphus; Type locality: Biliran).
lederi Eppelsheim, 1878: 118 (Edaphus; Type locality: Elisabetthal).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Reitter, 1914: 189 (Edaphus; characters; Transcaucasien region).
— Scheerpeltz, 1936: 201, 206 (Edaphus; characters; Caucasus).
— Puthz, 1975c: 173, 174, 176 (Edaphus; characters; Caucasus).
— Coiffait, 1984f: 358 (Edaphus; characters; Caucasus).
— Orousset, 1986b: 20 (Edaphus; notes).
▴ Distribution: Caucasus.
leileri Puthz, 1989c: 58 (Edaphus; Type locality: Malaysia: Insel Tioman: Tekek-Jaura).
leleupi Kistner, 1962a: 626 (Edaphus; Type locality: Congo Republic: Katanga, Kakungu).
limbicollis Puthz, 1990a: 216 (Edaphus; Type locality: Côte d’Ivoire: Forêt de Taï, 10 km ouest de la Station scientifique).
lineatocollis Puthz, 1979: 129 (Edaphus; Type locality: Nepal: Phulchoki, 2600 m).
lineatus Puthz, 1979: 135 (Edaphus; Type locality: Nepal: 5 km Manhari, 350 m).
lineolatus Puthz, 1987a: 448 (Edaphus; Type locality: Nepal: südliches Annapurna-Massiv, Kaski Distr., oberhalb Dhumpus, 2100 m).
lineolifer Puthz, 1998b: 51 (Edaphus; Type locality: China: Hongkong: Kadoorie Farm).
loebli Comellini, 1977: 256 (Edaphus; Type locality: Sri Lanka: province Uva, Inginiyagala).
longicollis Bernhauer, 1929, see: heikertingeri Bernhauer, 1929.
lorianus Puthz, 1990b: 278 (Edaphus; Type locality: N. Guinea mer.: Bujakori [nahe Kemp-Welch-River, etwa 147°46′E/9°42′S]).
luberoensis Puthz, 1992b: 19 (Edaphus; Type locality: Zaire: Forêt de Kasuo SW de Lubero).
lubukensis Kistner, 1962a: 627 (Edaphus; Type locality: Congo Republic: Kivu, Uvira territory, Lubuka, 2780 m).
luculentus Casey, 1884, see: nitidus Motschulsky, 1857.
luridus Blattný, 1925, see: dilutus Schaufuss, 1887.
luzonicus Bernhauer, 1929c: 94, 98 (Edaphus; Type locality: Luzon: Imugan).
lynceus Puthz, 1990a: 210 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
mabaliensis Kistner, 1962a: 628 (Edaphus; Type locality: Congo Republic: Equateur, Tshuapa, Mabali near Lake Tumba, 350 m).
magnipennis Bernhauer, 1929, see: dilutus Schaufuss, 1887.
mahnerti Puthz, 1990a: 214 (Edaphus; Type locality: Côte d’Ivoire: Forêt de Taï, env. de la Station).
major Fauvel, 1895: 205 (Edaphus; Type locality: Bírmanie, Bhamo).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Cameron, 1930: 448 (Edaphus; characters; Burma).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Coiffait, 1978b: 112 (Edaphus; Bhutan).
— Puthz, 1979: 158 (Edaphus; characters).
— Puthz, 1979b: 161 (Edaphus; characters; Burma).
▴ Distribution: Bhutan, Myanmar.
mandibularis Cameron, 1930, see: dentiventris Fauvel, 1905.
marlieri Kistner, 1962, see: africanus Eppelsheim, 1885.
marshalli Bernhauer, 1934, see: africanus Eppelsheim, 1885.
martensi Puthz, 1987a: 449 (Edaphus; Type locality: Nepal: Parbat Distr., Chitre, 2400 m).
mediopunctus Puthz, 1974s: 64 (Edaphus; Type locality: Congo-Brazzaville: Galerienwald bei Bampaka, westlich von Brazzaville, Talmulde).
melculus Oke, 1933: 110 (Edaphellus; Type locality: Australia: Victoria: Belgrave, Gembrook, Warburton, Preston, Macedon).
mendax Puthz, 1979: 137 (Edaphus; Type locality: Singapore. Also cited from Siam).
mendicus Puthz, 1979: 116 (Edaphus; Type locality: Zentral-Nepal: Pande-Sattel zwischen Biritanti und Naudanda westlich Pokhara, ca. 1800 m. Also cited from Thailand).
mendosus Puthz, 1998b: 52 (Edaphus; Type locality: China: Hongkong: Kadoorie Farm).
meo Orousset, 1988c: 186 (Edaphus; Type locality: Thaïlande: env. de Chiang Maï, chiang Dao, alt. 500 m).
mexicanus Puthz, 1973k: 54 (Edaphus; Type locality: Mexico: Chiapas, 8 mi. S. Simojovel).
microphthalmus Puthz, 1981g: 455 (Edaphus; Type locality: Nepal: Taoundja, massif du Lamjung Himal, 2860 m).
microps Puthz, 1976f: 156 (Edaphus; Type locality: Ceylon: Sabaragamuwa, Ratuapura).
micros Coiffait, 1981d: 332 (Edaphus; Type locality: Jagath, 1650 m).
mindoroensis Bernhauer, 1929c: 96, 106 (Edaphus; Type locality: Mindoro: S. Theodoro).
minutus Puthz, 1979: 130 (Edaphus; Type locality: Nepal: Phulchoki, Ost-Hang, ca. 2000 m, bei Dalikhel).
mjobergi Bernhauer, 1916a: 3 (Edaphus; cited as mjöbergi; Type locality: Queensland: Blackal Range).
— Scheerpeltz, 1933: 1204 (Edaphus; cited as mjöbergi; catalog).
— Oke, 1933: 110 (Edaphus; cited as mjobergi; notes on description).
— Puthz, 1978c: 126 (Edaphus; cited as mjöbergi; characters; Australia).
— Herman, 2001: 12 (Edaphus; spelling corrected to mjobergi).
▴ Distribution: Australia.
moloensis Puthz, 1974s: 756 (Edaphus; Type locality: Kenya: Molo, Mau Escarpment, 2150–2200 m).
monopticus Puthz, 1992b: 20 (Edaphus; Type locality: Zaire: Kivu: Itombwe, 2300 m., Mulenge, Nyalengwe).
monticola Puthz, 1975c: 177 (Edaphus; Type locality: China: West-Szechuan: Wassuland, Bezirk San-kiang-kou, Strecke S-S bis 3000 m).
montisamator Comellini, 1977: 266 (Edaphus; Type locality: Sri Lanka: province Central, Pidurutalagala, 2200 m).
montius Comellini, 1977: 263 (Edaphus; Type locality: Sri Lanka: province Central, Horton Plains, 2100 m).
mussardi Comellini, 1977: 252 (Edaphus; Type locality: Sri Lanka: province North Western, Rajakadaluwa).
najtae Orousset, 1986e: 75 (Edaphus; Type locality: Venezuela: Cordillera de Mérida: Sierra de la Culata, páramo de Piedras Blancas, alt. 4250 m).
nakayamaensis Puthz, 1980h: 243 (Edaphus; Type locality: Japan: Hokkaido: Nakayama-Pass südlich Sapporo, über 1000 m).
napporoensis Puthz, 1980h: 244 (Edaphus; Type locality: Japan: Hokkaido: Napporo National Park bei Sapporo, 60–70 m).
neboissi Puthz, 1978c: 123 (Edaphus; Type locality: S. Australia: Englebrook Nat-Trust Res. near Adelaide. Also cited from Victoria).
nepalensis Puthz, 1979: 121 (Edaphus; Type locality: Nepal: Barahbise gegen Ting Sang La [Wald oberhalb Durumtali am Aufstieg zum Ting Sang La, 2200–2300 m]).
nepalicus Coiffait, 1982: 104 (Edaphus; Type locality: Nepal central: Madi Khola, 2385 m).
nequam Puthz, 1990a: 202 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
nipponensis Puthz, 1975c: 181 (Edaphus; Type locality: Japan: Kanagawa).
nitidifrons Puthz, 1990c: 816 (Edaphus; Type locality: Sumatra: Aceh-Selatan, Bahahrot, 100 m).
nitidus LeConte, 1861, see: nitidus Motschulsky, 1857.
nitidus Motschulsky, 1857: 7 (Edaphus; Type locality: Am. bor.; [Note: See lectotype designation by Puthz, 1974t]).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Kistner, 1962a: 621 (Edaphus; characters; Pennsylvania; Illinois; Indiana; Washington D.C.; Louisiana; Mississippi).
— Puthz, 1974t: 915 (Edaphus; lectotype designation: Am. bor.; characters; notes; Pennsylvania, Indiana, Illinois, Iowa, District of Columbia, Virginia, West Virginia, North Carolina, South Carolina, Georgia, Florida, Kentucky, Louisiana, Texas).
— Downie and Arnett, 1996: 461 (Edaphus; characters; USA).
▴ Distribution: USA.
nitidus LeConte, 1861a: 68 (Edaphus; [preoccupied]; Type locality: Louisiana).
— LeConte, 1863: 25 (Edaphus; North America).
— LeConte, 1863a: 50 (Edaphus; characters; Alabama, Louisiana).
— Casey, 1884a: 29 (Edaphus; characters; Alabama; Louisiana).
— Horn, 1885b: 121 (Edaphus; characters).
— Herman, 2001: 00 (Edaphus; synonym of nitidus Motschulsky).
carinatus Casey, 1884a: 30 (Edaphus; Type locality: Washington, D.C.).
— Horn, 1885b: 121 (Edaphus; synonym of nitidus).
— Horn, 1885a: 109 (Edaphus; synonym of nitidus).
— Bernhauer and Schubert, 1911: 188 (Edaphus; synonym of nitidus).
— Kistner, 1962a: 621 (Edaphus; synonym of nitidus).
— Puthz, 1974t: 916 (Edaphus; synonym of nitidus).
luculentus Casey, 1884a: 31 (Edaphus; Type locality: District of Columbia; [Note: See lectotype designation by Puthz, 1974t]).
— Horn, 1885b: 121 (Edaphus; characters).
— Horn, 1885a: 109 (Edaphus; synonym of nitidus).
— Bernhauer and Schubert, 1911: 188 (Edaphus; synonym of nitidus).
— Kistner, 1962a: 621 (Edaphus; synonym of nitidus).
— Puthz, 1974t: 916 (Edaphus; lectotype designation; synonym of nitidus).
novaeguineae Fauvel, 1878f: 220 (Edaphellus; Type locality: Nouvelle-Guinée, Hatam).
— Bernhauer and Schubert, 1911: 188 (Edaphellus; catalog).
— Székessy, 1939: 98 (Edaphus; characters; New Guinea).
— Puthz, 1990: 275 (Edaphus; characters; New Guinea).
▴ Distribution: New Guinea.
noviesimpressus Puthz, 1998b: 53 (Edaphus; Type locality: China: Hongkong: Kadoorie Farm).
octopunctatus Kistner, 1962a: 628 (Edaphus; Type locality: Congo Republic: Kivu, counterfort S. Kahuzi, 2300 m).
oculatus Jarrige, 1963a: 122 (Edaphus; Type locality: Ceylan: I d’Alugtgama).
okei Puthz, 1979a: 75 (Edaphus; characters published under “Edaphus D” by Puthz, 1978c: 123, 129; Type locality: Queensland: Dividing Range, highway to Warwick, 800 m).
okinawaensis Puthz, 1980h: 241 (Edaphus; Type locality: Okinawa: Nishime-dake).
ophthalmicus Bernhauer, 1929c: 95, 102 (Edaphus; Type locality: Mindanao: Surigao).
pahangensis Puthz, 1979: 119 (Edaphus; Type locality: Malaya: Pahang, Cameron Highlands, Ringlet, 3200 feet).
papuanus Székessy, 1939: 100 (Edaphus; Type locality: Neu-Guinea: Stephansort, Astrolab-Bay).
parvipennis Bernhauer, 1929c: 96, 109 (Edaphus; Type locality: Luzon: Imugan).
peninsularis Puthz, 1979: 119 (Edaphus; Type locality: Malaya: Pahang, Temerloh District, Krau Game Reserve).
perminimus Puthz, 1990c: 822 (Edaphus; Type locality: West-Sumatra: Indrapoera).
permodestus Puthz, 1990a: 203 (Edaphus; Type locality: Côte d’Ivoire: Monogaga).
perpusillus Coiffait, 1981d: 333 (Edaphus; Type locality: Kali Gandaki, 1900 m).
perreti Puthz, 1990a: 218 (Edaphus; Type locality: Côte d’Ivoire: Forêt de Taï).
persimilis Puthz, 1992b: 15 (Edaphus; Type locality: Zaire: Kwango, terr. de Feshi, rive gauche Kwenge).
peruanus Puthz, 1973l: 148 (Edaphus; Type locality: Peru: Umgebung Tarapoto, Strasse nach Yutimagua am Huallaga).
perversus Puthz, 1994e: 390 (Edaphus; replacement name for japonicus Bernhauer).
japonicus Bernhauer, 1907: 380 (Euaesthetus; preoccupied; Type locality: Ocyama, 2000 feet).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Puthz, 1994e: 390 (Edaphus; synonym of perversus).
petzi Bernhauer, 1929c: 94, 98 (Edaphus; Type locality: Nonoc; Luzon: Balbalan).
peyrierasi Orousset, 1988: 42 (Edaphus; Type locality: Madagascar Est: col de Madiorano, à l’Est de Vondrozo, 700 m).
philippinus Bernhauer, 1929c: 96, 109 (Edaphus; Type locality: Luzon: Balbalan, Manila; Siargao: Dapa).
phloeocharis Puthz, 1986d: 619 (Edaphus; Type locality: Paraguay: Provinz Itapua, 10 km sud de Santa Maria).
piceus Cameron, 1933, see: beszedesi Reitter, 1914.
pilosus Puthz, 1979: 124 (Edaphus; Type locality: Nepal: Barahbise gegen Ting Sang La [Wald oberhalb Durumtali am Aufstieg zum Ting Sang La], 2200–2300 m).
planifrons Puthz, 1990f: 260 (Edaphus; Type locality: Brasilien: Floresta de Tijuca bei Rio de Janeiro, N. P. Alto de Boavista).
planus Puthz, 1979: 140 (Edaphus; Type locality: Vietnam: Cuc Phuong, Ninh binh. Also cited from Thailand).
plaumanni Puthz, 1978e: 213 (Edaphus; Type locality: Brazil: Santa Catarina, Nova Teutonia).
plicatulus Schaufuss, 1877: 25 (Tetratarsus; Type locality: Bangkok, Siam; [Note: See lectotype designation by Puthz, 1974r]).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1974r: 83 (Edaphus; lectotype designation: Bangkok, Siam; characters).
— Puthz, 1979: 156 (Edaphus; characters).
▴ Distribution: Thailand.
plicifer Puthz, 1979: 136 (Edaphus; Type locality: Malaya: Perak, Cameron Highlands, Cascade Sungei Simei).
plicipennis Puthz, 1978c: 42 (Edaphus; Type locality: Neuguinea: Papua: near Kokoda, 500 m).
politus Motschulsky, 1857: 7 (Edaphus; Type locality: Nouvelle-Orléans).
— Puthz, 1974t: 915, 918 (Edaphus; characters; notes; Kentucky; Louisiana; Texas).
— Downie and Arnett, 1996: 461 (Edaphus; characters; USA).
▴ Distribution: USA.
poulaini Orousset, 1986e: 78 (Edaphus; Type locality: Pérou: Loreto, Genaro Herrera, Río Ucayali, alt. 200 m).
praefectus Puthz, 1990f: 259 (Edaphus; Type locality: Ecuador: prov. Imbabura, Mojanda nord, 3470 m).
princeps Bernhauer, 1929c: 95, 100 (Edaphus; Type locality: Siargao: Dapa).
bakerianus Bernhauer, 1929c: 100 (Edaphus; subspecies of princeps Type locality: Luzon: Mt. Makiling).
pselaphoides Eichelbaum, 1913: 122 (Edaphus; Type locality: Amani, Berg Bomole).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Kistner, 1960: 31 (Edaphus; Tanganyika).
— Kistner, 1962a: 632 (Edaphus; incertae sedis).
— Puthz, 1975d: 39 (Edaphus; characters; Tanzania).
— Puthz, 1979b: 172 (Edaphus; characters; Kenya).
▴ Distribution: Kenya, Tanzania.
pubescens Bernhauer, 1929c: 96, 105 (Edaphus; Type locality: Luzon: Mt. Polis, Balbalan).
pulliclavis Puthz, 1990c: 821 (Edaphus; Type locality: Sumatra: Aceh-Selatan, Babahrot, 100 m).
pumilus Puthz, 1979: 138 (Edaphus; Type locality: Malaya: Gombak near Kuala Lumpur, ca. 800 feet).
punctatus Fauvel, 1895: 206 (Edaphus; Type locality: Pulo-Penang; [Note: See lectotype designation by Puthz, 1979b]. Also cited from Bírmanie, Bhamo).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Cameron, 1930: 449 (Edaphus; characters; Burma; Penang).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1979b: 162 (Edaphus; lectotype designation: Penang; characters).
— Coiffait, 1981d: 324 (Edaphus; Nepal).
▴ Distribution: Myanmar, Malaysia, Nepal.
punctatus Székessy, 1939, see: szekessyi Puthz, 1971.
puncticeps Puthz, 1974s: 765 (Edaphus; Type locality: Congo-Brazzaville: Galerienwald bei Bampaka, westlich Brazzaville, Talmulde).
puncticollis Puthz, 1979: 134 (Edaphus; Type locality: Thailand: Sakaerat Experimental Station).
punctulatus Puthz, 1979: 131 (Edaphus; Type locality: India: Darjeeling District, Chim-Khona, Ghum, 2200 m).
punctuliceps Puthz, 1998b: 54 (Edaphus; Type locality: China: Hongkong: Lantau Island: Tei Tong Tsai).
punctulifrons Puthz, 1990a: 199 (Edaphus; Type locality: Côte d’Ivoire: Forêt de Taï, env. de la Station).
puriventris Puthz, 1992b: 14 (Edaphus; Type locality: Rwanda: Forêt de la Rugege, 2150 m).
puthzi Comellini, 1977: 254 (Edaphus; Type locality: Sri Lanka: province Western, Colombo).
pygmaeus Székessy, 1939: 100 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay).
quadripunctus Kistner, 1962: 104 (Edaphus; Type locality: Angola: Parque Carisso, Dundo).
— Kistner, 1962a: 629 (Edaphus; characters).
— Kistner, 1963: 199 (Edaphus; Congo Republic).
— Puthz, 1974s: 761 (Edaphus; characters).
— Puthz, 1984g: 198 (Edaphus; characters; Angola; Zaire; Ghana).
▴ Distribution: Angola, Congo, Ghana.
quisquilius Puthz, 1990f: 262 (Edaphus; Type locality: Ecuador: Pichin[.], Rio Palenque).
rastellus Comellini, 1977: 260 (Edaphus; Type locality: Sri Lanka: province Central, Hasalaka près Weragamtola).
remilleti Orousset, 1983b: 116 (Edaphus; Type locality: Mont Grand-Matoury, 7 km S. de Cayenne, alt. 200 m, Guyane française).
remyi Jarrige, 1963a: 120 (Edaphus; Type locality: Ceylan: Ratnapura).
— Puthz, 1979: 132 (Edaphus; characters).
— Orousset, 1988: 47 (Edaphus; characters; Sri Lanka).
▴ Distribution: Sri Lanka; Réunion.
batrisoides Comellini, 1977: 256 (Edaphus; Type locality: Sri Lanka: province Western, Colombo).
etiennei Orousset, 1988: 50 (Edaphus; subspecies of remyi; Type locality: La Réunion: forêt de Mare Longue).
reticulatus Puthz, 1987: 30 (Edaphus; Type locality: Äthiopien: Chillalo, ca. 9000 feet).
riedeli Puthz, 1994d: 385 (Edaphus; Type locality: Indien: Kerala: Thekkady Peryar, W.L.S.).
rishwani Makhan, 1995, see: humeralis Puthz, 1973.
riukiuensis Puthz, 1980h: 242 (Edaphus; Type locality: Okinawa: Kanna).
rivalis Puthz, 1979: 133 (Edaphus; Type locality: Malaya: Gombak, near Kuala Lumpur, ca. 800 feet).
rosskotheni Wüsthoff, 1935, see: beszedesi Reitter, 1913.
rougemonti Puthz, 1998b: 54 (Edaphus; Type locality: China: Hongkong: Tai Po).
ruandae Puthz, 1992b: 24 (Edaphus; Type locality: Ruanda: Forêt de la Rugege, 2150 m).
ruwenzorianus Puthz, 1983f: 175 (Edaphus; Type locality: Zaire: Face N. Ruwenzori, Kilindera, 2750–2950 m).
scheerpeltzi Bernhauer, 1929, see: sulciceps Bernhauer, 1929.
scherpa Puthz, 1980i: 246 (Edaphus; Type locality: Nepal: Umgebung Pokhara: Phewa-See, Wald am Abhang gegenüber dem Tempelhügel bei Piota).
sechellarum Bernhauer, 1922b: 170 (Edaphus; Type locality: Mahé; [Note: See lectotype designation by Orousset, 1985b]. Originally cited from Seychelles: Silhouette: high forest above Mare aux Cochons and Mahé: near Morne Blanc).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Kistner, 1962a: 629 (Edaphus; characters).
— Orousset, 1985b: 250 (Edaphus; lectotype designation: Mahé, Seychelles).
— Orousset, 1988: 52 (Edaphus; characters; Seychelles).
▴ Distribution: Seychelles.
setosus Puthz, 1979: 135 (Edaphus; Type locality: Nepal: Sanghu Base, Camp area, ca. 6500 feet, 27°21′N, 87°33′E).
sexangulicollis Puthz, 1990b: 282 (Edaphus; Type locality: D’Entrecasteaux Islands: Fergus-son Island: Deidei, Gomwa Bay, 0–20 m, Camp 6).
sexpunctus Kistner, 1962, see: africanus Eppelsheim, 1885.
siamensis Puthz, 1979: 137 (Edaphus; Type locality: Thailand: Sakaerat Experimental Station).
similigenus Puthz, 1992b: 16 (Edaphus; Type locality: Zaire: Tshuapa, terr. Bikoro, Mabali).
similior Puthz, 1979: 115 (Edaphus; Type locality: India: W. Bengal: Darjeeling District, North Point, 1500 m. Also cited from Nepal).
similis Puthz, 1979a: 75 (Edaphus; characters published under “Edaphus E” by Puthz, 1978c: 127, 130; Type locality: Queensland: Dividing Range, highway to Warwick, 800 m).
simillimus Puthz, 1992b: 16 (Edaphus; Type locality: Zaire: Thysville).
— Puthz, 1983f: 173 (Edaphus; [Note: Misidentified as franzi according to Puthz, 1992b: 16]; characters; Zaire).
▴ Distribution: Congo.
singaporensis Puthz, 1979: 126 (Edaphus; Type locality: Singapore: Bukit Timah).
sparsus Puthz, 1979: 117 (Edaphus; Type locality: Malaya: Gombak near Kuala Lumpur, ca. 900 feet. Also cited from Singapore).
spectabilis Bernhauer, 1922b: 169 (Edaphus; Type locality: Praslin, Seychelles; [Note: See lectotype designation by Orousset, 1986]. Originally cited from Seychellen: Praslin: Côtes d’Or Estate; Silhouette: Mare aux Cochons).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Kistner, 1962a: 631 (Edaphus; characters).
— Orousset, 1986: 402 (Edaphus; lectotype designation: Praslin, Seychelles).
— Orousset, 1988: 50 (Edaphus; characters; Seychelles).
▴ Distribution: Seychelles.
spiniphallus Orousset, 1988: 43 (Edaphus; Type locality: Madagascar Centre: forêt d’Ambohiboatavo, à l’Est du lac de Mantasoa, 1340 m).
strangulatus Cameron, 1929b: 452 (Edaphus; Type locality: Penang).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1979: 158 (Edaphus; characters).
— Puthz, 1979b: 165 (Edaphus; characters).
▴ Distribution: Malaysia.
subbiformis Puthz, 1992b: 26 (Edaphus; Type locality: Zaire: Kivu, terr. Mwenga, riv. Lubushwa, 650 m).
subcarinatus Puthz, 1979: 118 (Edaphus; Type locality: Süd-Thailand: Kachong Experimental Station, 80 m).
sulciceps Bernhauer, 1929c: 95, 104 (Edaphus; Type locality: Leyte).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1979: 142 (Edaphus; characters).
▴ Distribution: Philippines.
scheerpeltzi Bernhauer, 1929c: 95, 101 (Edaphus; Type locality: Biliran).
sumatrensis Schaufuss, 1887: 110 (Edaphus; Type locality: Sumatra).
— Bernhauer and Schubert, 1911: 188 (Edaphus; catalog).
— Cameron, 1927c: 261 (Edaphus; Fiji; Sumatra; Timor).
— Cameron, 1930d: 330 (Edaphus; Sumatra).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Puthz, 1974r: 88 (Edaphus; lectotype designation; characters).
— Puthz, 1979: 156 (Edaphus; characters).
— Puthz, 1990c: 819 (Edaphus; characters; Sumatra).
▴ Distribution: Indonesia, Fiji.
sybaris Puthz, 1979: 120 (Edaphus; Type locality: India: W. Bengal: Darjeeling, North Point, 1300 m).
szechuanus Puthz, 1975c: 182 (Edaphus; Type locality: China: West-Szechuan, Wassuland, Bezirk San-kiang-kou, 1000 m).
szekessyi Puthz, 1971p: 203 (Edaphus; replacement name for punctatus Székessy).
punctatus Székessy, 1939: 104 (Edaphus; [preoccupied]; Type locality: Neu-Guinea: Marega).
taiwanensis Puthz, 1975c: 182 (Edaphus; Type locality: Formosa: Takao, 300 m).
tamoul Comellini, 1977: 263 (Edaphus; Type locality: Sri Lanka: province Sabaragamuwa, entre Ambepussa et Polganawela).
tasmani Puthz, 1991: 284 (Edaphus; Type locality: Fiji: Vanua Levu: Ndelaikoro, 500 m).
tavakiliani Orousset, 1983b: 117 (Edaphus; Type locality: Mont Grand-Matoury, 7 km S. de Cayenne, alt. 200 m, Guyane française).
taylori Puthz, 1979: 120 (Edaphus; Type locality: Malaya: Upper Gombak Valley near Kuala Lumpur, 1500 feet. Also cited from Singapore).
termitophilus Bernhauer, 1916a: 2 (Edaphus; Type locality: Queensland: Blackal Range).
— Scheerpeltz, 1933: 1204 (Edaphus; catalog).
— Seevers, 1957: 319 (Edaphus; doubtfully termitophilous).
— Puthz, 1978c: 124 (Edaphus; characters; Australia).
▴ Distribution: Australia.
camponoti Oke, 1933: 111 (Edaphellus; variety ? or species ? of melculus; Type locality: Australia: Victoria: Gern Tree Gully, Evelyn, Lilydale).
thai Orousset, 1988c: 184 (Edaphus; Type locality: Thaïlande: env. de Chiang Maï, Doï Inthanon, alt. 2500 m).
tongkingensis Puthz, 1979: 130 (Edaphus; Type locality: Vietnam: Luc yen).
topali Puthz, 1979: 133 (Edaphus; Type locality: India: W. Bengal, Darjeeling District, Ghum, 2200 m).
torresbolloi Puthz, 1985c: 355 (Edaphus; Type locality: Paraguay: alto Paraná, Pto Pte Stroessner).
transvaalensis Puthz, 1989a: 71 (Edaphus; Type locality: South Africa: E. Transvaal: Mariepskop).
tricornis Székessy, 1939: 102 (Edaphus; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen).
tshuapaensis Kistner, 1962a: 631 (Edaphus; Type locality: Congo Republic: Tshuapa, Mabali, near Lake Tumba, 350 m).
— Kistner, 1963: 199 (Edaphus; Congo Republic).
— Puthz, 1974s: 762 (Edaphus; characters).
— Puthz, 1984g: 202 (Edaphus; Zaire).
▴ Distribution: Congo.
tuberculicollis Puthz, 1990b: 280 (Edaphus; Type locality: Neu Guinea: Huon Golf: Sattelberg).
tuberculifer Puthz, 1990a: 197 (Edaphus; Type locality: Côte d’Ivoire: env. Bingerville).
ulcilabris Puthz, 1992b: 28 (Edaphus; Type locality: Zaire: Kivu, Mwenga, 1250 m. Also cited from Rwanda).
unicornis Székessy, 1939: 105 (Edaphus; Type locality: Neu-Guinea: Marega).
uvaensis Comellini, 1977: 266 (Edaphus; Type locality: Sri Lanka: province Uva, Monaragala).
uvirensis Kistner, 1962a: 632 (Edaphus; Type locality: Congo Republic: Kivu, near Uvira, 800 m).
variabilis Puthz, 1974s: 761 (Edaphus; Type locality: Angola: Lunda: Alto Cuílo Cacola, Brüke über den Cuvembo).
— Puthz, 1984g: 201 (Edaphus; characters; Ghana; Tanzania; Uganda).
— Puthz, 1990a: 211 (Edaphus; Ivory Coast; Gabon; Cameroon).
▴ Distribution: Angola, Gabon, Ghana, Ivory Coast, Cameroon, Tanzania, Uganda.
discipunctus Puthz, 1974s: 766 (Edaphus; Type locality: Uganda: zwischen Kampala und Jinja).
— Puthz, 1983f: 173 (Edaphus; Angola; Rhodesia; Kenya).
— Puthz, 1984g: 201 (Edaphus; synonym of variabilis).
— Puthz, 1992b: 31 (Edaphus; synonym of variabilis).
variatus Puthz, 1990a: 215 (Edaphus; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).
variicollis Puthz, 1986c: 311 (Edaphus; Type locality: Rwanda: Cyangugu, Nyakabuye. Also cited from Zaire, Angola, and Cameroon).
varius Puthz, 1990a: 211 (Edaphus; Type locality: Côte d’Ivoire: Agboville: forêt de Yapo, près Yapo-Gare).
vedda Comellini, 1977: 259 (Edaphus; Type locality: Sri Lanka: province Central, Hasalaka près de Weragamtola).
ventriosus Puthz, 1989a: 73 (Edaphus; Type locality: South Africa: Anthrax Camp, Pafuri, Kruger National Park, 15 km W junction Limpopo River and Levuuhu River).
vietnamensis Puthz, 1979: 139 (Edaphus; Type locality: Vietnam: Cuc phuong, Ninh binh).
vietus Puthz, 1974r: 85 (Edaphus; Type locality: Vietnam: Provinz Quang Tri, 7 mi. W. of Dong Ha, Cam Lo).
vulcanicola Puthz, 1979b: 167 (Edaphus; Type locality: Sumatra: Sibajak vulkan).
vulneratus Bernhauer, 1929c: 94, 99 (Edaphus; Type locality: Leyte).
werneri Puthz, 1990d: 33 (Edaphus; Type locality: Rwanda: Rangiro).
wernerianus Puthz, 1990d: 34 (Edaphus; Type locality: Rwanda: Rangiro).
wittmeri Puthz, 1979: 124 (Edaphus; Type locality: India: Assam, Megalaya, Garo Hills, Darugiri, 450 m).
xiao Puthz, 1983g: 163 (Edaphus; Type locality: China: Guanxi [Kwangsi]: Guilin, Yaoshan).
zukwalanus Puthz, 1990d: 39 (Edaphus; Type locality: Ethiopia: Shoa Province: L. Helene S. Mt. Zughala).
zwicki Puthz, 1978c: 121 (Edaphus; Type locality: Victoria: Dandenong Range, Belgrave. Also cited from Queensland).
zwickianus Puthz, 1978c: 125 (Edaphus; Type locality: Victoria: Tolmie near Whitfield).
Euaesthetotyphlus
[1 species; Palaearctic Region]
Euaesthetotyphlus Coiffait and Decou, 1970: 377 (species included: almajensis). Type species: Euaesthetotyphlus almajensis Coiffait and Decou, fixed by original designation.
almajensis Coiffait and Decou, 1970: 379 (Euaesthetotyphlus; Type locality: Roumaine: Ogasul Porcariului, Vallée de la Nera).
Euaesthetus
[41 species; Neotropical, Nearctic, Palaearctic, and Oriental Regions]
Euaesthetus Gravenhorst, 1806: 201 (species included: scaber). Type species: Euaesthetus scaber Gravenhorst, fixed by monotypy.
— Gyllenhal, 1810: 461 (characters).
— Dejean, 1821: 25 (list of species in collection).
— Latreille, 1825: 245 (cited as Euaesthète; characters).
— Berthold, 1827: 331 (characters).
— Le Peletier and Serville, 1828: 483 (characters; notes).
— Zetterstedt, 1828: 91 (characters).
— Curtis, 1829: 30 (catalog; Britain).
— Stephens, 1829a: 24 (catalog; Britain).
— Stephens, 1829: 294 (catalog; Britain).
— Latreille, 1829: 437 (characters).
— C. Sahlberg, 1832: 420 (characters).
— Griffith and Pidgeon, 1832: 294 (characters).
— Dejean, 1833: 66 (list of species).
— Stephens, 1834: 328 (characters; habitat).
— Gistel, 1834: 9 (list of species).
— Westwood, 1838a: 18 (characters; type species).
— Zetterstedt, 1838: 72 (characters).
— Erichson, 1839a: 574 (characters).
— Heer, 1839: 228 (characters).
— Stephens, 1839: 407 (characters).
— Erichson, 1840: 746 (characters).
— Agassiz, 1847: 145 (cited as Evaesthetus).
— Redtenbacher, 1849: 731 (characters).
— Schaum, 1852: 28 (list of species; Europe).
— Lacordaire, 1854: 109 (characters; notes; list of species).
— Fairmaire and Laboulbène, 1856: 597 (characters).
— Gistel, 1856: 389 (list of species).
— Jacquelin du Val, 1857: 53 (characters).
— Kraatz, 1857d: 734 (characters).
— Redtenbacher, 1857: 226 (characters; Austrian species).
— Thomson, 1858: 39 (characters).
— G. Waterhouse, 1858: 27 (catalog of British species).
— Thomson, 1859: 42 (characters; type species: ruficapillus).
— Schaum, 1859: 29 (catalog; European species).
— LeConte, 1861a: 67 (characters).
— Thomson, 1861: 115 (characters).
— Gredler, 1863: 112 (list of species from Tirol).
— LeConte, 1863: 25 (list of North American species).
— Wencker and Silbermann, 1866: 32 (list of species; collecting notes; France).
— Gemminger and Harold, 1868: 631 (catalog).
— Jacquelin du Val, 1868: 76 (catalog; European species).
— Sharp, 1871d: 12 (list of British species).
— Fauvel, 1873: 63 [= 1873a: 221] (characters; key to species).
— Redtenbacher, 1874: 236 (characters; key to Austrian species).
— Fauvel, 1875a: XIV [= 1875b: 216] (catalog).
— Mulsant and Rey, 1878a: 303 (characters; key to French species).
— Heyden, 1880: 79 (list of species of Siberian region).
— Duvivier, 1883: 178 (catalog).
— Casey, 1884a: 19 (key to North American species).
— Fowler, 1888: 324 (notes; key to British species).
— Seidlitz, 1891: 359 (key to some European species).
— Seidlitz, 1891a: 383 (key to some European species).
— Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus).
— Heyden, 1893: 46 (list of species of Siberian region).
— Ganglbauer, 1895: 544 (characters; key to middle European species).
— Heyden, 1896: 35 (list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 151 (list of species of Europe and Caucasus).
— Reitter, 1909: 153 (characters).
— Blatchley, 1910: 458 (characters; key to Indiana species).
— Sainte-Claire Deville, 1910a: 127 (notes; key to and annotated list of species of Seine Basin, France).
— Bernhauer and Schubert, 1911: 188 (world catalog; 21 species).
— Petri, 1912: 63 (list of species of Siebenbürgen).
— Johansen, 1914: 475 (characters; key to Danish species).
— Leng, 1920: 100 (catalog of North American species).
— Winkler, 1925: 357 (catalog for Palaearctic region).
— Porta, 1926: 65 (characters; key to species of Italy).
— Bernhauer, 1928c: 38 (key to North American species).
— Portevin, 1929: 390 (characters; key to French species).
— Roubal, 1930: 348 (catalog; Slovakia).
— Joy, 1932: 14 (key to British species).
— Scheerpeltz, 1933: 1204 (world catalog supplement).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Kloet and Hincks, 1945: 164 (list of British species).
— Tottenham, 1949: 366, 413 (type species: bipunctatus Ljungh = scaber Gravenhorst; notes; checklist of British species).
— Blackwelder, 1952: 152 (type species: scaber).
— Tottenham, 1954: 74 (characters; collecting notes; key to British species).
— Hatch, 1957: 240 (characters; key to species of Pacific Northwest).
— Palm, 1961a: 117 (characters; key to species of Sweden).
— Horion, 1963: 402 (list of central European species).
— Székessy, 1963: 39 (characters).
— Lohse, 1964: 131 (characters; key to central European species).
— Kasule, 1966: 277 (larval characters).
— Scheerpeltz, 1968a: 46 (catalog of Austrian species).
— Scheerpeltz, 1974: 106 (characters).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Moore and Legner, 1975: 262 (catalog; North America north of Mexico).
— Shibata, 1976: 211 (checklist of species of Japan).
— Pope, 1977: 26 (list of British species).
— Topp, 1978: 307 (larval characters).
— Moore and Legner, 1979: 244 (characters; notes).
— Uhlig and Vogler, 1981: 102 (list of some species collected in Germany; habitat and collecting notes).
— Coiffait, 1984f: 358 (characters; key to species of western Palaearctic region).
— Ádám, 1985: 250 (list of species from southeast Hungary).
— Frank, 1986a: 367 (checklist of species of Florida).
— Muona and Viramo, 1986: 22 (list of species of northeastern Finland).
— Segers, 1986: 34 (checklist of species; Belgium).
— Ádám, 1987: 143 (list of four species with collecting notes; Hungary).
— Lohse and Lucht, 1989: 149 (supplement for key to genera of Lohse, 1964).
— Newton, 1990b: 1164, 1170 (larval characters).
— Frank, 1991: 345 (larval characters).
— Silfverberg, 1992: 20 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Siitonen, 1993: 230 (list of Finnish species).
— Boháč, 1993: 45 (list of species of Czech Republic and Slovakia).
— Puthz, 1994e: 392 (key to species of Old World).
— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).
— Terlutter, 1995: 55 (list of some species of Germany).
— Zanetti, 1995: 19 (list of species of Italy).
— M. Hansen, 1996: 102 (list of Danish species).
— Downie and Arnett, 1996: 460 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 48 (list of species from Florida; records for other States listed for each species).
— M. Hansen, Palm, Pedersen, and Runge, 1998: 71 (list of species collected in 1997; Denmark).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (27 Nearctic species; notes; characters in key).
Eristhetus Leach, 1819: 174 (unjustified emendation; cited by Blackwelder, 1952: 152 as an emendation of Euaesthetus because it also had been treated as such by Mannerheim, 1830: 41). Type species: Euaesthetus scaber Gravenhorst, fixed by objective synonymy.
— Mannerheim, 1831: 422, 455 (emendation of Euaesthetus; [Note: Unjustified emendation]).
— Lacordaire, 1835: 438 (characters).
— Laporte, 1840: 184 (characters).
— Lacordaire, 1854: 109 (synonym of Euaesthetus).
— Fauvel, 1873: 63 [= 1873a: 221] (synonym of Euaesthetus).
— Bernhauer and Schubert, 1911: 188 (synonym of Euaesthetus; [Note: Mannerheim, 1830 cited as the author of genus]).
— Coiffait, 1984f: 358 (synonym of Euaesthetus).
albertae Puthz, 1998a: 229 (Euaesthetus; Type locality: Canada: Alberta: 2 mi. S. Ponaka, Battle River marsh).
americanus Erichson, 1840: 747 (Euaesthetus; Type locality: Pensylvania).
— LeConte, 1850: 220 (Euaesthetus; Lake Superior).
— Casey, 1884a: 27 (Euaesthetus; characters; Michigan; Massachusetts; Washington, D.C.; Florida; New York; Delaware).
— Blatchley, 1910: 459 (Euaesthetus; characters; Indiana).
— Bernhauer and Schubert, 1911: 188 (Euaesthetus; catalog).
— Notman, 1919a: 95 (Euaesthetus; Ontario).
— Notman, 1920a: 180 (Euaesthetus; New York).
— Bernhauer, 1928c: 39 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Frank, 1982: 16 (Euaesthetus; list of parasites).
— Campbell and Davies, 1991: 112 (Euaesthetus; USA; Canada).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
— Puthz, 1998a: 242 (Euaesthetus; lectotype designation; synonyms).
▴ Distribution: Canada, USA.
subiridipennis Bernhauer, 1928c: 38 (Euaesthetus; Type locality: Massachusetts: Marion).
— Scheerpeltz, 1933: 1206 (Euaesthetus; catalog).
— Downie and Arnett, 1996: 460 (Euaesthetus; cited as subviridipennis; [Note: Listed from Florida but perhaps erroneously]; USA).
— Puthz, 1998a: 242 (Euaesthetus; synonym of americanus).
andersoni Puthz, 1998a: 233 (Euaesthetus; Type locality: Mexico: Guerrero, 10.3 km SW Filo de Caballo, 2700 m).
atomus Casey, 1884a: 28 (Euaesthetus; Type locality: Sand Point and Cedar Keys, Florida).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 40 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Puthz, 1998a: 225 (Euaesthetus; character figured).
▴ Distribution: USA.
bicoloratus Casey, 1924: 150 (Euaesthetus; Type locality: Florida: near Indian River).
bierigi Puthz, 1998a: 225 (Euaesthetus; Type locality: Costa Rica: Vara Blanca, 2000 m).
bipunctatus Ljungh, 1804: 68 (Stenus; Type locality: in glacie paludis Sjärsjöensis).
— Gyllenhal, 1810: 462 (Euaesthetus; synonym of scaber Gravenhorst).
— C. Sahlberg, 1832: 420 (Eristhetus; synonym of scaber Gravenhorst).
— Erichson, 1839a: 575 (Euaesthetus; synonym of scaber).
— Erichson, 1840: 746 (Euaesthetus; synonym of scaber).
— Kraatz, 1857d: 736 (Euaesthetus; synonym of scaber).
— Fauvel, 1873: 64 [= 1873a: 222] (Euaesthetus; characters; France; Scandinavia; Britain; Germany; Austria; Transbaikal).
— Fauvel, 1873b: 129 [= 1873c: 22] (Euaesthetus; Russia).
— Mulsant and Rey, 1878a: 308 (Euaesthetus; characters; France).
— Ganglbauer, 1895: 545 (Euaesthetus; characters; north and middle Europe).
— Poppius, 1909: 20 (Euaesthetus; Lena Valley, Siberia).
— Reitter, 1909: 153 (Euaesthetus; characters; Germany).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Portevin, 1929: 391 (Euaesthetus; characters; France).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Palm, 1961a: 119 (Euaesthetus; characters; Sweden; Denmark; Norway; Finland).
— Horion, 1963: 402 (Euaesthetus; France; Italy; Dalmatia; Romania; Germany; Austria).
— Lohse, 1964: 131 (Euaesthetus; characters; central Europe).
— Smetana, 1964d: 68 (Euaesthetus; Czechoslovakia).
— Szujecki, 1968a: 722 (Euaesthetus; Poland).
— Jarrige, 1968a: 141 (Euaesthetus; characters).
— Puthz, 1972q: 293 (Euaesthetus; characters).
— Ashworth, 1972: 215 (Euaesthetus; late-glacial subfossil; Britain).
— Puthz, 1973m: 138 (Euaesthetus; characters; notes).
— Outerelo, 1976: 265 (Euaesthetus; Spain).
— Pope, 1977: 26 (Euaesthetus; Britain).
— Coiffait, 1984f: 362 (Euaesthetus; characters).
— Lucht, 1987: 99 (Euaesthetus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 149 (Euaesthetus; characters).
— Nowosad, 1990: 69 (Euaesthetus; collected from mole nest; notes; Poland).
— Sabella and Zanetti, 1991: 274 (Edaphus; notes).
— Boháč, 1993: 45 (Euaesthetus; Slovakia; Czech Republic).
— Puthz, 1994e: 393 (Euaesthetus; characters).
— Sparacio, 1995: 149 (Euaesthetus; characters; notes; Sicily).
— Zanetti, 1995: 19 (Euaesthetus; Italy).
— Ádám, 1996: 242 (Euaeathetus; Hungary).
▴ Distribution: Europe, European and Asia Russia.
scaber Gravenhorst, 1806: 202 (Euaesthetus; Type locality: Brunsvigae).
— Gyllenhal, 1810: 462 (Euaesthetus; characters; Sweden).
— Germar, 1823a: 13 (Euaesthetus; characters; Germany; Sweden).
— Gyllenhal, 1827: 499 (Staphylinus; note).
— Zetterstedt, 1828: 92 (Euaesthetus; characters; Lapland).
— Latreille, 1829: 437 (Euaesthetus; France).
— Mannerheim, 1830: 41 (Eristhetus; Sweden).
— Mannerheim, 1831: 455 (Eristhetus; Sweden).
— C. Sahlberg, 1832: 420 (Eristhetus; characters; Finland).
— Stephens, 1834: 328 (Euaesthetus; characters; England).
— Lacordaire, 1835: 439 (Eristhetus; characters; France).
— Westwood, 1838a: 18 (Euaesthetus; type species).
— Zetterstedt, 1838: 72 (Eristhetus; characters; habitat; Lapland).
— Erichson, 1839a: 574 (Euaesthetus; characters; Germany).
— Heer, 1839: 228 (Euaesthetus; characters; Switzerland).
— Stephens, 1839: 415 (Euaesthetus; characters; England).
— Erichson, 1840: 746 (Euaesthetus; characters; Germany; Sweden).
— Laporte, 1840: 184 (Eristhetus; characters; France).
— Guérin-Méneville, 1844a: 31 (Euaesthetus; Europe).
— Redtenbacher, 1849: 731 (Euaesthetus; characters; Austria).
— Jacquelin Duval, 1850: 50 (Euaesthetus; characters; Germany; France).
— Fairmaire and Laboulbène, 1856: 597 (Euaesthetus; characters; France).
— Kraatz, 1857d: 736 (Euaesthetus; characters; Germany).
— Redtenbacher, 1857: 226 (Euaesthetus; characters; Austria).
— Thomson, 1861: 115 (Euaesthetus; characters; Scandinavia).
— Fauvel, 1873: 64 [= 1873a: 222] (Euaesthetus; synonym of bipunctatus).
— Redtenbacher, 1874: 237 (Euaesthetus; characters; Austria).
— Seidlitz, 1875: 252 (Euaesthetus; characters; Baltic region).
— J. Sahlberg, 1876: 171 (Euaesthetus; Finland).
— Mulsant and Rey, 1878a: 306 (Euaesthetus; characters; France).
— Fowler, 1888: 324 (Euaesthetus; characters; Britain).
— Ganglbauer, 1895: 545 (Euaesthetus; synonym of bipunctatus).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; synonym of bipunctatus).
— Johansen, 1914: 476 (Euaesthetus; characters; Denmark).
— Scheerpeltz, 1933: 1205 (Euaesthetus; synonym of bipunctatus).
— Pope, 1977: 26 (Euaesthetus; synonym of bipunctatus).
— Coiffait, 1984f: 362 (Euaesthetus; synonym of bipunctatus).
elongatus Motschulsky, 1860: 557 (Euaesthetus; Type locality: Baical; [Note: See lectotype designation by Puthz, 1973m]. Originally cited from environs du lac Baical en Daourie).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Puthz, 1973m: 138 (Euaesthetus; lectotype designation: Baikal; synonym of bipunctatus).
— Coiffait, 1984f: 362 (Euaesthetus; synonym of bipunctatus).
mariae Bethe, 1867: 308 (Euaesthetus; Type locality: Dürkheim).
— Fauvel, 1873: 64 [= 1873a: 222] (Euaesthetus; synonym of bipunctatus).
— Redtenbacher, 1874: 237 (Euaesthetus; characters).
— Ganglbauer, 1895: 545 (Euaesthetus; synonym of bipunctatus).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; synonym of bipunctatus).
— Scheerpeltz, 1933: 1205 (Euaesthetus; synonym of bipunctatus).
— Coiffait, 1984f: 362 (Euaesthetus; synonym of bipunctatus).
brevipennis Mulsant and Rey, 1878a: 308 (Euaesthetus; Type locality: Bresse).
— Ganglbauer, 1895: 545 (Euaesthetus; synonym of bipunctatus).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; synonym of bipunctatus).
— Coiffait, 1984f: 362 (Euaesthetus; synonym of bipunctatus).
brevelytratus Outerelo and Gamarra, 1986: 592 (Euaesthetus; Type locality: Puerto de Honduras, Cáceres, entre Hervás y Cabezuela del Valle).
brevipennis Mulsant and Rey, 1878, see: bipunctatus Ljungh, 1804.
brevipennis Casey, 1884, see: duplex Herman, 2001.
chandleri Puthz, 1998a: 234 (Euaesthetus; Type locality: USA: New Hampshire: Strafford Co., Spruce Hole, 3 mi SW Durham).
chantali Puthz, 1998a: 236 (Euaesthetus; Type locality: USA: Massachusetts: Tyngsboro. Also cited from New Hampshire, Iowa, Mississippi, Texas, Ontario, Quebec; Cuba).
columbus Puthz, 1998a: 226 (Euaesthetus; Type locality: Cuba: Caimito del Guayabal).
cubanus Puthz, 1998a: 224 (Euaesthetus; Type locality: Cuba: Aspiro, Sierra del Rosario).
dampfi Bernhauer, 1929h: 190 (Euaesthetus; Type locality: Mexiko: Ajusco Berg, in einer
Seehöhe von 4000 m).
duplex Herman, 2001: 36 (Euaesthetus; replacement name for brevipennis Casey).
brevipennis Casey, 1884a: 20 (Euaesthetus; [preoccupied]; Type locality: Detroit, Michigan).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 39 (Euaesthetus; characters in key).
— Campbell and Davies, 1991: 112 (Euaesthetus; Canada).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
— Herman, 2001: 36 (Euaesthetus; synonym of duplex).
dybasi Puthz, 1998a: 230 (Euaesthetus; Type locality: USA: New York: Hamilton Co., Hope, N. Hope Falls Road. Also cited from New Hampshire, Illinois, Michigan, Wisconsin, Ontario and Quebec).
elongatus Motschulsky, 1860, see: bipunctatus Ljungh, 1804.
fauveli Puthz, 1979b: 172 (Euaesthetus; Type locality: Tonkin).
— J. Li, 1993: 27 (Euaesthetus; China).
— Puthz, 1994e: 392 (Euaesthetus; characters).
▴ Distribution: Vietnam, China.
floridae Casey, 1884a: 21 (Euaesthetus; Type locality: Florida, Tampa, Enterprise).
— Blatchley, 1910: 458 (Euaesthetus; characters; Indiana).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 39 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Campbell and Davies, 1991: 112 (Euaesthetus; Canada).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
— Puthz, 1998a: 240 (Euaesthetus; illustration of aedeagus).
▴ Distribution: USA, Canada.
franki Puthz, 1998a: 241 (Euaesthetus; Type locality: USA: Florida: Highlands Co., Fish-eating Creek, 4 mi. W. of Venus).
frigidus Poppius, 1909, see: laeviusculus Mannerheim, 1844.
fungicola Motschulsky, 1860, see: Gyrophaena.
fulvus Motschulsky, 1860: 558 (Euaesthetus; Type locality: montagnes Oulou Tau dans les parties méridionales des Steppes des Kirguises).
— Fauvel, 1873: 65 [= 1873a: 223] (Euaesthetus; synonym of ruficapillus).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Coiffait, 1984f: 361 (Euaesthetus; characters; Caucasus; southern Russia; Kirghistan).
— Puthz, 1994e: 393 (Euaesthetus; characters).
▴ Distribution: Caucasus, Kyrgyzstan.
ganglbaueri Bernhauer, 1928c: 39 (Euaesthetus; Type locality: Michigan).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Campbell and Davies, 1991: 112 (Euaesthetus; Canada).
— Downie and Arnett, 1996: 460 (Euaesthetus; cited as ganglbauri; USA).
▴ Distribution: USA, Canada.
granulipennis Eppelsheim, 1887: 429 (Euaesthetus; Type locality: Wladiwostok).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— L. Benick, 1924: 257 (Euaesthetus; Siberia).
— Puthz, 1973m: 139 (Euaesthetus; characters; notes; Siberia).
— Puthz, 1994e: 394 (Euaesthetus; characters).
▴ Distribution: Russia.
hammondi Puthz, 1976d: 46 (Euaesthetus; Type locality: Japan: Yodogawa, Osaka. Also cited from China).
— J. Li, 1993: 27 (Euaesthetus; cited as hammionidi; China).
— Puthz, 1994e: 392 (Euaesthetus; characters).
▴ Distribution: Japan, China.
hispanicus Coiffait, 1984b: 256 (Euaesthetus; Type locality: Espagne Picos de Europa, Lhoso, 1900 m).
— Outerelo and Gamarra, 1986: 598 (Euaesthetus; characters).
— Puthz, 1994e: 394 (Euaesthetus; characters).
▴ Distribution: Spain.
immarginatus Erichson, 1840, see: Stenaesthetus.
iripennis Casey, 1884a: 25 (Euaesthetus; Type locality: Ross County, Ohio).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 38 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
▴ Distribution: USA.
japonicus Bernhauer, 1907, see: Edaphus.
juanaensis Puthz, 1998a: 240 (Euaesthetus; Type locality: Cuba: Aspiro, Sierra del Rosario).
kincaidi Hatch, 1957, see: pacificus Fall, 1926.
laeviusculus Mannerheim, 1844a: 195 (Euaesthetus; Type locality: Kavantholm, Finlande).
— Jacquelin Duval, 1850: 50 (Euaesthetus; characters; Finland).
— Fairmaire and Laboulbène, 1856: 598 (Euaesthetus; characters; France).
— Redtenbacher, 1857: 227 (Euaesthetus; characters; Germany).
— Kraatz, 1857d: 736 (Euaesthetus; characters; Austria, France, Germany).
— Thomson, 1861: 116 (Euaesthetus; [Note: Misidentification: Bernhauer and Schubert, 1911: 189 cited ruficollis Thomson as a synonym of laeviusculus; Thomson did not describe the species as new, he attributed it to Erichson who attributed it to Lacordaire]; characters; Scandinavia).
— Thomson, 1867: 286 (Euaesthetus; ruficapillus cited by Thomson, 1860: 116 is misidentification of laeviusculus characters; Scandinavia).
— Fauvel, 1873: 66 [= 1873a: 224] (Euaesthetus; characters; France; Scandinavia; Britain; Germany; Russia; Switzerland).
— Redtenbacher, 1874: 237 (Euaesthetus; characters).
— Seidlitz, 1875: 252 (Euaesthetus; characters; Baltic region).
— J. Sahlberg, 1876: 171 (Euaesthetus; Finland).
— Mulsant and Rey, 1878a: 314 (Euaesthetus; characters; France).
— Fowler, 1888: 325 (Euaesthetus; characters; Britain).
— Ganglbauer, 1895: 546 (Euaesthetus; characters; north and middle Europe).
— Reitter, 1909: 153 (Euaesthetus; characters; Germany).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Johansen, 1914: 477 (Euaesthetus; characters; Denmark).
— L. Benick, 1924: 257 (Euaesthetus; Siberia).
— Portevin, 1929: 391 (Euaesthetus; characters; France).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Palm, 1961a: 119 (Euaesthetus; characters; Sweden; Denmark; Norway; Finland).
— Horion, 1963: 403 (Euaesthetus; Fennoscandia; France; Italy; Romania; Germany; Austria).
— Lohse, 1964: 131 (Euaesthetus; characters; central Europe).
— Smetana, 1964d: 69 (Euaesthetus; Czechoslovakia).
— Szujecki, 1968a: 722 (Euaesthetus; Poland).
— Puthz, 1972q: 293, 294 (Euaesthetus; characters; notes).
— Ashworth, 1972: 215 (Euaesthetus; late-glacial subfossil; Britain).
— Pope, 1977: 26 (Euaesthetus; Britain).
— Coiffait, 1984f: 361 (Euaesthetus; characters; Europe; Britain; Scandinavia; Siberia).
— Lucht, 1987: 99 (Euaesthetus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Campbell and Davies, 1991: 112 (Euaesthetus; USA; Canada).
— Boháč, 1993: 45 (Euaesthetus; Slovakia; Czech Republic).
— Puthz, 1994e: 393 (Euaesthetus; characters).
— Zanetti, 1995: 19 (Euaesthetus; Italy).
— Puthz, 1998a: 231 (Euaesthetus; note).
▴ Distribution: Europe, Russia, Canada, USA.
frigidus Poppius, 1909: 21 (Euaesthetus; Type locality: Lena bei Tjulah-haja).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Puthz, 1972q: 294 (Euaesthetus; synonym of laeviusculus).
lespesii Jacquelin du Val, 1850, see: ruficapillus Lacordaire, 1835.
mandschuricus Puthz, 1976d: 46 (Euaesthetus; Type locality: China: Heilungjiang, Harbin).
— J. Li, 1993: 27 (Euaesthetus; China).
— Puthz, 1994e: 394 (Euaesthetus; characters).
▴ Distribution: China.
mariae Bethe, 1867, see: bipunctatus Ljungh, 1804.
marionensis Bernhauer, 1928c: 40 (Euaesthetus; Type locality: Massachusetts: Marion).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Downie and Arnett, 1996: 460 (Euaesthetus; USA).
▴ Distribution: USA.
mundulus Casey, 1884a: 23 (Euaesthetus; Type locality: Cambridge, Massachusetts).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 40 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
— Puthz, 1998a: 243 (Euaesthetus; synonyms).
▴ Distribution: USA.
robustulus Casey, 1884a: 24 (Euaesthetus; Type locality: Cambridge, Massachusetts).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 40 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Downie and Arnett, 1996: 460 (Euaesthetus; characters; USA).
— Puthz, 1998a: 243 (Euaesthetus; synonym of mundulus).
neomexicanus Fall, 1907, see: similis Casey, 1884.
nesterovi Petrenko, 1984, see: Ctenomastax.
newtoni Puthz, 1998a: 231 (Euaesthetus; Type locality: Mexico: Jalisco, 10 mi. W. Atenquique, 6700 feet).
nitidulus Sharp, 1874: 78 (Euaesthetus; Type locality: Nagasaki).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Nakane, 1963a: 87 (Euaesthetus; characters; habitus photograph; Japan).
— Puthz, 1994e: 392 (Euaesthetus; characters).
▴ Distribution: Japan.
pacificus Fall, 1926a: 62 (Euaesthetus; Type locality: Scow Bay, Alaska).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Campbell and Davies, 1991: 112 (Euaesthetus; Canada).
▴ Distribution: USA, Canada.
kincaidi Hatch, 1957: 240 (Euaesthetus; Type locality: Washington: Seaview).
— Puthz, 1980j: 21 (Euaesthetus; synonym of pacificus).
— Puthz, 1998a: 243 (Euaesthetus; synonym of pacificus).
pugetensis Hatch, 1957: 241 (Euaesthetus; Type locality: Washington: Cedar Mt. Also cited from British Columbia).
— Campbell and Davies, 1991: 112 (Euaesthetus; USA; Canada).
— Puthz, 1998a: 227 (Euaesthetus; illustration of aedeagus).
▴ Distribution: Canada, USA.
pullus Thomson, 1857, see: ruficapillus Lacordaire, 1835.
punctatus Casey, 1884a: 21 (Euaesthetus; Type locality: Crescent City, Florida).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 39 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
▴ Distribution: USA.
robustulus Casey, 1884, see: mundulus Casey, 1884.
ruficapillus Lacordaire, 1835: 439 (Eristhetus; Type locality: Paris).
— Erichson, 1839a: 575 (Euaesthetus; characters; Germany).
— Heer, 1839: 228 (Euaesthetus; characters; Switzerland).
— Erichson, 1840: 747 (Euaesthetus; characters; Germany; France).
— Laporte, 1840: 184 (Eristhetus; characters; France).
— Redtenbacher, 1849: 731 (Euaesthetus; characters; Austria).
— Jacquelin Duval, 1850: 50 (Euaesthetus; characters; Germany; France).
— Fairmaire and Laboulbène, 1856: 597 (Euaesthetus; characters; France).
— Thomson, 1857: 234 (Euaesthetus; characters; Sweden).
— Kraatz, 1857d: 737 (Euaesthetus; characters; Germany).
— Thomson, 1859: 42 (type species of Euaesthetus).
— Hochhuth, 1862: 80 (Euaesthetus; Caucasus; Ukraine).
— Thomson, 1867: 287 (Euaesthetus; characters; Scandinavia).
— Fauvel, 1873: 65 [= 1873a: 223] (Euaesthetus; characters; France; Scandinavia; Britain; Germany; Russia; Austria; Italy; Caucasus).
— Redtenbacher, 1874: 237 (Euaesthetus; characters; Austria).
— Seidlitz, 1875: 252 (Euaesthetus; characters; Baltic region).
— J. Sahlberg, 1876: 171 (Euaesthetus; Finland).
— Mulsant and Rey, 1878a: 311 (Euaesthetus; characters; France).
— Fowler, 1888: 325 (Euaesthetus; characters; Britain).
— Ganglbauer, 1895: 545 (Euaesthetus; characters; north and middle Europe; Italy; Caucasus).
— Reitter, 1909: 153 (Euaesthetus; characters; Germany).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Johansen, 1914: 477 (Euaesthetus; characters; Denmark).
— Portevin, 1929: 391 (Euaesthetus; characters; France).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Palm, 1961a: 119 (Euaesthetus; characters; Sweden; Denmark; Norway; Finland).
— Horion, 1963: 403 (Euaesthetus; France to Poland-Russia; Italy; Germany; Austria).
— Lohse, 1964: 131 (Euaesthetus; characters; central Europe).
— Smetana, 1964d: 68 (Euaesthetus; Czechoslovakia).
— Puthz, 1972q: 293 (Euaesthetus; characters).
— Outerelo, 1976a: 260 (Euaesthetus; notes; Spain).
— Pope, 1977: 26 (Euaesthetus; Britain).
— Frank, 1982: 16 (Euaesthetus; list of parasites).
— Coiffait, 1984f: 359 (Euaesthetus; characters; Europe; Turkey; Caucasus; Siberia).
— Lucht, 1987: 99 (Euaesthetus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Boháč, 1993: 45 (Euaesthetus; Slovakia; Czech Republic).
— Puthz, 1994e: 393 (Euaesthetus; characters).
— Zanetti, 1995: 19 (Euaesthetus; Italy).
— Bordoni, 1995: 82, 83 (Euaesthetus; habitus; ecological notes; Italy).
— Owen, 1997a: 149 (Euaesthetus; collecting sites; Ireland).
▴ Distribution: Europe, Turkey, Caucasus, Russia.
lespesii Jacquelin du Val, 1850: 48, 51 (Euaesthetus; Type locality: Toulouse; environs de Paris).
— Fairmaire and Laboulbène, 1856: 598 (Euaesthetus; characters; France).
— Mulsant and Rey, 1878a: 312 (Euaesthetus; synonym of ruficapillus).
— Ganglbauer, 1895: 545 (Euaesthetus; cited as lespesi; synonym of ruficapillus).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; cited as lespesi; synonym of ruficapillus).
— Coiffait, 1984f: 360 (Euaesthetus; cited as lespesi; synonym of ruficapillus).
pullus Thomson, 1857: 235 (Euaesthetus; Type locality: Ringsjön i Skåne).
— Thomson, 1861: 116 (Euaesthetus; characters; Scandinavia).
— Fauvel, 1867c: 114 [= 1868b: 213] (Euaesthetus; synonym of ruficapillus).
— Thomson, 1867: 287 (Euaesthetus; synonym of ruficapillus).
— Mulsant and Rey, 1878a: 312 (Euaesthetus; synonym of ruficapillus).
— Ganglbauer, 1895: 545 (Euaesthetus; synonym of ruficapillus).
— Bernhauer and Schubert, 1911: 190 (Euaesthetus; synonym of ruficapillus).
ruficollis Motschulsky, 1860: 558 (Euaesthetus; Type locality: Transbaicalie).
— Fauvel, 1873: 64 [= 1873a: 222] (Euaesthetus; synonym of bipunctatus).
— J. Sahlberg, 1880: 97 (Euaesthetus; characters; Siberia).
— Ganglbauer, 1895: 545 (Euaesthetus; synonym of bipunctatus).
— Poppius, 1909: 20 (Euaesthetus; Lena Valley, Siberia).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; synonym of bipunctatus).
— Puthz, 1973m: 139 (Euaesthetus; valid species; characters; notes; Siberia).
— Puthz, 1994e: 393 (Euaesthetus; characters).
— Ryabukhin, 1999: 42 (Euaesthetus; catalog; habitat; distribution; northeastern Russia).
▴ Distribution: Asian Russia.
wuorentausi Bernhauer, 1927b: 90 (Euaesthetus; Type locality: Wuorentaus bei Dudinka).
— Scheerpeltz, 1933: 1206 (Euaesthetus; catalog).
— Puthz, 1973m: 139 (Euaesthetus; synonym of ruficollis).
scaber Gravenhorst, 1806, see: bipunctatus Ljungh, 1804.
schawalleri Puthz, 1994e: 389 (Euaesthetus; Type locality: Russland: Khabarovsk Province, SE Boitsovo, 12 km NE Bikin, 250–350 m).
similis Casey, 1884a: 22 (Euaesthetus; Type locality: Tallahassee, Florida).
— Bernhauer and Schubert, 1911: 190 (Euaesthetus; catalog).
— Bernhauer, 1928c: 40 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1206 (Euaesthetus; catalog).
— Puthz, 1974: 11 (Euaesthetus; characters; Illinois; Florida; Texas; New Mexico; Louisiana; Maryland; North Carolina).
▴ Distribution: USA.
texanus Casey, 1884a: 26 (Euaesthetus; Type locality: Columbus, Texas).
— Bernhauer and Schubert, 1911: 190 (Euaesthetus; catalog).
— Bernhauer, 1928c: 40 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1206 (Euaesthetus; catalog).
— Puthz, 1974: 11 (Euaesthetus; synonym of similis).
neomexicanus Fall, 1907: 219 (Euaesthetus; Type locality: Las Vegas, New Mexico).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Bernhauer, 1928c: 39 (Euaesthetus; characters in key).
— Scheerpeltz, 1933: 1205 (Euaesthetus; catalog).
— Puthz, 1974: 11 (Euaesthetus; synonym of similis).
smetanai Puthz, 1998a: 238 (Euaesthetus; Type locality: USA: Louisiana: Calcasieu Pa., Choupique Bayon, 7 mi N. Hackleberry. Also cited from Texas and Mississippi).
spinicollis Coiffait, 1983d: 174 (Euaesthetus; Type locality: Nepal: Ramche, 1430 m).
subiridipennis Bernhauer, 1928, see: americanus Erichson, 1840.
superlatus Peyerimhoff, 1937: 44 (Euaesthetus; Type locality: France: Alpes-Maritimes: col. de Mulinès au-dessus de Beuil, vers la cote 2000).
— Jarrige, 1968a: 139 (Euaesthetus; characters; notes; France).
— Puthz, 1969i (Euaesthetus; characters; notes; Germany; Austria; France; Yugoslavia).
— Puthz, 1972q: 291, 293 (Euaesthetus; characters; Hungary).
— Puthz, 1976j: 189 (Euaesthetus; distribution in Germany).
— Coiffait, 1984f: 363 (Euaesthetus; characters; France; Austria; Germany; Hungary).
— Lohse and Lucht, 1989: 149 (Euaesthetus; characters).
— Boháč, 1993: 45 (Euaesthetus; Slovakia; Czech Republic).
— Puthz, 1994e: 394 (Euaesthetus; characters).
— Assing, Frisch, Kahlen, et al., 1998: 131 (Euaesthetus; note).
— Assing, 2001: 75 (Euaesthetus; collected from xerothermous site; Germany).
▴ Distribution: Austria, France, Germany, Hungary, Yugoslavia.
texanus Casey, 1884, see: similis Casey, 1884.
wuorentausi Bernhauer, 1927, see: ruficollis Motschulsky, 1860.
Macroturellus
[1 species; Ethiopian Region]
Macroturellus Orousset, 1987: 219 (species included: pulcher). Type species: Macroturellus pulcher Orousset, fixed by original designation and monotypy.
pulcher Orousset, 1987: 222 (Macroturellus; Type locality: Cameroun: Yaoundé).
Neocoiffaitia
[1 species; Madagascan Region]
Neocoiffaitia Orousset, 1988: 69 (species included: anosyana). Type species: Neocoiffaitia anosyana Orousett, fixed by original designation and monotypy.
anosyana Orousset, 1988: 69 (Neocoiffaitia; Type locality: Madagascar Est: massif de l’Andohahelo, 1600 m).
Octavius
[194 species; Palaearctic, Ethiopian, Madagascan, Oriental, Oceanic, and Neotropical Regions]
Octavius Fauvel, 1873: 62 [= 1873a: 220] (species included: pyrenaeus, insularis, grandiceps).
Type species: Octavius pyrenaeus Fauvel, fixed by subsequent designation by R. Lucas, 1920: 449.
— Fauvel, 1875a: XIV [= 1875b: 216] (catalog).
— Mulsant and Rey, 1878a: 316 (characters; key to French species).
— Duvivier, 1883: 179 (catalog).
— Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus).
— Heyden, Reitter, and Weise, 1906: 151 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1911: 187 (world catalog; 7 species).
— Winkler, 1925: 357 (catalog for Palaearctic region).
— Porta, 1926: 65 (characters; key to species of Italy).
— Portevin, 1929: 390 (characters; key to French species).
— Scheerpeltz, 1933: 1202 (world catalog supplement).
— Normand, 1935: 364 (list of species of Tunisia).
— Blackwelder, 1952: 266 (type species).
— Coiffait, 1958a (revision of European species; characters; ecology; distribution; checklist of species).
— Kistner, 1960a: 117 (notes; key to subsaharan species).
— Székessy, 1963: 38 (characters).
— Coiffait, 1965f: 100 (key to species groups; list of species for each group).
— Bordoni, 1973j: 676 (list of species from islands near Sicily).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Scheerpeltz, 1974: 104 (characters).
— Puthz, 1977h: 106 (characters; notes).
— Puthz, 1977h: 108, 111, 122, 123 (key to species of Neotropical region and Africa; list of species).
— Puthz, 1979c: 50 (key to African species).
— Puthz, 1980j: 22 (discussion; synonyms; characters).
— Puthz, 1980g (Octavius; notes; Nepal; Philippines).
— Orousset, 1983: 48 (comments on and characters for infrageneric divisions).
— Coiffait, 1984f: 369 (characters; key to species of western Palaearctic region).
— Puthz, 1985d: 703 (key to species of India and Himalayan region).
— Shibata, 1986a: 121 (supplemental list of species from Taiwan).
— Puthz, 1986f: 194 (key to species of Africa south of the Sahara).
— Orousset, 1988: 71 (characters; key to species of Madagascar; notes on world distribution and habitat).
— Orousset, 1990a: 124 (key to species of Corsica).
— Zanetti, 1995: 19 (list of species of Italy).
Turellus Sharp, 1876: 423 (species included: batesi). Type species: Turellus batesi Sharp, fixed by monotypy.
— Duvivier, 1883: 206 (catalog).
— Bernhauer and Schubert, 1911: 187 (world catalog; 1 species).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 400 (type species: batesi).
— Kistner, 1962: 101 (characters).
— Kistner, 1967: 55 (notes).
— Scheerpeltz, 1974: 104 (characters).
— Puthz, 1980j: 26 (synonym of Octavius).
— Orousset, 1988: 71 (synonym of Octavius).
Anillosthetus Mulsant and Rey, 1876: 146 [= 1877: 2] ([Note: Nomen nudum; the genus was cited without species (Article 12.2) and although a case might be made that characters were presented (see Blackwelder, 1952: 53), the only included species (corsicus by Mulsant and Rey, 1878a: 318) was cited first as a synonym of insularis and is also unavailable (Article 11.6)]).
Exoctavius Bierig, 1934b: 221 (species included: bermudezi). Type species: Exoctavius bermudezi Bierig, by original designation and monotypy.
— Blackwelder, 1943: 227 (type species: bermudezi).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 163 (type species: bermudezi).
— Puthz, 1973k: 68 (characters; notes).
— Puthz, 1977h: 106 (synonym of Octavius).
— Puthz, 1980j: 26 (synonym of Octavius).
— Orousset, 1988: 71 (synonym of Octavius).
Thaxterius Bernhauer, 1935f: 213 (species included: sulcicollis). Type species: Thaxterius sulcicollis Bernhauer, by original designation and monotypy.
— Blackwelder, 1952: 384 (type species: sulcicollis).
— Kistner, 1960a: 117 (synonym of Octavius).
— Scheerpeltz, 1974: 106 (characters).
— Puthz, 1977h: 106 (synonym of Octavius).
— Puthz, 1980j: 26 (synonym of Octavius).
— Orousset, 1988: 71 (synonym of Octavius).
Doletica Cameron, 1938: 1 (species included: bicolor). Type species: Doletica bicolor Cameron, fixed by monotypy.
— Cameron, 1950f: 112 (characters).
— Blackwelder, 1952: 131 (type species: bicolor).
— Kistner, 1961: 13 (characters; key to species; type species: bicolor).
— Kistner, 1962: 101 (synonym of Turellus).
— Scheerpeltz, 1974: 104 (characters).
— Puthz, 1980j: 26 (synonym of Octavius).
— Orousset, 1988: 71 (synonym of Octavius).
Traegardhius Scheerpeltz, 1974: 93, 105 (species included: natalensis). Type species: Traegardhius natalensis Scheerpeltz, fixed by original designation.
Brinckiellus Scheerpeltz, 1974: 98, 105 (species included: capensis). Type species: Brinckiellus capensis Scheerpeltz, fixed by original designation.
Nepalopeplus Coiffait, 1982: 129 (species included: himalayicus). Type species: Nepalopeplus himalayicus Coiffait, fixed by original designation and monotypy.
abditus Orousset, 1990a: 122 (Octavius; Type locality: Corse du Sud: source entre Chiova d’Asinu et Suartone, au Nord du golfe de Santa Manza, alt. 120 m).
abruzzensis Coiffait, 1965, see: vitalei Bernhauer, 1908.
africanus Cameron, 1950f: 119 (Octavius; Type locality: Angola: Dundo: Luachimo Forest).
— Kistner, 1960a: 118 (Octavius; characters; Angola).
— Puthz, 1977h: 111 (Octavius; characters; Angola).
— Puthz, 1979c: 53 (Octavius; characters).
— Puthz, 1986f: 199 (Octavius; characters).
▴ Distribution: Angola.
alzadae Kistner, 1963: 201 (Octavius; Type locality: Congo Republic: Katanga, Albertville territory, Bendera, 1000 m).
— Puthz, 1977h: 111 (Octavius; characters; Zaire).
— Puthz, 1979c: 56 (Octavius; characters).
— Puthz, 1986f: 198 (Octavius; characters).
▴ Distribution: Congo.
ambitsika Orousset, 1988: 120 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt d’Ambitsika, 2100 m).
amindramiova Orousset, 1988: 90 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, Amindramiova, 1950 m).
andohahelo Orousset, 1988: 113 (Octavius; Type locality: Madagascar Est, massif de l’Andohahelo).
andringitra Orousset, 1988: 104 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, pic Boby, 2650 m).
anjavidilava Orousset, 1988: 100 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, Anjavidilava, 1900 m).
ankaratra Orousset, 1988: 140 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt de Betay, 2100 m).
anophthalmus Puthz, 1991b: 83 (Octavius; Type locality: Ghana: Central Region: Kibi, 274 m, 6°10′N, 0°34′W).
apicalis Orousset, 1983, see: spiniphallus Orousset, 1981.
apicidens Puthz, 1985d: 701 (Octavius; Type locality: Indien: West Bengal: Darjeeling district: entre Ghoom et Lopchu, à 13 km de Ghoom, versant nord, 2000 m).
armatus Orousset, 1988: 99 (Octavius; Type locality: Madagascar Centre: entre Ihosy et Ivohibe, col d’Ivohibory, 1000 m).
asiaticus Coiffait, 1958a: 90 (Octavius; Type locality: Turquie d’Asie: Karamursel, Villayet d’Izmit).
— Puthz, 1977h: 122 (Octavius; characters; Turkey).
— Coiffait, 1984f: 391 (Octavius; characters; Turkey).
▴ Distribution: Turkey.
assamensis Puthz, 1985, see: flavescens Kistner, 1961.
atomus Puthz, 1986f: 186 (Octavius; Type locality: Südafrika: Cape Province, Garden of Eden, 34°02′S, 23°12′E).
bacillus Puthz, 1986f: 187 (Octavius; Type locality: Südafrika: Southern Cape: Keurboom-strand near Plettenberg Bay, 34°00′S, 23°27′E).
balazuci Jarrige, 1964: 189 (Octavius; Type locality: dans la terre de la terrasse bordant
l’entrée de la grotte de Baume Grenas, commune de Ruoms, Ardèche).
— Orousset, 1981: 21 (Octavius; characters).
— Coiffait, 1984f: 395 (Octavius; characters; France).
— Orousset, 1984a: 129 (Octavius; collecting notes; France).
— Orousset, 1987b: 17 (Octavius; France).
▴ Distribution: France.
baloghi Puthz, 1977h: 119 (Octavius; Type locality: New Guinea NE: Wau, Kilolo Creek, 1000 m).
baloghianus Puthz, 1991a: 256 (Octavius; Type locality: New Guinea NE: Angoram).
batesi Sharp, 1876: 424 (Turellus; Type locality: Ega).
— Bernhauer and Schubert, 1911: 187 (Turellus; catalog).
— Puthz, 1980j: 24, 27 (Octavius; characters; Peru).
— Orousset, 1986b: 20 (Octavius; Surinam).
▴ Distribution: Brazil, Peru, Surinam.
berardi Lavagne, 1917, see: grandiceps Mulsant and Rey, 1853.
bermudezi Bierig, 1934b: 222 (Exoctavius; Type locality: Sierra del Rosario, rio Taco-Taco).
— Blackwelder, 1943: 227 (Exoctavius; Cuba).
— Puthz, 1977h: 108 (Octavius; characters).
▴ Distribution: Cuba.
bessoni Orousset, 1986d: 100 (Octavius; Type locality: Odita Lezia, gorges de Kakoueta, alt. 850 m, commune de Saint-Engrâce, Pyrénées-Atlantiques).
besucheti Puthz, 1977h: 121 (Octavius; Type locality: Türkei: Bolu, Düzce-Akcakoca, 400 m).
betschi Orousset, 1988: 135 (Octavius; Type locality: Madagascar Centre: tampoketsa d’Ambohitantely, au Nord d’Ankazobe, 1500 m).
betsileo Orousset, 1988: 111 (Octavius; Type locality: Madagascar Centre: pays Betsileo, Ambatofitoharana, RN 7, pk 292, 1600 m).
bicolor Cameron, 1938: 1 (Doletica; Type locality: Belgian Congo: Eala).
— Kistner, 1961: 17 (Doletica; characters; Belgian Congo).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1984h: 563 (Octavius; characters; Zaire).
— Puthz, 1986f: 196 (Octavius; characters).
▴ Distribution: Congo.
bifidus Puthz, 1979c: 57 (Octavius; Type locality: Elfenbeinküste: Man, Cascade).
biroi Puthz, 1977h: 114 (Octavius; Type locality: Neu Guinea: Seleo Berlinhafen).
bordei Sainte-Claire Deville, 1911: 214 (Octavius; Type locality: environs de Grasse, Alpes-Martimes).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1958a: 94 (Octavius; characters; Pyrenees).
— Coiffait, 1958a: 42 (Octavius; characters).
— Hervé, 1964: 294 (Octavius; France).
— Orousset, 1983c: 136 (Octavius; notes; holotype is male).
— Coiffait, 1984f: 404 (Octavius; characters; France).
— Orousset, 1987b: 19 (Octavius; France).
— Zanetti, 1995: 19 (Octavius; Italy ?).
▴ Distribution: France.
boryi Orousset, 1988: 107 (Octavius; Type locality: Madagascar Centre: massif de l’Andrin-gitra, vallée située sous le pic Bory, 2500 m).
breviceps Jarrige, 1968: 876 (Octavius; Type locality: Madagascar-Est, Italaviana, Fanovana).
brevipennis Cameron, 1956: 182 (Doletica; Type locality: Ruanda: Tshuruyaga, forêt du Rugege, 2400 m, terr. Astrida).
— Kistner, 1961: 17 (Doletica; characters; Ruanda).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 197 (Octavius; characters).
▴ Distribution: Rwanda.
brincki Puthz, 1976e: 240 (Turellus; replacement name for capensis Scheerpeltz).
— Puthz, 1976e: 240 (Turellus; characters).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 199 (Octavius; characters).
▴ Distribution: South Africa.
capensis Scheerpeltz, 1974: 100 (Brinckiellus; [preoccupied]; Type locality: Cape Prov.: Cape Peninsula, Hout Bay Skorsteenkop).
bryophilus Orousset, 1988: 106 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, cirque Boby, 2550 m).
capdeviellei Orousset, 1979: 127 (Octavius; Type locality: grotte de Gourgue, commune d’Arbas près d’Aspet, Haute-Garonne).
capensis Kistner, 1967: 59 (Turellus; Type locality: Republic of South Africa, Cape Province, Beaufort District, Katberg).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 200 (Octavius; characters).
▴ Distribution: South Africa.
capensis Scheerpeltz, 1974, see: brincki Puthz, 1976.
capitocleftus Kistner, 1961: 19 (Doletica; Type locality: Congo: Parc National de la Garamba).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 195 (Octavius; characters).
▴ Distribution: Congo.
carpetanicus Outerelo, 1989: 522 (Octavius; Type locality: España: Villanueva de la Vera, Cáceres, Extremadura).
castrii Coiffait, 1963c: 182 (Octavius; Type locality: Italie: province de Vicenza, Monte Spitz dans les Monte Lessini, Recoaro).
— Pace, 1983: 106, 107 (Octavius; characters; Italy).
— Coiffait, 1984f: 396 (Octavius; characters; Italy).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Italy.
euganensis Coiffait, 1972a: 79 (Octavius; Type locality: Colli Euganei, Italie du Nord).
— Puthz, 1972i: 129 (Octavius; Italy).
— Pace, 1983: 107 (Octavius; subspecies of castrii; characters; Italy).
— Coiffait, 1984f: 396 (Octavius; cited as valid species; characters; Italy).
— Zanetti, 1995: 19 (Octavius; subspecies of castrii; Italy).
catalanus Coiffait, 1958a: 91 (Octavius; Type locality: Barcelone: Parc de Tibidabo).
catharus Coiffait, 1963c: 178 (Octavius; Type locality: Ariège, Montferrier, vers 900 m à la lisière de la Forêt de Montmenier, sur le versant Nord d’un contrefort du Pic Saint-Barthélemy).
cerberus Coiffait, 1958a: 86 (Octavius; Type locality: Pyrénées Orientales: Cerbère).
— Coiffait, 1984f: 393 (Octavius; characters; Pyrenees).
— Orousset, 1987b: 17 (Octavius; France).
▴ Distribution: France, Spain.
coecus Jarrige, 1968: 874 (Octavius; Type locality: Madagascar-Centre, Mamjakatompo, Ankaratra).
coiffaiti Puthz, 1985b: 299 (Octavius; Type locality: Gabon: Belinga).
coiffaitianus Orousset, 1987b: 12 (Octavius; Type locality: Alos, au sud de Saint-Girons, Ariège).
comellinii Coiffait, 1963c: 186 (Octavius; Type locality: Espagne: Province de Tarragone, Sierra de Prades, Montréal, à l’Est de la Vallée du Rio Francoli).
confusus Coiffait, 1958a: 85 (Octavius; Type locality: Basses-Pyrénées: Gotein).
— Coiffait, 1959a: 44 (Octavius; France).
— Puthz, 1977h: 122 (Octavius; notes; Pyrenees, Spain).
— Coiffait, 1984f: 390 (Octavius; characters; Pyrenees).
— Orousset, 1986d: 102 (Octavius; characters; France).
— Orousset, 1987b: 17 (Octavius; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France, Spain.
corsicus Orousset, 1983: 54 (Octavius; Type locality: Petreto-Bicchisano, route du Col de Saint-Eustache, alt. 800 m).
— Coiffait, 1984f: 403 (Octavius; characters; Corsica).
— Orousset, 1990a: 126 (Octavius; characters).
▴ Distribution: Corsica.
cossyrensis Coiffait, 1965, see: vitalei Bernhauer, 1908.
crenicollis Fauvel, 1876a: 59 [= 1876: 244] (Octavius; Type locality: Pyrénées Orientales, La Massane).
— Mulsant and Rey, 1878a: 321 (Octavius; characters; France).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Portevin, 1929: 390 (Octavius; characters; France).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1958a: 94 (Octavius; characters; Pyrenees).
— Coiffait, 1984f: 403 (Octavius; characters; Pyrenees).
— Orousset, 1987b: 19 (Octavius; France).
— Orousset, 1991: 167 (Octavius; notes; Pyrenees).
▴ Distribution: France, Spain.
croaticus Puthz, 1972j: 127 (Octavius; Type locality: Jugoslawien: Kroatien: Agram [Zagreb]).
— Puthz, 1977h: 122 (Octavius; notes).
— Coiffait, 1984f: 398 (Octavius; characters; Croatia).
— Orousset, 1991: 168 (Octavius; Slovenia; Croatia).
▴ Distribution: Croatia, Slovenia.
crucialis Kistner, 1967: 57 (Turellus; Type locality: Republic of South Africa, Natal, Bergville District, Drakensberg, Ogalweni Forest, 1800 m).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 200 (Octavius; characters).
▴ Distribution: South Africa.
cruciatus Orousset, 1988: 129 (Octavius; Type locality: Madagascar Centre: forêt d’Ambo-hiboatavo, à l’Ent du lac de Mantasoa).
dajozi Orousset, 1988: 125 (Octavius; Type locality: Madagascar Est: région de Périnet, forêt de la Grande Ile, 900 m).
debilis Orousset, 1988: 122 (Octavius; Type locality: Madagascar Centre: massif de l’Anka-ratra, forêt aux environs du lac Froid, 1800 m).
deharvengi Orousset, 1981: 17 (Octavius; Type locality: grotte proche du hameau de Rieu-fourcand en forêt de Bélesta, alt. 700 m, Ariège).
delicatus Puthz, 1979c: 54 (Octavius; Type locality: Elfenbeinküste: Abidjan, Parc national du Banco).
dentatus Puthz, 1985d: 701 (Octavius; Type locality: Indien: West Bengal: Darjeeling district: 13 km au nord de Ghoom, route pour Bijanbari, 1500 m).
denticulatus Puthz, 1985d: 703 (Octavius; Type locality: Nepal: Kosi: District de Sankhuwa-sawa, Col au nord-est de Mangmaya, 2300 m).
descarpentriesi Jarrige, 1968: 877 (Octavius; Type locality: Madagascar-Centre, Cirque Boby, 2500 m, Andringitra).
deuvei Orousset, 1983: 58 (Octavius; Type locality: Sainte-Lucie-de-Tallano, alt. 350 m).
— Coiffait, 1984f: 401 (Octavius; characters; Corsica).
— Orousset, 1990a: 127 (Octavius; characters).
▴ Distribution: Corsica.
distortus Orousset, 1988: 121 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt de Manjakatompo, 1650 m).
doderoi Coiffait, 1965f: 100 (Octavius; Type locality: Sardaigne, Teulada).
— Coiffait, 1984f: 405 (Octavius; characters; Sardinia).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Sardinia.
dolosus Orousset, 1988: 98 (Octavius; Type locality: Madagascar Est: forêt de Mahabo au Sud-Est de Maropaika).
doriai Poggi, 1977: 301 (Octavius; Type locality: Isola del Giglio, Arcipelago toscano, prov. di Grosseto).
— Coiffait, 1984f: 406 (Octavius; characters; Giglio Island).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Italy.
dybasi Puthz, 1977h: 112 (Octavius; Type locality: Palau Inseln: Angaur, 6°55′N, 134°09′O).
edentulus Puthz, 1985d: 700 (Octavius; Type locality: Indien: West Bengal: Darjeeling district: Mahanadi près de Kurseong, versant sud, 1000 m).
endrodyi Jacobson and Kistner, 1974: 90 (Octavius; Type locality: Ghana: Ashanti Region, Kumasi).
— Puthz, 1977h: 111 (Octavius; characters; Ghana).
— Puthz, 1979c: 57 (Octavius; characters).
— Puthz, 1986f: 198 (Octavius; characters; Ghana; Ivory Coast).
▴ Distribution: Ghana, Ivory Coast.
endroedyanus Puthz, 1986f: 190 (Octavius; Type locality: Südafrika: Natal: Cathedral Peak, 28°57′S, 29°12′E).
euganensis Coiffait, 1972, see: castrii Coiffait, 1963.
exilis Orousset, 1988: 112 (Octavius; Type locality: Madagascar Est: massif de l’Andohahelo).
exspectatus Puthz, 1984h: 569 (Octavius; Type locality: Zaire: terr. Bikoro, 350 m, LacTumba).
falcifer Orousset, 1988: 121 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt de Betay, 2100 m).
fallax Orousset, 1988: 91 (Octavius; Type locality: Madagascar Est: massif de l’Andohahelo).
fauveli Puthz, 1977h: 115 (Octavius; Type locality: New Guinea NE: Lae).
ferus Orousset, 1988: 141 (Octavius; Type locality: Madagascar Centre: forêt de Vanjamanitra, à 8 km au Sud-Est d’Anjozorobe, 1380 m).
flavescens Kistner, 1961: 20 (Doletica; Type locality: Sumatra).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1980g: 749 (Octavius; notes; Philippines; Formosa; India).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1985d: 704 (Octavius; characters).
— Puthz, 1986f: 197 (Octavius; characters).
▴ Distribution: India, Indonesia, Philippines, Taiwan.
assamensis Puthz, 1985d: 698 (Octavius; subspecies of flavescens; Type locality: Indien: Assam: Manas, 200 m).
minor Puthz, 1985d: 698 (Octavius; subspecies of flavescens; Type locality: Indien: Assam: Manas, 200 m).
franzi Coiffait, 1963, see: tarragonensis Coiffait, 1963.
frater Puthz, 1979c: 56 (Octavius; Type locality: Zaire: Katanga, Bendera, Terr. Albertville
[Kalemie], 1000 m).
gamai Puthz, 1986f: 189 (Octavius; Type locality: Südafrika: Natal: Cathedral Peak, 28°27′S, 29°12′E).
gavarrensis Coiffait, 1963c: 186 (Octavius; Type locality: Espagne: Province de Gérone, massif des Montes Gavarras).
— Coiffait, 1984f: 404 (Octavius; characters; Spain).
— Orousset, 1991: 168 (Octavius; Spain).
▴ Distribution: Spain.
gemellus Orousset, 1988: 130 (Octavius; Type locality: Madagascar Centre: forêt d’Ambo-hiboatavo, à l’Est du lac de Mantasoa).
goanus Puthz, 1991, see: indicus Puthz, 1980.
gomyi Orousset, 1988: 93 (Octavius; Type locality: Madagascar Est: massif de l’Andohahelo).
grandiceps Mulsant and Rey, 1853: 39 (Homalota; Type locality: environs de Lyon).
— Fauvel, 1873: 62 [= 1873a: 220] (Octavius).
— Mulsant and Rey, 1878a: 323 (Octavius; characters; France).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1984f: 407 (Octavius; characters; France).
— Orousset, 1987b: 14 (Octavius; characters; notes; France).
— Orousset, 1991: 167 (Octavius; characters; France).
▴ Distribution: France.
berardi Lavagne, 1917: 139 (Octavius; variety of pyrenaeus; Type locality: Avèze; [Note: See lectotype designation by Orousset, 1983c]).
— Scheerpeltz, 1933: 1203 (Octavius; variety of pyrenaeus).
— Coiffait, 1958a: 84 (Octavius; characters; France).
— Orousset, 1983c: 135 (Octavius; lectotype designation).
— Coiffait, 1984f: 390 (Octavius; characters; France).
— Orousset, 1987b: 17 (Octavius; compared to grandiceps; France).
— Orousset, 1991: 167 (Octavius; synonym of grandiceps).
guamensis Puthz, 1977h: 112 (Octavius; Type locality: Marianen: Guam, Amantes Point).
gutierrezi Orousset, 1988: 132 (Octavius; Type locality: Madagascar Centre: tampoketsa d’Ambohitantely, au Nord d’Ankazobe, 1550 m).
longulus Orousset, 1988: 132 (Octavius; subspecies of gutierrezi; Type locality: Madagascar Est: route de Moramanga à Ambatondrazaka, à 10 km à l’Est d’Ambohidray, forêt, d’Ankarongambe, 900 m).
hervei Ochs, 1949: 38 (Octavius; Type locality: Sospel, A.-M.).
— Coiffait, 1958a: 83 (Octavius; characters; France; Italy).
— Coiffait, 1984f: 391 (Octavius; characters; Alps).
— Orousset, 1987b: 18 (Octavius; France).
— Orousset, 1991: 168 (Octavius; Italy).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: France, Italy.
himalayicus Coiffait, 1982: 130 (Nepalopeplus; Type locality: Nepal central: Seti Khola, 2235 m).
— Puthz, 1985d: 704 (Octavius; characters).
— Puthz, 1987a: 453 (Octavius; characters; notes; Nepal).
▴ Distribution: Nepal.
horrendus Orousset, 1988: 97 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, base de la forêt du cirque de Manjarivolo, 1200 m).
hypogaeus Orousset, 1988: 87 (Octavius; Type locality: Madagascar Centre: Antanifotsy, au N. du massif de l’Andringitra, forêt d’Ambohipo, 1500 m).
indicus Puthz, 1980g: 750 (Octavius; Type locality: Indien: Kerala: Palghat Hills, 10 km au
Nord de Malampuzha Dam, 150 m).
goanus Puthz, 1991a: 254 (Octavius; subspecies of indicus; Type locality: Indien: Goa: Molem.).
indifferens Puthz, 1979d: 124 (Octavius; Type locality: Vietnam: Cuc phuong, Ninh binh).
insignis Orousset, 1988: 127 (Octavius; Type locality: Madagascar Centre: forêt d’Ambo-hiboatavo, à l’Est du lac de Mantasoa).
insularis Fauvel, 1873: 62 [= 1873a: 220] (Octavius; Type locality: Corse: Omessa; [Note: See lectotype designation by Orousset, 1983]. Originally cited from Corse, Rogliano et Omessa; Sardaigne, sassari et Bonnari).
— Fauvel, 1876a: 59 [= 1876: 244] (Octavius; characters).
— Saulcy, 1878: 123 (Octavius; characters; Corsica; Sardinia).
— Mulsant and Rey, 1878a: 318 (Octavius; characters; France).
— Sainte-Claire Deville, 1906: 79 (Octavius; Corsica).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1958a: 90 (Octavius; characters; Corsica).
— Orousset, 1983: 56 (Octavius; lectotype designation: Omessa; characters; Corsica).
— Coiffait, 1984f: 398 (Octavius; characters; Corsica).
— Orousset, 1986c: 104 (Octavius; Corsica).
— Orousset, 1990a: 126 (Octavius; characters).
▴ Distribution: Corsica; Sardinia.
interfector Orousset, 1988: 101 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, Anjavidilava, 2000 m).
italus Fauvel, 1907a: 105 (Octavius; Type locality: Italie méridionale: Oriolo; [Note: See lectotype designation by Orousset, 1991: 167]).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Coiffait, 1965f: 98 (Octavius; characters).
— Puthz, 1972i: 129 (Octavius; Italy).
— Coiffait, 1984f: 402 (Octavius; characters; Italy).
— Orousset, 1991: 167 (Octavius; lectotype designation).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Italy.
ituriensis Kistner, 1960a: 119 (Octavius; Type locality: Belgian Congo: Kibali-Ituri, between Mambasa and Mungbere, near Epulu, 900 m).
— Puthz, 1977h: 111 (Octavius; characters; Zaire).
— Puthz, 1979c: 55 (Octavius; characters).
— Puthz, 1986f: 198 (Octavius; characters; Zaire; Congo; Gabon).
▴ Distribution: Congo, Congo Republic, Gabon.
janae Janák, 1996: 187 (Octavius; Type locality: Madagascar Centre: Ambohimahasoa env., RN 7 [= Nationalstrasse No 7], pk 344 [= km 344], 1300–1400 m).
javanus Puthz, 1978d: 41 (Octavius; Type locality: Java: Udjong-Kulon, Pulau Pentjang, 2 m).
joffrei Orousset, 1981: 25 (Octavius; Type locality: Rivesaltes, Pyrénées-Orientales).
kaboboensis Kistner, 1963: 199 (Turellus; Type locality: Congo Republic: Mont Kabobo, Albertville territory, Hte. Kiymbi, 1700 m).
kenyamontis Puthz, 1991a: 253 (Octavius; Type locality: Kenya: Mount Kenya, Met. Station Q, 3050 m).
kenyanus Puthz, 1984h: 570 (Octavius; Type locality: Kenya: Emba, Irangi Forest Station, 2000 m).
khasimontium Puthz, 1985d: 700 (Octavius; Type locality: Indien: Maghalaya: Khasi Hills, entre Mawsynram et Balat, à 16 km de Mawsynram, 1000 m).
kistneri Puthz, 1984h: 567 (Octavius; Type locality: Zaire: Tshuapa, Mabali, terr. Bikoro, Lac Tumba).
leenhardti Orousset, 1983: 50 (Octavius; Type locality: col de Sorro à l’Est de Vico, 525 m).
— Coiffait, 1984f: 394 (Octavius; characters; Corsica).
— Orousset, 1986c: 103 (Octavius; characters; notes; Corsica).
— Orousset, 1990a: 126 (Octavius; characters).
▴ Distribution: Corsica.
leleupi Kistner, 1961: 21 (Doletica; Type locality: Congo: Kivu, Nyakasiba, Kabare, 1800 m).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1977h: 111 (Octavius; characters; Zaire).
— Puthz, 1979c: 57 (Octavius; characters).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 197 (Octavius; characters).
▴ Distribution: Congo.
leleupi Kistner, 1963, see: leleupianus Puthz, 1980.
leleupianus Puthz, 1980j: 27 (Octavius; replacement name for leleupi Kistner, 1963).
leleupi Kistner, 1963: 202 (Octavius; [preoccupied]; Type locality: Congo Republic: Kivu, Mwenga, 1250 m).
lepidus Orousset, 1986c: 106 (Octavius; Type locality: Zonza, alt. 750 m, Corse-du-sud).
lichtensteini Lavagne, 1917: 139 (Octavius; subspecies of pyrenaeus; Type locality: La Gardiole; [Note: See lectotype designation by Orousset, 1983c]. Originally cited from Gardiole à Mireval, La Paillade près Montpellier, Le Bousquet-d’Orb; La Salvetat).
— Scheerpeltz, 1933: 1203 (Octavius; subspecies of pyrenaeus).
— Coiffait, 1958a: 87 (Octavius; cited as valid species; characters; France).
— Orousset, 1983c: 136 (Octavius; lectotype designation: La Gardiole).
— Coiffait, 1984f: 395 (Octavius; characters; France).
— Orousset, 1987b: 18 (Octavius; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France.
meridionalis Coiffait, 1958a: 88 (Octavius; subspecies of lichtensteini; Type locality: Ariège: Lacave).
— Coiffait, 1984f: 395 (Octavius; subspecies of lichtensteini; characters; Pyrenees).
— Orousset, 1987b: 18 (Octavius; subspecies of lichtensteini; France).
perroti Coiffait, 1963c: 182 (Octavius; subspecies of lichtensteini; Type locality: Hérault, Saint-Pons de Mauchiens).
rabili Orousset, 1981: 19 (Octavius; Type locality: Forêt de la Grésigne, alt. 250 m, Tarn).
llinaresi Orousset, 1988: 108 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt de Manjakatompo, 1650 m).
loebli Puthz, 1979c: 58 (Octavius; Type locality: Elfenbeinküste: Agboville, Forêt de Yapo, près Yapo-Gare).
loeblianus Puthz, 1979c: 58 (Octavius; Type locality: Elfenbeinküste: Abidjan, Parc national du Banco).
loebliellus Puthz, 1984h: 573 (Octavius; Type locality: Côte d’Ivoire: Man, 2 km à l’est de Sangoviné).
longipes Coiffait, 1963c: 180 (Octavius; Type locality: Ariège, Sentein, Vallon de l’Izard, vers 800 m).
longispinosus Orousset, 1988: 138 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, station de Manjakatompo).
longulus Orousset, 1988, see: gutierrezi Orousset, 1988.
lueboensis Kistner, 1962: 102 (Turellus; Type locality: Congo Republic: Kasai, route Luluabourg-Tshikapa, along the Luebo stream).
machadoi Cameron, 1950f: 112 (Doletica; Type locality: Angola: Dundo).
— Kistner, 1961: 21 (Doletica; characters; Angola).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 196 (Octavius; characters).
▴ Distribution: Angola.
madagascariensis Orousset, 1988: 90 (Octavius; Type locality: Madagascar Centre: Antanifotsy, forêt d’Ambohipo, 1500 m).
madecassus Orousset, 1988: 126 (Octavius; Type locality: Madagascar Est: route de Lakato, 1100 m).
major Coiffait, 1958a: 84 (Octavius; Type locality: Ariège: Aulus).
princeps Coiffait, 1963c: 76 (Octavius; subspecies of major; Type locality: Ariège, Rimont, Forêt de Bellissens vers 1000 m sur le versant Nord).
malgassicus Orousset, 1988: 97 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, Anjavidilava, 1800 m).
manjarivolo Orousset, 1988: 100 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, cirque de Manjarivolo).
marositry Orousset, 1988: 104 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, Marositry Est, rivière Antsifotra).
massatensis Coiffait, 1959a: 38 (Octavius; Type locality: France: Col des Caougnous, commune de Bousenac près de Massat, Ariège).
— Coiffait, 1984f: 390 (Octavius; characters; Pyrenees; France).
— Orousset, 1987b: 19 (Octavius; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France.
mendax Puthz, 1989b: 75 (Octavius; Type locality: Côte d’Ivoire: Monogaga).
meridionalis Coiffait, 1958, see: lichtensteini Lavage, 1917.
microps Kistner, 1967: 55 (Turellus; Type locality: Republic of South Africa, Transvaal, Pietersburg District, Woodbush Forest).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 199, 200 (Octavius; characters).
▴ Distribution: South Africa.
mindorensis Orousset, 1980a: 182 (Octavius; Type locality: Puerto Galera, Tabinay, alt. 300 m, île de Mindoro, Philippines).
minor Puthz, 1985, see: flavescens Kistner, 1961.
minutus Kistner, 1961: 23 (Doletica; Type locality: Congo: Kibali-Ituri, Mambasa-Mungbere, near Epulu, 900 m).
— Kistner, 1962: 102 (Turellus).
— Kistner, 1963: 201 (Turellus; Congo Republic).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 194 (Octavius; characters; Angola; Zaire).
▴ Distribution: Angola, Congo.
monticola Orousset, 1988: 114 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, 2000 m).
morio Kistner, 1967: 58 (Turellus; Type locality: Republic of South Africa, Transvaal, Pilgrim's Rest District, Graskop Forest).
— Puthz, 1976e: 240 (Turellus; synonym of brincki).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 199 (Octavius; characters).
▴ Distribution: South Africa.
mostarensis Bernhauer, 1912a: 398 (Octavius; Type locality: Mostars).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1965f: 95 (Octavius; characters).
— Puthz, 1972i: 129 (Octavius; cited as mostariensis; Herczgowina).
— Orousset, 1981: 19 (Octavius; cited as mostariensis; Herzegowina).
— Coiffait, 1984f: 391 (Octavius; characters; Herzegowina).
▴ Distribution: Bosnia Hercegovina.
mutatus Coiffait, 1963c: 180 (Octavius; Type locality: Ariège, Bousenac, Col des Caougnous).
— Coiffait, 1984f: 393 (Octavius; characters; Pyrenees).
— Orousset, 1987b: 19 (Octavius; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France.
natalensis Scheerpeltz, 1974, see: natalensis Kistner, 1967.
natalensis Kistner, 1967: 57 (Turellus; Type locality: Republic of South Africa, Natal, Pietermaritzburg, Townbush Forest).
— Puthz, 1976e: 240 (Turellus; characters).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 200 (Octavius; characters).
▴ Distribution: South Africa.
natalensis Scheerpeltz, 1974: 95 (Traegardhius; [preoccupied]; Type locality: Natal: Sweetwaters).
— Puthz, 1976e: 240 (Turellus; synonym of natalensis).
— Puthz, 1980j: 27 (Octavius; synonym of natalensis).
neotropicus Puthz, 1977h: 107 (Octavius; Type locality: Costa Rica: Asahar de Sabanilla).
novaebritanniae Puthz, 1977h: 120 (Octavius; Type locality: New Britain: Rabaul Keravat).
novaeguineae Puthz, 1977h: 117 (Octavius; Type locality: New Guinea NE: Wau, Kilolo Creek, 1000 m).
quadridenticulatus Puthz, 1977h: 118 (Octavius; subspecies of novaeguineae; Type locality: New Guinea NE: Lae).
quinquedenticulatus Puthz, 1977h: 118 (Octavius; subspecies of novaeguineae; Type locality: New Guinea NE: Wau, Kilolo Creek, 1000 m).
sexdenticulatus Puthz, 1977h: 118 (Octavius; subspecies of novaeguineae; Type locality: New Guinea NE: Bayer River, Sanctuary).
oculocallus Coiffait, 1959a: 40 (Octavius; Type locality: France: Laguiolle, Aveyron).
— Coiffait, 1984f: 395 (Octavius; characters; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France.
orbensis Orousset, 1986c: 104 (Octavius; Type locality: Isolaccio-di-Fiumorbo, alt. 550 m, Haute-Corse).
orcinus Orousset, 1988: 95 (Octavius; Type locality: Madagascar Centre: massif de l’Andringitra, base de la forêt du cirque de Manjarivolo, 1200 m).
pandora Puthz, 1984h: 572 (Octavius; Type locality: Zaire: Kivu, terr. Uvira, 800 m. Also cited from Chad).
paraguayanus Puthz, 1986d: 617 (Octavius; Type locality: Paraguay: Prov. Cordillera, env. 6 km S. Tobati, Reposoir).
parvulus Orousset, 1988: 93 (Octavius; Type locality: Madagascar Est: massif de l’Ando-hahelo).
pauliani Jarrige, 1968: 875 (Octavius; Type locality: Madagascar-Centre, Manjakatompo, Ankaratra).
pectinifer Puthz, 1986f: 191 (Octavius; Type locality: Südafrika: Cape Province: Garden of Eden, 34°02′S, 23°12′E).
perroti Coiffait, 1963, see: lichtensteini Lavagne, 1917.
peyrierasi Orousset, 1988: 112 (Octavius; Type locality: Madagascar Est: environs de Périnet, station du lac Vert).
piriensis Kistner, 1967: 60 (Turellus; Type locality: Republic of South Africa, Kingwilliamstown District, Pirie Forest).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 199 (Octavius; characters).
▴ Distribution: South Africa.
pluto Orousset, 1988: 136 (Octavius; Type locality: Madagascar Centre: pays Betsileo, Ambatofitoharana, RN 7, pk 292, 1600 m).
portensis Coiffait, 1959a: 42 (Octavius; Type locality: Corse, environs de Porto).
— Orousset, 1983: 58 (Octavius; characters; Corsica).
— Coiffait, 1984f: 405 (Octavius; characters; Corsica).
— Orousset, 1986c: 106 (Octavius; Corsica).
— Orousset, 1990a: 126 (Octavius; characters).
▴ Distribution: Corsica.
princeps Coiffait, 1963, see: major Coiffait, 1958.
priotoni Coiffait, 1966e: 164 (Octavius; [Note: Cited as piotoni at the head of the description but as priotoni on pages 163 and in the caption for the figures on page 165; the species was named for the collector Prioton]; Type locality: Mons, gorge d’Héric, Hérault).
pseudopauliani Orousset, 1988: 136 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, station de Manjakatompo).
puthzi Pace, 1983: 104 (Octavius; Type locality: Umgb. Görz).
— Coiffait, 1984f: 397 (Octavius; characters; Italy).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Italy.
puthzianus Orousset, 1988: 140 (Octavius; Type locality: Madagascar Centre: Sud-Est d’Ambatolampy, Tsinjoarivo).
pyrenaeus Fauvel, 1873: 62 [= 1873a: 220] (Octavius; Type locality: Hautes Pyrénées, L’Héris près Bagnères de Bigorre; [Note: See lectotype designation by Orousset, 1986d]).
— Mulsant and Rey, 1878a: 322 (Octavius; characters; France).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Portevin, 1929: 390 (Octavius; characters; France).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1958a: 86 (Octavius; characters; Pyrenees).
— Coiffait, 1984f: 389 (Octavius; characters; Pyrenees).
— Orousset, 1986d: 99 (Octavius; lectotype designation; characters; notes; France).
— Orousset, 1991: 168 (Octavius; France).
▴ Distribution: France.
quadridenticulatus Puthz, 1977, see: novaeguineae Puthz, 1977.
quinquedenticulatus puthz, 1977, see: novaeguineae Puthz, 1977.
rabili Orousset, 1981, see: lichtensteini Lavagne, 1917.
raja Puthz, 1986e: 369 (Octavius; Type locality: Indien: Kerala: Nelliampathi Hills, au nord-ouest des Anaimalai Hills, Kaikatty, 900 m).
rapax Orousset, 1988: 116 (Octavius; Type locality: Madagascar Est: route de Moramanga à Ambatondrazaka, à 10 km à l’Est d’Ambohidray, forèt d’Ankarongambe, 1050 m).
raymondi Saulcy, 1878: 124 (Octavius; Type locality: Bonnari, en Sardaigne; [Note: See lectotype designation by Orousset, 1983c]).
— Bernhauer and Schubert, 1911: 187 (Octavius; cited as raimondi; catalog).
— Scheerpeltz, 1933: 1203 (Octavius; catalog; Tunisia).
— Coiffait, 1958a: 92 (Octavius; characters; Sardinia; Corsica; Tunisia; Algeria).
— Orousset, 1983: 62 (Octavius; notes; Sardinia).
— Orousset, 1983c: 136 (Octavius; lectotype designation: Bonnari).
— Coiffait, 1984f: 401 (Octavius; characters; Sardinia; Corsica record may be erroneous).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Corsica(?), Sardinia, Tunisia, Algeria.
reali Orousset, 1989a: 219 (Octavius; Type locality: Haute-Corse: Zilia, au Nord-Est de Calenzana, alt. 325 m).
rostrifer Puthz, 1986f: 192 (Octavius; Type locality: Cape Province: Nature's Valley, 33°58′S, 23°33′E).
rugosus Orousset, 1988: 89 (Octavius; Type locality: Madagascar Centre: Antanifotsy, forêt de Viloboa).
sardous Coiffait, 1965f: 96 (Octavius; Type locality: Espagne: Province de Barcelone, Tordera).
— Orousset, 1983: 62 (Octavius; notes; Sardinia).
— Coiffait, 1984f: 402 (Octavius; characters; Sardinia).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Sardinia, Spain.
sculptilis Orousset, 1988: 127 (Octavius; Type locality: Madagascar Est: région de Périnet, forêt de la Grande Ile, 900 m).
secqi Orousset, 1991: 168 (Octavius; Type locality: Estours près de Seix [sud de Saint-Girons], Ariège).
semipiceus Jarrige, 1968: 876 (Octavius; Type locality: Madagascar-Centre, Manjakatompo, Ankaratra).
senyensis Coiffait, 1958a: 92 (Octavius; Type locality: Prov. de Barcelone: Figaro, au pied de la Sierra de Montseny).
serraticollis Puthz, 1984h: 565 (Octavius; Type locality: Côte d’Ivoire: Sassandra, à 10 km de Monogaga).
serratus Orousset, 1988: 91 (Octavius; Type locality: Madagascar Est: massif de l’Andohahelo).
sexdenticulatus Puthz, 1977, see: novaeguineae Puthz, 1977.
solomonensis Puthz, 1977h: 120 (Octavius; Type locality: Solomon Islands: Rendova Island).
spiniphallus Orousset, 1981: 22 (Octavius; Type locality: fontaine de Sendo, Corse).
— Orousset, 1983: 60 (Octavius; characters; Corsica).
— Coiffait, 1984f: 402 (Octavius; characters; Corsica).
— Orousset, 1990a: 126 (Octavius; characters).
▴ Distribution: Corsica.
apicalis Orousset, 1983: 62 (Octavius; subspecies of spiniphallus; Type locality: Rezza, au Sud de Guagno, alt. 250 m).
— Coiffait, 1984f: 402 (Octavius; subspecies of spiniphallus; characters; Corsica).
— Orousset, 1990a: 126 (Octavius; subspecies of spiniphallus; characters).
spinulosus Coiffait, 1963c: 175 (Octavius; Type locality: Ariège, Bordes-sur-Lez, Vallée du Ribérot vers 1000 m).
strigosus Orousset, 1990a: 124 (Octavius; Type locality: Corse du Sud: environs s’Arca, au sud-ouest de Porto-Vecchio, alt. 20 m).
subtilis Puthz, 1979c: 54 (Octavius; Type locality: Elfenbeinküste: Abidjan, Parc national du Banco).
sulcicollis Bernhauer, 1935f: 214 (Thaxterius; Type locality: Kamerun).
— Kistner, 1960a: 121 (Octavius; characters; Cameroon).
— Puthz, 1977h: 111 (Octavius; characters; Congo).
— Puthz, 1979c: 55 (Octavius; characters; Ivory Coast).
— Puthz, 1986f: 198 (Octavius; characters; Cameroon; Ivory Coast).
— Frank, 1982: 16 (Octavius; list of parasites).
▴ Distribution: Cameroon, Ivory Coast.
sulcifrons Jarrige, 1968: 874 (Octavius; Type locality: Madagascar-Est, Périnet).
sumatranus Puthz, 1989e: 115 (Octavius; Type locality: Sumatra: Aceh-Selatan, Babahrot, 100 m).
tarragonensis Coiffait, 1963c: 185 (Octavius; Type locality: Espagne, Province de Tarragone, Rodôna, à l’ouest de la vallée du Rio Francoli).
franzi Coiffait, 1963c: 185 (Octavius; subspecies of tarragonensis; Type locality: Espagne, Province de Barcelone, Costas de Garraf).
tavakiliani Orousset, 1983b: 114 (Octavius; Type locality: Mont Grand-Matoury, 7 km S. de Cayenne, alt. 200 m, Guyane française).
tenebricosus Orousset, 1988: 139 (Octavius; Type locality: Madagascar Centre: massif de l’Ankaratra, forêt de Betay, 2100 m).
tergestinus Coiffait, 1963c: 184 (Octavius; Type locality: Slovénie: Kozina).
— Pace, 1983: 105, 108 (Octavius; characters; Slovenia).
— Coiffait, 1984f: 397 (Octavius; characters; Slovenia).
▴ Distribution: Slovenia.
terrensis Kistner, 1960a: 119 (Octavius; Type locality: Parc National de l’Upemba: Kabwe-sur-Muye, affl. Mitembo, 1320 m).
— Puthz, 1977h: 111 (Octavius; characters; Zaire).
— Puthz, 1979c: 54 (Octavius; characters).
— Puthz, 1986f: 199 (Octavius; characters).
▴ Distribution: Congo.
terrigena Orousset, 1988: 107 (Octavius; Type locality: Madagascar Centre: pays Betsileo, Ambatofitoharana, RN 7, pk 292, 1600 m).
tibialis Kistner, 1967: 59 (Turellus; Type locality: Republic of South Africa, Cape Province, Swellendam District, Grootvatersbos).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 200 (Octavius; characters).
▴ Distribution: South Africa.
togoensis Puthz, 1977h: 108 (Octavius; Type locality: Togo: Palimé, Forêt de Kpimé).
torderensis Coiffait, 1965f: 96 (Octavius; Type locality: Espagne: Province de Barcelone, Tordera).
tortilis Orousset, 1988: 115 (Octavius; Type locality: Madagascar Centre: station forestière d’Angavokely, Carion, 1700 m).
torulosus Orousset, 1988: 122 (Octavius; Type locality: Madagascar Centre, [pays Betsileo], Ambatofitorahana, RN 7, pk 292, 1600 m).
transadriaticus Breit, 1911: 170 (Octavius; Type locality: Monte Maggiore [= Mont Ucka] en Istria; [Note: See lectotype designation by Orousset, 1991: 167]).
— Scheerpeltz, 1933: 1203 (Octavius; catalog).
— Coiffait, 1958a: 89 (Octavius; characters; Istrie: Monte Maggiore).
— Puthz, 1972i: 129 (Octavius; Istria).
— Orousset, 1981: 22 (Octavius; characters; Istria).
— Pace, 1983: 106, 107 (Octavius; characters; Italy).
— Coiffait, 1984f: 397 (Octavius; characters; Istria).
— Orousset, 1991: 167 (Octavius; lectotype designation: Monte Maggiore [= Mont Ucka] en Istria; characters; Yugoslavia).
▴ Distribution: Croatia.
vanjamanitra Orousset, 1988: 141 (Octavius; Type locality: Madagascar Centre, forêt de Vanjamanitra, à 8 km au Sud-Est d’Anjozorobe, 1380 m).
velebiticus Breit, 1913: 236 (Octavius; race of transadriaticus; Type locality: Velebitgebirge bei Starigrad).
— Pace, 1983: 107 (Octavius; Yugoslavia).
— Coiffait, 1984f: 407 (Octavius; valid species; characters; Yugoslavia).
▴ Distribution: Croatia.
venator Orousset, 1988: 137 (Octavius; Type locality: Madagascar Centre: forêt de Tsarafidy, aux environs d’Ambohimahasoa, 1450 m).
verschureni Kistner, 1961: 25 (Doletica; Type locality: Congo: Parc National de la Garamba).
— Kistner, 1962: 102 (Turellus).
— Puthz, 1974s: 769 (Turellus; characters; Chad).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 196, 197 (Octavius; characters).
▴ Distribution: Congo.
viduus Puthz, 1986f: 188 (Octavius; Type locality: Südafrika: Natal: Cathedral Peak, 28°57′S, 29°12′E).
vietnamensis Puthz, 1979d: 123 (Octavius; Type locality: Vietnam: Luc yen).
vitalei Bernhauer, 1908a: 25 (Octavius; Type locality: Sizilien, Umgebung von Messina).
— Bernhauer and Schubert, 1911: 187 (Octavius; catalog).
— Coiffait, 1958a: 94 (Octavius; characters; Sicily).
— Coiffait, 1963c: 188 (Octavius; Algeria; Tunisia).
— Coiffait, 1965f: 97 (Octavius; characters).
— Puthz, 1972i: 129 (Octavius; Sicily; Italy).
— Bordoni, 1973j: 676 (Octavius; notes; discussion of distribution of subspecies).
— Coiffait, 1984f: 405 (Octavius; characters; Sicily; Italy; Tunisia; Algeria).
— Orousset, 1991: 168 (Octavius; notes; Algeria; Italy).
— Sparacio, 1995: 149 (Octavius; characters; notes; Sicily).
— Zanetti, 1995: 19 (Octavius; Italy).
▴ Distribution: Sicily, Italy, Algeria, Tunisia.
cossyrensis Coiffait, 1965f: 98 (Octavius; subspecies of vitalei; Type locality: Ile de Pantellaria).
— Coiffait, 1984f: 405 (Octavius; subspecies of vitalei; characters; Pantellaria Island).
— Zanetti, 1995: 19 (Octavius; subspecies of vitalei; Italy).
abruzzensis Coiffait, 1965f: 98 (Octavius; subspecies of vitalei; Type locality: Abruzzes, environs de Chieti).
— Coiffait, 1984f: 405 (Octavius; subspecies of vitalei; characters; Italy).
— Zanetti, 1995: 19 (Octavius; subspecies of vitalei; Italy).
viti Puthz, 1977h: 110 (Octavius; Type locality: Togo: Palimé, Forêt de Klouto).
vivariensis Coiffait, 1959a: 43 (Octavius; Type locality: Corse: Vivario, à 10 km au Nord de Vizzavona).
— Orousset, 1983: 52 (Octavius; characters; Corsica).
— Coiffait, 1984f: 401 (Octavius; characters; Corsica).
— Orousset, 1990a: 127 (Octavius; characters).
▴ Distribution: Corsica.
wittei Kistner, 1960a: 116 (Doletica; Type locality: Parc National de l’Upemba: gorges de la Pelenge, 1150 m).
— Kistner, 1961: 26 (Doletica; characters).
— Kistner, 1962: 102 (Turellus).
— Kistner, 1963: 201 (Turellus; Congo Republic).
— Puthz, 1980j: 27 (Octavius).
— Puthz, 1986f: 197 (Octavius; characters).
▴ Distribution: Congo.
zariquieyi Coiffait, 1958a: 94 (Octavius; Type locality: Catalogne: Arenys de Mar).
Phaenoctavius
[1 species; Neotropical Region]
Phaenoctavius Pace, 1986: 45 (species included: brasiliensis). Type species: Phaenoctavius brasiliensis Pace, fixed by original designation and monotypy.
brasiliensis Pace, 1986: 46 (Phaenoctavius; Type locality: Brasile: Nova Teutonia).
Protopristus
[3 species; Australian Region]
Protopristus Broun, 1909: 225 (species included: minutus) Type species: Protopristus minutus Broun, fixed by monotypy.
— Blackwelder, 1952: 323 (type species: minutus).
— Puthz, 1980j: 21 (notes; characters).
— Newton, 1985: 205 (notes on austral distribution).
Geosthethus Oke, 1933: 111 (species included: attenuatus). Type species: Geosthethus attenuatus Oke, by original designation and monotypy.
attenuatus Oke, 1933: 111 (Geosthethus; Type locality: Australia: Victoria: Belgrave, Warburton).
— Puthz, 1978c: 121 (Geosthethus; characters; Australia).
— Puthz, 1980j: 22 (Protopristus; notes).
▴ Distribution: Australia.
gracilis Puthz, 1979a: 75 (Geosthethus; Type locality: Australia: N.S.W.: Jenolan State Forest entre Jenolan et Hampton).
minutus Broun, 1909: 226 (Protopristus; Type locality: Broken River, Canterbury; [Note: See lectotype designation by Puthz, 1980j]).
Schatzmayrina
[2 species; Ethiopian Region]
Schatzmayrina C. Koch, 1934: 63 (species included: oxyclypea). Type species: Schatzmayrina oxyclypea C. Koch, fixed by monotypy.
— Blackwelder, 1952: 345 (type species: oxyclypea).
— Kistner, 1961: 27 (characters; key to species; type species: oxyclypea).
— Scheerpeltz, 1974: 107 (characters).
— Coiffait, 1984f: 354 (characters; species of western Palaearctic region).
angolensis Kistner, 1962, see: oxyclypea C. Koch, 1934.
boops Puthz, 1989b: 76 (Schatzmayrina; Type locality: Côte d’Ivoire: Abidjan: Parc National du Banco).
ceylonica Puthz, 1978, see: oxyclypea C. Koch, 1934.
congoensis Kistner, 1961, see: oxyclypea C. Koch, 1934.
indica Puthz, 1978, see: oxyclypea C. Koch, 1934.
nigritula Kistner, 1961, see: oxyclypea C. Koch, 1934.
oxyclypea C. Koch, 1934: 69 (Schatzmayrina; Type locality: Pyramiden von Ghizeh; Sakkarah).
— Kistner, 1961: 30 (Schatzmayrina; characters; Egypt).
— Puthz, 1978f: 47 (Schatzmayrina; characters; Egypt; Ethiopia; Tanzania; Kenya; Burma).
— Coiffait, 1984f: 355 (Schatzmayrina; characters; Egypt; Morocco; intertropical Africa [?]).
▴ Distribution: Angola, Myanmar, Congo, Egypt, Ethiopia, India, Kenya, Morocco, Sri Lanka, Tanzania.
nigritula Kistner, 1961: 31 (Schatzmayrina; Type locality: Congo: Parc National de la Garamba, Mabanga).
— Puthz, 1978f: 47 (Schatzmayrina; synonym of oxyclypea).
— Coiffait, 1984f: 355 (Schatzmayrina; synonym of oxyclypea).
congoensis Kistner, 1961: 33 (Schatzmayrina; Type locality: Congo: Kivu, Uvira, 800 m).
— Puthz, 1978f: 47 (Schatzmayrina; synonym of oxyclypea).
— Coiffait, 1984f: 355 (Schatzmayrina; synonym of oxyclypea).
angolensis Kistner, 1962: 103 (Schatzmayrina; Type locality: Angola: Alto Cuilo, Tcha-Muchito).
— Puthz, 1978f: 47 (Schatzmayrina; synonym of oxyclypea).
— Coiffait, 1984f: 355 (Schatzmayrina; synonym of oxyclypea).
ceylonica Puthz, 1978f: 50 (Schatzmayrina; subspecies of oxyclypea; Type locality: Ceylon: Nuwara Eliya, 1900 m).
indica Puthz, 1978f: 50 (Schatzmayrina; subspecies of oxyclypea; Type locality: India: Madras, Nilgiri, 15 km E. Coonoor, 900 m).
Tamotus
[5 species; Neotropical Region]
Tamotus Schaufuss, 1872: 248 (without species). Type species: Tamotus femoratus Schaufuss, fixed by Schaufuss, 1874: 289 by subsequent monotypy.
— Schaufuss, 1874: 289 (first included species: femoratus).
— Schaufuss, 1877: 24 (characters).
— Schaufuss, 1877a: 460 (characters).
— Bernhauer and Schubert, 1911: 188 (world catalog; 1 species).
— Blackwelder, 1943: 227 (type species: femoratus).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 372 (type species).
— Puthz, 1973k: 60 (characters; key to species).
— Scheerpeltz, 1974: 107 (characters).
Paredaphus Bernhauer, 1915l: 291 (species included: tropicus). Type species: Paredaphus tropicus Bernhauer, fixed by monotypy.
— Scheerpeltz, 1933: 1204 (world catalog supplement).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 294 (type species: tropicus).
— Puthz, 1973k: 60 (synonym of Tamotus).
— Scheerpeltz, 1974: 107 (characters).
brasilianus Puthz, 1973k: 66 (Tamotus; Type locality: Brazil [SE]: Blumenau).
carinifrons Puthz, 1986d: 618 (Tamotus; Type locality: Paraguay: Provinz Paraguari, Estancia Montiel Potrero, 15 km est de Cerrito, 40 km sud-ouest de Caapucu).
crassus Puthz, 1973k: 65 (Tamotus; Type locality: Costa Rica: Santa Clara Province: Hamburgfarm. Also cited from Panama).
femoratus Schaufuss, 1874: 289 (Tamotus; Type locality: Cuba).
— Bernhauer and Schubert, 1911: 188 (Tamotus; catalog).
— Blackwelder, 1943: 228 (Tamotus; Cuba).
— Puthz, 1973k: 64 (Tamotus; characters; Cuba).
▴ Distribution: Cuba. tropicus Bernhauer, 1915l: 292 (Paredaphus; Type locality: Brasilien: S. Paulo).
— Scheerpeltz, 1933: 1204 (Paredaphus; catalog).
— Puthz, 1973k: 64 (Tamotus; characters; Brazil).
— Orousset, 1986b: 20 (Tamotus; Brazil).
▴ Distribution: Brazil.
Tribe Fenderiini
Fenderiini Scheerpeltz, 1974: 103 (genera included: Fenderia). Type genus: Fenderia (see Newton and Thayer, 1992: 56).
Fenderia
[1 species; Nearctic Region]
Fenderia Hatch, 1957: 245 (species included: capizzii). Type species: Fenderia capizzii Hatch, fixed by original designation and monotypy.
— Steel, 1959: 82 (transferred from Osoriinae to Euaesthetinae).
— Moore and Legner, 1974c: 118 (characters; notes).
— Scheerpeltz, 1974: 103 (characters).
— Puthz, 1974t: 924 (genus of Euaesthetinae).
— Moore and Legner, 1975: 172 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 245 (characters; notes).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (1 Nearctic species; notes; characters in key).
capizzii Hatch, 1957: 246 (Fenderia; Type locality: Oregon: Curry Co.: Humbug Mt. Also cited from Washington and California).
Stictocranius
[3 species; Palaearctic and Nearctic Regions]
Stictocranius LeConte, 1866: 374 (species included: puncticeps). Type species: Stictocranius puncticeps LeConte, fixed by monotypy.
— Gemminger and Harold, 1868: 632 (catalog).
— Bernhauer and Schubert, 1911: 187 (world catalog; 1 species).
— Leng, 1920: 100 (catalog of North American species).
— Blackwelder, 1952: 361 (type species: puncticeps).
— Scheerpeltz, 1974: 103 (characters).
— Moore and Legner, 1975: 264 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 247 (characters; notes).
— Watt and McColl, 1982: 14, 20 (notes; [Note: Misidentified according to Newton, 1996: 159]).
— Downie and Arnett, 1996: 461 (characters; species of Northeastern North America).
— Newton, 1996: 159 (notes).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (2 Nearctic species; notes; characters in key).
chinensis Puthz, 1974p: 36 (Stictocranius; Type locality: China: West Szechuan, Wassuland, Bezirk Sankiang-kou, Strecke S.-S., 2000 m).
mariae Hatch, 1957: 245 (Paralispinus; Type locality: Oregon: Benton Co.: Marys Peak).
— Newton, 1989: 145 (Stictocranius).
— Puthz, 1989d: 165 (Stictocranius; characters of male; California).
▴ Distribution: USA.
puncticeps LeConte, 1866: 374 (Stictocranius; Type locality: Washington, D.C.).
— Bernhauer and Schubert, 1911: 187 (Stictocranius; catalog).
— Puthz, 1974p: 37 (Stictocranius; characters; West Virginia; North Carolina; District of Columbia; Kentucky; Pennsylvania; Illinois).
— Puthz, 1974t: 928 (Stictocranius; characters; New Jersey; New York; Pennsylvania; Illinois; North Carolina; District of Columbia; Virginia; West Virginia; Kentucky; Alabama).
— Newton, 1989: 145 (Stictocranius; Arkansas).
— Puthz, 1989d: 165 (Stictocranius; Indiana; Ohio; New Hampshire; Maryland; South Carolina).
— Campbell and Davies, 1991: 112 (Stictocranius; Canada).
— Downie and Arnett, 1996: 461 (Stictocranius; characters; USA).
▴ Distribution: USA, Canada.
Tribe Nordenskioldiini
Nordenskioldiini Bernhauer and Schubert, 1911: 186 (genera included: Nordenskioldia). Type genus: Nordenskioldia (see Newton and Thayer, 1992: 56).
— C. Koch, 1934: 67 (characters in key).
— Scheerpeltz, 1940: 31 (key to Palaearctic genera).
— Cameron, 1944b: 69 (characters).
— Scheerpeltz, 1974: 102 (characters).
— Coiffait, 1984f: 353 (characters).
— Newton and Thayer, 1992: 56 (tribe of Euaesthetinae).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (characters; notes).
Edaphosoma
[6 species; Palaearctic and Oriental Regions]
Edaphosoma Scheerpeltz, 1976: 25 (species included: janetscheki). Type species: Edaphosoma janetscheki Scheerpeltz, fixed by original designation.
Orosthetus Puthz, 1979e: 9 (species included: coriaceus, nepalensis, sikkimensis). Type species: Orosthetus coriaceus Puthz, fixed by original designation.
cassagnaui Coiffait, 1983d: 175 (Edaphosoma; Type locality: Nepal: Trisuli Gosaikund, 3660 m).
coriaceum Puthz, 1979e: 10 (Orosthetus; Type locality: Nepal: Topke Gola, 27°38′N, 87°35′E, ca. 12,500 feet).
janetscheki Scheerpeltz, 1976: 27 (Edaphosoma; Type locality: Nepal: Mingbo-Tal beim Airstrip der Hillary-Makalu-Expedition, ca. 4800 m).
— Puthz, 1980j: 16 (Edaphosoma; characters).
— Puthz, 1986: 138 (Edaphosoma; characters).
▴ Distribution: Nepal.
nepalense Puthz, 1979e: 11 (Orosthetus; Type locality: West-Nepal: Gebiet von Jumla, Mahidoela-Pass, 5000 m).
— Puthz, 1980j: 16 (Edaphosoma).
— Puthz, 1980j: 17 (Edaphosoma).
— Puthz, 1986: 138 (Edaphosoma; characters).
▴ Distribution: Nepal.
scherpa Puthz, 1986: 135 (Edaphosoma; Type locality: Nepal: Provinz Bagmati: Yangri Ridge, 4350 m).
sikkimense Puthz, 1979e: 11 (Orosthetus; Type locality: Indien: Sikkim, Dzongri, 13,000 feet).
Nordenskioldia
[2 species; Nearctic and Palaearctic Regions]
Nordenskioldia J. Sahlberg, 1880: 96 (cited as Nordenskiöldia; species included: glacialis).
Type species: Nordenskioldia glacialis J. Sahlberg, fixed by monotypy.
— Heyden, 1880: 79 (cited as Nordenskiöldia; list of species of Siberian region).
— Duvivier, 1883: 178 (cited as Nordenskioeldia; catalog).
— Bernhauer and Schubert, 1911: 186 (world catalog; 1 species).
— Winkler, 1925: 357 (cited as Nordenskioeldia; catalog for Palaearctic region).
— Blackwelder, 1952: 262 (cited as Nordenskiöldia; type species: glacialis).
— Tikhomirova, 1973a: 171 (cited as Nordenskioeldia; checklist of species of USSR).
— Scheerpeltz, 1974: 102 (cited as Nordenskioeldia; characters).
— Puthz, 1974t: 926 (cited as Nordenskioldia; notes; key to species).
— Moore and Legner, 1979: 246 (cited as Nordensioldia and Nordenskioldia; characters; notes).
— Newton and Thayer, 1992: 56 (cited as Nordenskioldia; name emended).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (1 Nearctic species; notes; characters in key).
columbiana Puthz, 1974t: 926 (Nordenskioldia; Type locality: British Columbia: Yoho National Park, 5500′, Otterhead River).
glacialis J. Sahlberg, 1880: 96 (Nordenskioldia, cited as Nordenskiöldia; Type locality: vicum Chantaika).
Tribe Stenaesthetini
Stenaesthetini Bernhauer and Schubert, 1911: 186 (genera included: Stenaesthetus, Stictocranius). Type genus: Stenaesthetus (see Newton and Thayer, 1992: 56).
— Scheerpeltz, 1933: 1202 (world catalog supplement).
— C. Koch, 1934: 67 (characters in key).
— Scheerpeltz, 1940: 31 (key to Palaearctic genera).
— Blackwelder, 1944: 115 (checklist of species from Mexico, Central and South America, and the West Indies).
— Cameron, 1944b: 69 (characters).
— Arnett, 1963: 266 (characters; list of North American genera).
— Scheerpeltz, 1974: 103 (characters).
— Coiffait, 1984f: 353 (characters).
— Newton and Thayer, 1992: 56 (tribe of Euaesthetinae).
Agnosthaetus
[5 species; Australian Region]
Agnosthaetus Bernhauer, 1939f: 213 (species included: whitehorni, cariniceps, vicinus, stilbus, bisulciceps, brouni). Type species: Agnosthaetus brouni Bernhauer, by original designation.
— Blackwelder, 1952: 42 (type species: brouni).
— Puthz, 1980j: 20 (characters).
— Newton, 1985: 206 (characters).
— Klimaszewski, Newton, and Thayer, 1996: 150 (endemic to New Zealand).
— Newton, 1996: 159 (notes).
bisulciceps Broun, 1917: 374 (Dimerus; Type locality: Mos Basin, Canterbury).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Bernhauer, 1939f: 214 (Agnosthaetus; characters in key).
▴ Distribution: New Zealand.
brouni Bernhauer, 1939f: 215 (Agnosthaetus; Type locality: Neu-Seeland).
— Kuschel, 1990: 26 (Agnosthaetus; endemic; flightless; habitat; New Zealand).
▴ Distribution: New Zealand.
cariniceps Bernhauer, 1939f: 214 (Agnosthaetus; Type locality: Neu-Seeland: South Island: Routeburn, north of Lake Wakatipu).
stilbus Broun, 1910: 15 (Dimerus; Type locality: Waimarino).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; cited as stielus; catalog).
— Bernhauer, 1939f: 214 (Agnosthaetus; characters in key).
— Puthz, 1980j: 20 (Agnosthaetus; characters).
▴ Distribution: New Zealand.
vicinus Broun, 1921: 483 (Dimerus; Type locality: Kiwi Bush, near Glenhope).
Gerhardia
[7 species; Ethiopian Region]
Gerhardia Kistner, 1960: 32 (species included: africana). Type species: Stenaesthetus africana Bernhauer, fixed by original designation and monotypy.
—Puthz, 1980j: 20 (synonym of Stenaesthetus).
— Orousset, 1988: 146 (synonym of Stenaesthetus).
— Orousset, 1990: 11 (synonym of Stenaesthetus).
— Puthz, 1995a: 120 (valid genus; characters; list of species; discussion of genital structures).
afra Kistner, 1961: 33 (Stenaesthetus; Type locality: Congo: Parc National de la Garamba, Mabanga).
— Kistner, 1962: 98 (Stenaesthetus; characters; Congo).
— Puthz, 1974s: 767 (Stenaesthetus; characters; Chad).
— Puthz, 1984g: 202 (Stenaesthetus; Ghana).
— Puthz, 1995a: 122 (Gerhardia).
▴ Distribution: Chad, Congo, Ghana, Senegal. senegalensis Kistner, 1962: 99 (Stenaesthetus; Type locality: Senegal: Badoumbé).
— Jacobson and Kistner, 1974: 90 (Stenaesthetus; aedeagal characters; Ghana).
— Puthz, 1995a: 123 (Gerhardia; synonym of afer).
africana Bernhauer, 1915d: 106 (Stenaesthetus; Type locality: Deutsch-Ostafrika: Westl. Kilimandscharo, 1400–1500 m).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Kistner, 1960: 31, 34 (Gerhardia; notes; charactes; Tanganyika; Belgian Congo).
— Puthz, 1995a: 123 (Gerhardia).
▴ Distribution: Congo, Rwanda, Tanzania.
ruandae Puthz, 1979b: 176 (Stenaesthetus; Type locality: Rwanda: Kayove, 2100 m).
—Puthz, 1980j: 20 (Stenaesthetus; synonym of africanus).
— Puthz, 1995a: 123 (Gerhardia; synonym of africanus).
arrowi Bernhauer, 1927: 51 (Stenaesthetus; Type locality: Natal: Kloof, 1500 ft.).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Kistner, 1962: 100 (Stenaesthetus; characters; South Africa).
— Kistner, 1967: 52 (Stenaesthetus; South Africa).
— Puthz, 1995a: 123 (Gerhardia).
▴ Distribution: South Africa.
coiffaiti Puthz, 1986b: 118 (Stenaesthetus; Type locality: Gabun: Belinga. Also cited from Ivory Coast).
gerardi Bernhauer, 1929f: 108 (Stenaesthetus; Type locality: Elisabethville: Pweto).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Kistner, 1960a: 116 (Stenaesthetus; Belgian Congo).
— Kistner, 1962: 100 (Stenaesthetus; characters; Congo Republic; Ruanda; Angola; Northern Rhodesia; Southern Rhodesia; Sierra Leone).
— Kistner, 1967: 52 (Stenaesthetus; Zambia).
— Puthz, 1995a: 123 (Gerhardia).
▴ Distribution: Congo, Congo Republic; Ruanda; Angola; Northern Rhodesia; Southern Rhodesia; Sierra Leone, Zambia.
lohsei Puthz, 1995a: 124 (Gerhardia; Type locality: Zaire: Parc National Albert, Secteur Tshiaberima, riv. Talya Nord, 2340 m, affl. g. Semliki).
natalensis Puthz, 1979b: 175 (Stenaesthetus; Type locality: Natal).
ruandae Puthz, 1979, see: africanus Bernhauer, 1915.
senegalensis Kistner, 1962, see: afer Kistner, 1961.
Stenaesthetus
[70 species; Neotropical, Palaearctic, Ethiopian, Madagascan, and Oriental Regions]
Stenaesthetus Sharp, 1874: 79 (species included: sunioides). Type species: Stenaesthetus sunioides Sharp, fixed by monotypy.
— Sharp, 1876: 356 (characters; notes).
— Duvivier, 1883: 179 (catalog).
— Bernhauer and Schubert, 1911: 186 (world catalog; 3 species).
— Cameron, 1921a: 352, 401 (characters; key to Singapore species; catalog).
— Winkler, 1925: 357 (catalog for Palaearctic region).
— Cameron, 1925: 31 (catalog of Indian species).
— Cameron, 1930: 445 (characters; key to species of British India).
— Scheerpeltz, 1933: 1202 (world catalog supplement).
— Blackwelder, 1943: 228 (type species: sunioides).
— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 357 (type species: sunioides).
— Kistner, 1962: 96 (revision of Old World species; key Oriental and Ethiopian species; characters).
— Puthz, 1973k: 66 (notes; key to some New World species).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Scheerpeltz, 1974: 101 (characters).
— Shibata, 1976: 209 (checklist of species of Japan).
— Puthz, 1980j: 17 (characters; notes; synonyms).
— Orousset, 1988: 146 (characters; key to species of Madagascar; notes on world distribution and biology).
— Orousset, 1990: 11 (characters; revision of and key to species of neotropics).
— Puthz, 1995b: 183 (key to species of China).
Aulacosthaetus Bernhauer, 1939f: 211 (species included: klimai, nitidiceps, rambouseki, pauloensis, mrazi). Type species: Aulacosthaetus rambouseki Bernhauer, by original designation.
— Blackwelder, 1944: 115 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 70 (type species: rambouseki).
— Kistner, 1965: 358 (characters; revision of and key to species).
— Scheerpeltz, 1974: 103 (cited as Aulacoaesthetus; characters).
— Puthz, 1980j: 20 (synonym of Stenaesthetus).
— Orousset, 1988: 146 (synonym of Stenaesthetus).
— Orousset, 1990: 11 (synonym of Stenaesthetus).
aethiopicus Puthz, 1988f: 12 (Stenaesthetus; Type locality: Ethiopia: Kaffa Province: Belleta F. 40 km S. W. Jimma, 2100 m).
afer Kistner, 1961, see: Gerhardia.
africanus Bernhauer, 1915, see: Gerhardia.
amedegnatoae Orousset, 1990: 36 (Stenaesthetus; Type locality: Pérou: Estiron, Rio Ampiyacu. Also cited from Colombia).
anjavidilava Orousset, 1988: 177 (Stenaesthetus; Type locality: Madagascar Centre: massif de l’Andringitra, Anjavidilava, 2000 m).
apterus Puthz, 1988f: 12 (Stenaesthetus; Type locality: Ethiopia: Wolamo Province: spur of Mt. Damota, over 8000 feet).
arrowi Bernhauer, 1927, see: Gerhardia.
atrox Orousset, 1988: 171 (Stenaesthetus; Type locality: Madagascar Est: région de Périnet, station du lac Vert).
badius Orousset, 1988: 182 (Stenaesthetus; Type locality: Madagascar Est: massif du Marojejy, 600 m).
bernhaueri Orousset, 1990: 42 (Stenaesthetus; Type locality: Brésil: état de São Paulo).
— Kistner, 1965: 361 (Aulacosthaetus; [Note: Misidentified as mrazi, according to Orousset, 1990: 42]; characters; Brazil).
▴ Distribution: Brazil.
betschi Orousset, 1988: 177 (Stenaesthetus; Type locality: Madagascar Centre: massif de l’Andringitra, chaîne du Vohidray, 2000 m).
brasilianus Orousset, 1983, see: fauveli Puthz, 1979.
campbelli Orousset, 1990: 48 (Stenaesthetus; Type locality: Brésil: état de São Paulo, Salesopolis).
canaliculatus Coiffait, 1975a: 185 (Stenaesthetus; Type locality: Phulchoki près Katmandou).
taurulus Puthz, 1987a: 447 (Stenaesthetus; subspecies of canaliculatus; Type locality: Nepal: Kathmandu-Tal, Nagarjung, Berg Jamacok, 1900–2100 m).
castaneus Orousset, 1990: 24 (Stenaesthetus; Type locality: Paraguay: dep. de Concepcion, à l’est de Estrellas. Also cited from Peru and Brazil).
coiffaiti Puthz, 1986, see: Gerhardia.
comoranus Lecoq, 1996: 135 (Stenaesthetus; Type locality: l’archipel des Comores: Mohéli, Miringoni).
condei Wendeler, 1930: 184 (Stenaesthetus; Type locality: Brasilien, Espirito Santo, Sta. Thereza).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Orousset, 1990: 45 (Stenaesthetus; characters; Brazil).
▴ Distribution: Brazil.
conflictatus Puthz, 1995b: 177 (Stenaesthetus; Type locality: China: Yunnan: Xishuangbanna).
confragrosus Orousset, 1990: 23 (Stenaesthetus; Type locality: Brésil: état d’Amazonas: 20 km au S.O. d’Itapiranga).
deharvengi Orousset, 1988c: 183 (Stenaesthetus; Type locality: Thaïlande: env. de Chiang Maï, Doï Inthanon, alt. 2000 m).
descarpentriesi Orousset, 1988: 173 (Stenaesthetus; Type locality: Madagascar Centre: massif de l’Andringitra, cirque de Manjarivolo, 1650 m).
dolosus Orousset, 1988: 163 (Stenaesthetus; Type locality: Madagascar Est: baie d’Antongil, région de Maroantsetra, Ambodivoangy).
dunayi Janák, 1996a: 337 (Stenaesthetus; Type locality: Madagascar Centre: Ankaratra: Mandritsara, Route Antanifotsy - Faratsiho, 1800–2000 m).
elegans Orousset, 1988: 186 (Stenaesthetus; Type locality: Madagascar Est: route de Lakato, 1100 m).
eximius Orousset, 1990: 46 (Stenaesthetus; Type locality: Brésil: état de São Paulo, Salesopolis).
fauveli Puthz, 1979b: 174 (Stenaesthetus; Type locality: Antilles: St. Vincent).
— Orousset, 1990: 27 (Stenaesthetus; characters; St. Vincent; Brazil).
▴ Distribution: Brazil, West Indies.
brasilianus Orousset, 1983b: 118 (Stenaesthetus; Type locality: Matto-Grosso, Brésil).
frontalis Cameron, 1950d: 9 (Aulacosthaetus; Type locality: Selangor: The Gap).
— Kistner, 1965: 360, 367 (Aulacosthaetus; characters; Malaya).
— Puthz, 1980j: 20 (Stenaesthetus).
▴ Distribution: Malaysia.
gerardi Bernhauer, 1929, see: Gerhardia.
griveaudi Jarrige, 1968: 873 (Aulacosthaetus; Type locality: Madagascar-Est, Ambinanitelo, Marojéjy, distr. de Sambava, 500 m).
— Puthz, 1980j: 20 (Stenaesthetus).
— Orousset, 1988: 161 (Stenaesthetus; characters; Madagascar).
▴ Distribution: Madagascar.
haribe Janák, 1996: 190 (Stenaesthetus; Type locality: Madagascar Nord: Massiv Anjana-haribe Nord, sentier Ambodihasina-Ambalarombe, crête, 1250–1300 m).
heilongjiangensis J. Li, 1993: 55 (Stenaesthetus; Type locality: Mount Laoye, Shangzhi, Heilongjiang).
horrifer Orousset, 1988: 161 (Stenaesthetus; Type locality: Madagascar Est: baie d’Antomgil, région de Maroantsetra, Fampanambo).
illatus Sharp, 1876: 357 (Stenaesthetus; Type locality: Tapajos, Brazil; [Note: See lectotype designation by Orousset, 1990]. Also cited from Lages near Manaos).
— Bernhauer and Schubert, 1911: 186 (Stenaesthetus; catalog).
— Puthz, 1973k: 67 (Stenaesthetus; characters; Brazil).
— Irmler, 1979b: 6, 7 (Stenaesthetus; ecological notes).
— Orousset, 1990: 20 (Stenaesthetus; lectotype designation: Tapajos, Brazil; characters; Brazil; Peru; Paraguay).
▴ Distribution: Brazil, Peru, Paraguay.
imitator Orousset, 1988: 158 (Stenaesthetus; Type locality: Madagascar Est: baie d’antongil, Nosy Mangabe).
immarginatus Erichson, 1840: 748 (Euaesthetus; Type locality: Vallis Araguensis Columbiae).
— Bernhauer and Schubert, 1911: 189 (Euaesthetus; catalog).
— Blackwelder, 1943: 228 (Stenaesthetus; St. Vincent).
— Puthz, 1973k: 67 (Stenaesthetus; characters; Colombia).
— Orousset, 1990: 27 (Stenaesthetus; characters; Colombia).
▴ Distribution: Colombia, West Indies.
immigratorius Kistner, 1967: 52 (Stenaesthetus; Type locality: Republic of South Africa: Transvaal, Pietersburg District, Woodbush Forest, Calyey ravine).
indicus Cameron, 1943: 5 (Aulacosthaetus; Type locality: Darjeeling district: Ghum).
— Kistner, 1965: 365 (Aulacosthaetus; characters; India).
— Puthz, 1980j: 20 (Stenaesthetus).
▴ Distribution: India.
kistneri Jarrige, 1968, see: rugosus Kistner, 1965.
klimai Bernhauer, 1939f: 212 (Aulacosthaetus; Type locality: Brasilien: Minas Geraës).
— Kistner, 1965: 363 (Aulacosthaetus; characters; Brazil).
— Puthz, 1980j: 20 (Stenaesthetus).
— Orousset, 1990: 49 (Stenaesthetus; characters; Brazil).
▴ Distribution: Brazil.
kraatzi Bernhauer, 1908d: 300 (Stenaesthetus; Type locality: Paraguay).
— Bernhauer and Schubert, 1911: 186 (Stenaesthetus; catalog).
— Puthz, 1973k: 67, 68 (Stenaesthetus; characters; Paraguay).
— Orousset, 1990: 32 (Stenaesthetus; characters; Paraguay).
▴ Distribution: Paraguay.
leleupi Kistner, 1967: 54 (Aulacosthaetus; Type locality: Republic of South Africa, Cape Province, Peninsula, Table Mountain, Bats Cave).
— Puthz, 1979b: 177 (Aulacosthaetus; characters; South Africa).
— Puthz, 1980j: 20 (Stenaesthetus).
▴ Distribution: South Africa.
lilliputanus Orousset, 1990: 51 (Stenaesthetus; Type locality: Brésil: état de São Paulo).
llinaresi Orousset, 1988: 188 (Stenaesthetus; Type locality: Madagascar Centre: pays Betsileo, Ambatofitoharana, RN 7, pk 292, 1600 m).
machomenos Orousset, 1988: 185 (Stenaesthetus; Type locality: Madagascar Centre: Est du lac de Mantasoa, forêt d’Ambohiboatavo).
malayanus Puthz, 1986a: 297 (Stenaesthetus; Type locality: Malaysia: Pahang: Cameron Highlands, Ringlette, 3200 feet).
marojejy Orousset, 1988: 180 (Stenaesthetus; Type locality: Madagascar Est: massif du Marojejy, 600 m).
martensi Puthz, 1987a: 445 (Stenaesthetus; Type locality: Nepal: Manang Distr.: Thimang oberhalb Bagarchap, 2550 m).
mendax Orousset, 1988: 169 (Stenaesthetus; Type locality: Madagascar Est: presqu’île de Masoala, Ambohitsitondroina, 1100 m).
microphtalmus Jarrige, 1968: 873 (Aulacosthaetus; Type locality: Madagascar-Est, Ambodivoangy).
microphthalmus Orousset, 1988: 163 (Stenaesthetus; emendation of microphtalmus).
microphthalmus Orousset, 1988, see: microphtalmus Jarrige, 1968.
minax Orousset, 1988: 175 (Stenaesthetus; Type locality: Madagascar Centre: massif de l’Andringitra, base de la forêt du cirque de Manjarivolo, 1200 m).
miskoi Janák, 1996a: 340 (Stenaesthetus; Type locality: Madagascar Est: P. N. Ranomafana env., Sahavondrona).
monticola Orousset, 1988: 158 (Stenaesthetus; Type locality: Madagascar Est: presqu’île de Masoala, Ambohitsitondroina, 1100 m).
mrazi Rambousek, 1915: 54 (Stenaesthetus; Type locality: São Paolo, Brazil; [Note: See lectotype designation by Orousset, 1990]).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Bernhauer, 1939f: 212 (Aulacosthaetus; characters).
— Puthz, 1980j: 20 (Stenaesthetus).
— Orousset, 1990: 39 (Stenaesthetus; lectotype designation; characters; Brazil).
▴ Distribution: Brazil.
rambouseki Bernhauer, 1939f: 212 (Aulacosthaetus; Type locality: Not cited).
— Kistner, 1965: 363 (Aulacosthaetus; characters; Brazil).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1990: 39 (Stenaesthetus; synonym of mrazi).
pauloensis Bernhauer, 1939f: 212 (Aulacosthaetus; Type locality: Brasilien: Minas Geraës).
— Kistner, 1965: 363 (Aulacosthaetus; synonym of rambouseki).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1990: 39 (Stenaesthetus; synonym of mrazi).
natalensis Puthz, 1979, see: Gerhardia.
nepalensis Puthz, 1990e: 65 (Stenaesthetus; Type locality: Nepal: near Sanghu).
nitidiceps Bernhauer, 1939f: 212 (Aulacosthaetus; Type locality: Brasilien: Minas Geraës).
— Kistner, 1965: 363 (Aulacosthaetus; synonym of rambouseki).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1990: 43 (Stenaesthetus; valid species; characters; Brazil).
▴ Distribution: Brazil.
nitidifrons Puthz, 1987a: 447 (Stenaesthetus; Type locality: Nepal: Kaski Distr., oberhalb Dhumpus, 2100 m).
nomurai Puthz, 1995b: 181 (Stenaesthetus; Type locality: Taiwan: Taoyuan Hsien: Paling).
okinawaensis Puthz, 1986a: 299 (Stenaesthetus; Type locality: Japan: Okinawa: Yonaha-dake).
panamensis Puthz, 1973k: 68 (Stenaesthetus; Type locality: Panama: Canal Zone: Bohio).
pauliani Jarrige, 1968: 872 (Aulacosthaetus; Type locality: Madagascar-Est: Ambodivoangy, distr. de Maroantsétra).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1988: 185 (Stenaesthetus; characters; Madagascar).
▴ Distribution: Madagascar.
pauloensis Bernhauer, 1939, see: mrazi Rambousek, 1915.
peruanus Orousset, 1990: 35 (Stenaesthetus; Type locality: Pérou: dep. de Loreto, Genaro Herrera, Rio Ucayali).
peyrierasi Orousset, 1988: 171 (Stenaesthetus; Type locality: Madagascar Est: région de Périnet, forêt de la Grande Ile, 900 m).
praestans Orousset, 1990: 29 (Stenaesthetus; Type locality: Brésil: état de Santa Catarina: Nova Teutonia, alt. 300–500. Also cited from Paraguay).
puthzi Orousset, 1990: 34 (Stenaesthetus; Type locality: Pérou: dép. de Cuzco: Consuelo, route de Manu, km 165. Also cited from Brazil and Ecuador).
quadrisulcatus Cameron, 1930: 446 (Stenaesthetus; Type locality: Almora: Kali Valley, alt. 9000 feet).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Cameron, 1943: 5 (Aulacosthaetus; transferred from Stenaesthetus).
— Cameron, 1950d: 9 (transfer to Aulacosthaetus).
— Kistner, 1965: 360, 367 (Aulacosthaetus; characters; notes).
— Abdullah and Qadri, 1968a: 309 (Stenaesthetus; characters; West Pakistan).
— Coiffait, 1975a: 156 (Stenaesthetus; Nepal).
— Puthz, 1980j: 21 (Stenaesthetus).
— Coiffait, 1982: 37 (Stenaesthetus; Nepal).
— Puthz, 1987a: 443 (Stenaesthetus; characters; notes; Nepal).
▴ Distribution: India, Nepal, Pakistan.
rambouseki Bernhauer, 1939, see: mrazi Rambousek, 1915.
robinsoni Orousset, 1988: 157 (Stenaesthetus; Type locality: Madagascar Nord: 50 km au Sud-Est de Diégo-Suarez, forêt d’Analamerana, 80 m).
ruandae Puthz, 1979, see: Gerhardia.
rugatulus Orousset, 1988: 159 (Stenaesthetus; Type locality: Madagascar Est: baie d’Antongil, région de Maroantsetra, Ambodivoangy).
rugosus Kistner, 1965: 367 (Aulacosthaetus; Type locality: East Madagascar: Baie d’Antongil).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1988: 156 (Stenaesthetus; characters; Madagascar).
▴ Distribution: Madagascar.
kistneri Jarrige, 1968: 873 (Aulacosthaetus; Type locality: Madagascar-Est, Ambodivoangy, distr. de Maroant setra).
schuhi Orousset, 1990: 31 (Stenaesthetus; Type locality: Brésil: état de Para: 5 km à l’est de Belém).
scotti Puthz, 1988f: 9 (Stenaesthetus; Type locality: Ethiopia: Gamo Province, Gughé highlands, south of Bonghé, ca. 9500–10,000 feet).
senegalensis Kistner, 1962, see: Gerhardia.
sogai Jarrige, 1968: 871 (Aulacosthaetus; Type locality: Madagascar-Est: R. N. XII, Col Central, 1700 m, Marojéjy, distr. de Sambava).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1988: 180 (Stenaesthetus; characters; Madagascar).
▴ Distribution: Madagascar.
sunioides Sharp, 1874: 80 (Stenaesthetus; Type locality: Mitzuyama [Japan]; North China).
— Bernhauer and Schubert, 1911: 186 (Stenaesthetus; catalog).
— Cameron, 1921a: 369, 401 (Stenaesthetus; characters; Singapore).
— Cameron, 1930: 446 (Stenaesthetus; characters; Ceylon; India; Sierra Leone).
— Scheerpeltz, 1933: 1202 (Stenaesthetus; catalog).
— Watanabe, 1961: 350 (Stenaesthetus; Izu Islands).
— Kistner, 1962: 98 (Stenaesthetus; characters; China; Formosa; Japan; India).
— Nakane, 1963a: 87 (Stenaesthetus; characters; habitus photograph; Japan).
— Abdullah and Qadri, 1968a: 309 (Stenaesthetus; characters; West Pakistan).
— Watanabe and Shibata, 1972: 61 (Stenaesthetus; Yaku-shima Island).
— Puthz, 1973k: 67 (Stenaesthetus; characters).
— Coiffait, 1982: 37 (Stenaesthetus; Nepal).
— J. Li, 1993: 26 (Stenaesthetus; China).
— Puthz, 1995b: 180, 183 (Stenaesthetus; characters; Taiwan; China).
▴ Distribution: China, Japan, Taiwan, Nepal, Pakistan, India, Sri Lanka, Singapore. taurulus Puthz, 1987, see: canaliculatus Coiffait, 1975.
taiwanensis Puthz, 1995b: 180 (Stenaesthetus; Type locality: Taiwan: Ilan Hsien: Taiping-shan).
thrips Puthz, 1986b: 117 (Stenaesthetus; Type locality: Kenya: Taita Hills, Umgebung Wundanyi, 1350 m).
vadoni Kistner, 1965: 369 (Aulacosthaetus; Type locality: East Madagascar: Baie d’Antongil).
— Puthz, 1980j: 21 (Stenaesthetus).
— Orousset, 1988: 167 (Stenaesthetus; characters; Madagascar).
▴ Distribution: Madagascar.
viduus Puthz, 1986b: 118 (Stenaesthetus; Type locality: Elfenbeinküste: 7–17 km W. Abidjan).
Tyrannomastax
[2 species; Madagascan Region]
Tyrannomastax Orousset, 1988: 189 (species included: hova, excelsus). Type species: Tyrannomastax hova Orousset, fixed by original designation.
excelsus Orousset, 1988: 196 (Tyrannomastax; Type locality: Madagascar Est: Sud-Est de Maropaika, forêt de Mahabo, entre Ivohibe et Vondrozo).
hova Orousset, 1988: 193 (Tyrannomastax; Type locality: Madagascar Est: Est de Vondrozo, col de Madiorano, 700 m).
LEPTOTYPHLINAE
[43 genera; 515 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Oriental, and Australian Regions]
Leptotyphlinae Fauvel, 1874.
—Eichelbaum, 1909: 130 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).
— Bernhauer and Schubert, 1911: 146 (world catalog; Leptotyphlus, Mayetia, Cyrtotyphlus).
— Porta, 1926: 49 (characters; species of Italy).
— Portevin, 1929: 391 (characters; key to French genera).
— Scheerpeltz, 1933: 1138 (world catalog supplement).
— Scheerpeltz, 1940: 28 (key to Palaearctic genera).
— Coiffait, 1955c (notes on distribution).
— Coiffait, 1956d (notes).
— Horion, 1963: 295 (notes; list of central European species).
— Lohse, 1964: 106 (characters; key to central European genera and species).
— Moore, 1964a: 87 (characters; notes).
— Coiffait and Saiz, 1965a: 136 (key to Chilean genera).
— Coiffait, 1963d: 380, 381 (key to tribes; biogeographic notes for each tribe).
— Coiffait and Saiz, 1968: 346 (list of Chilean species).
— Scheerpeltz, 1968a: 34 (catalog of Austrian species).
— Coiffait, 1972: 381 (characters; key to tribes, genera and species of western Palaearctic region).
— Coiffait and Decou, 1972: 136 (list of genera of Cuba).
— Decou, 1973: 360 (list of genera of Cuba; comments on distribution).
— Saiz, 1973b (characteristics of edaphic species; dendrogram of genera; distribution).
— Moore and Legner, 1974g: 553 (key to genera of America north of Mexico).
— Saiz, 1974: 47 (revision and key to genera and species of Chile).
— Pace, 1975a: 110 (supplement to key to tribes of Coiffait, 1972).
— Moore and Legner, 1975: 268 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 249 (characters; tabular key to genera of North America north of Mexico).
— Frank and Thomas, 1984a: 1413 (checklist of New World taxa).
— Orousset, 1984: 297 (comments on history of group; checklist of species in Corsica).
— Outerelo and Gamarra, 1985: 15 (key to genera of Iberian Peninsula).
— Smetana, 1986: 181 (discussion of distributions in far north of northern Hemisphere).
— Frank, 1991: 346 (larvae undescribed).
— Newton and Thayer, 1992: 13, 56 (summary of current classification).
— Lawrence and Newton, 1995: 829 (notes).
— Sparacio, 1995: 159 (characters; species of Sicily).
— M. Hansen, 1996: 37 (listed in Staphylinine group of subfamilies).
— Pace, 1996 (systematic position; morphology of adult and larva; general natural history and behavior; distribution; key to tribes, genera, species of Italy; review of Italian fauna).
— M. Hansen, 1997: 80, 81, 130, 175, 177, 180 (characters supporting clade; autapomorphic features; key adult and larval characters; notes).
— Newton, Thayer, Ashe, and Chandler, 2000: 323, 382 (key to Nearctic genera; characters; notes).
— Klimaszewski, 2000: 62 (characters; notes on classification, natural history, and distribution in Alaska).
Tribe Cephalotyphlini
Cephalotyphlini Coiffait, 1963d: 380, 381 (genera included: Cephalotyphlus). Type genus: Cephalotyphlus (see Newton and Thayer, 1992: 56).
Cephalotyphlus
[5 species; Palaearctic Region]
Cephalotyphlus Coiffait, 1955a: 68 (species included: revelieri). Type species: Leptotyphlus revelieri Saulcy, fixed by original designation and monotypy.
— Coiffait, 1957: 61 (characters).
— Coiffait, 1959: 322 (characters; type species: revelieri).
— Coiffait, 1972: 490 (characters; type species: revelieri; key to species of western Palaearctic region; references).
— Orousset, 1984: 300 (list of species of Corsica).
— Pace, 1975a: 110 (characters).
— Zanetti, 1995: 14 (list of species of Italy).
— Pace, 1996: 60 (characters; key to species of Italy).
Cyclotyphlus Scheerpeltz, 1955b: 461 (species included: revelieri, cribriceps, and one unavailable name). Type species: Leptotyphlus revelieri Saulcy, fixed by original designation.
— Coiffait, 1959: 322 (synomym of Cephalotyphlus; type species: revelieri).
— Coiffait, 1972: 490 (synonym of Cephalotyphlus).
— Pace, 1996: 60 (synonym of Cephalotyphlus).
Italicotyphlus Pace, 1975a: 104 (species included: vicetinus). Type species: Italicotyphlus vicetinus Pace, fixed by original designation.
bergamascus Pace, 1991: 112 (Cephalotyphlus; Type locality: Alpi Bergamasche, Oltre il Colle).
— Scheerpeltz, 1955: 461 (Cyclotyphlus; nomen nudum).
— Zanetti, 1995: 14 (Cephalotyphlus; Italy).
— Pace, 1996: 67 (Cephalotyphlus; characters; Italy).
▴ Distribution: Italy.
florentinus Pace, 1979b: 435 (Italicotyphlus; Type locality: versante nord occcidentale del Monte Morello, nord-ovest di Firenze, presso il villaggio di Volmiano, a circa 150 metri d’altitudine).
— Pace, 1991: 112 (Cephalotyphlus).
— Zanetti, 1995: 14 (Cephalotyphlus; Italy).
— Pace, 1996: 67 (Cephalotyphlus; characters; Italy).
▴ Distribution: Italy.
foroiuliensis Pace, 1976: 114 (Italicotyphlus; Type locality: versante settentrionale del M. Jof, presso Maniago a circa 410 m).
— Pace, 1991: 112 (Cephalotyphlus).
— Zanetti, 1995: 14 (Cephalotyphlus; Italy).
— Pace, 1996: 71 (Cephalotyphlus; characters; Italy).
▴ Distribution: Italy.
revelierei Saulcy, 1878: 120 (Leptotyphlus; Type locality: Omessa).
— Croissandeau, 1891: 150 (Leptotyphlus; characters; Corsica).
— Sainte-Claire Deville, 1906: 78 (Leptotyphlus; Corsica).
— Bernhauer, 1908e: 332 (Leptotyphlus; characters).
— Dodero, 1908: 634, 637 (Leptotyphlus; characters; Italy).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Dodero, 1916: 341 (Leptotyphlus; Sardinia).
— Peyerimhoff, 1926: 119 (Leptotyphlus; characters).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1955a: 68 (Cephalotyphlus).
— Scheerpeltz, 1955b: 461 (Cyclotyphlus).
— Coiffait, 1959: 324 (Cephalotyphlus; notes; characters; Corsica, Sardinia).
— Coiffait, 1972: 491 (Cephalotyphlus; characters; Corsica).
— Pace, 1991: 112 (Cephalotyphlus).
— Pace, 1996: 65 (Cephalotyphlus; characters; Corsica).
▴ Distribution: Italy, Corsica, Sardinia.
vicetinus Pace, 1975a: 106 (Italicotyphlus; Type locality: nord-est dalla città di Marostica, provincia di Vicenza, 160 m).
Tribe Entomoculiini
Entomoculiini Coiffait, 1957: 61 (characters; key to genera; genera included: Entomoculia, Stenotyphlus, Allotyphlus, Cephalotyphlus, Cyrtotyphlus, Mesotyphlus, Paratyphlus, Ischnotyphlus). Type genus: Entomoculia (see Newton and Thayer, 1992: 56).
— Coiffait, 1959: 274 (characters; key to genera; morphology; discription).
— Coiffait, 1960a: 512 (biogeography).
— Coiffait, 1963d: 380, 381 (characters; biogeographic notes).
— Coiffait, 1972: 383, 385 (characters; key to genera and species of western Palaearctic region).
— Saiz, 1973b: 289 (distribution).
— Pace, 1975a: 110 (characters).
— Orousset, 1984: 300 (list of taxa of Corsica).
— Outerelo and Gamarra, 1985: 15 (key to genera of Iberian Peninsula).
— Newton and Thayer, 1992: 56 (tribe of Leptotyphlinae).
— Pace, 1996: 59, 60, 71 (characters; key to genera).
Allotyphlus
[22 species; Palaearctic Region]
Allotyphlus Coiffait, 1955a: 67 (species included: aspericeps). Type species: Leptotyphlus aspericeps Scheerpeltz, fixed by original designation and monotypy.
— Coiffait, 1957: 61 (characters).
— Coiffait, 1959: 310 (characters; key to species; type species: aspericeps).
— Coiffait, 1972: 450 (characters; type species: aspericeps; key to species of western Palaearctic region; references).
— Pace, 1977: 197 (key to species of Apennines).
— Zanetti, 1995: 14 (list of species of Italy).
— Pace, 1996: 124 (characters; key to species of Italy).
Tropidotyphlus Scheerpeltz, 1955b: 459 (species included: aspericeps along with 4 unavailable names). Type species: Leptotyphlus aspericeps Scheerpeltz, fixed by original designation and monotypy.
— Coiffait, 1959: 310 (synomym of Allotyphlus; type species: aspericeps).
— Coiffait, 1972: 450 (synonym of Allotyphlus).
— Pace, 1996: 124 (synonym of Allotyphlus).
Moreotyphlus Coiffait, 1959: 310, 312 (subgenus of Allotyphlus; species included: spartensis, naupliensis, patrensis). Type species: Allotyphlus (Moreotyphlus) spartensis Coiffait, fixed by original designation.
— Coiffait, 1972: 450, 452, 454 (subgenus of Allotyphlus; characters; type species: spartensis; key to species of western Palaearctic region).
— Pace, 1996: 131 (subgenus of Allotyphlus; key to species of Italy).
achileus Coiffait, 1959: 402 (Allotyphlus; subgenus Allotyphlus; Type locality: Corfou: Ayos Deka).
ambracius Scheerpeltz, 1958c: 392 (Tropidotyphlus; Type locality: Umgebung von Arta im südlichen Epirus).
aspericeps Scheerpeltz, 1931: 396 (Leptotyphlus; Type locality: Kaligoni auf Levkas).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1955a: 67 (Allotyphlus).
— Scheerpeltz, 1955b: 459 (Tropidotyphlus).
— Coiffait, 1959: 312 (Allotyphlus; characters; Ionian Islands).
— Coiffait, 1972: 453 (Allotyphlus; characters; Ionian Islands).
— Pace, 1983a: 454 (Allotyphlus; subgenus Moreotyphlus; characters).
▴ Distribution: Ionian Islands.
baldensis Pace, 1978, see: pacei Coiffait, 1973.
bericiensis Coiffait, 1973: 110 (Allotyphlus; subgenus Moreotyphlus; Type locality: Lumignano, Monti Berici, Italie).
— Pace, 1975b: 9 (Allotyphlus; subgenus Moreotyphlus; characters).
— Zanetti, 1995: 14 (Allotyphlus; Italy).
— Pace, 1996: 132 (Allotyphlus; subgenus Moreotyphlus; characters; Italy).
▴ Distribution: Italy.
bipunctatus Coiffait, 1973: 110 (Mesotyphlus; Type locality: Zovencedo, Monti Berici, Italie).
bipunctatus Coiffait, 1973, see: bericiensis Coiffait, 1973.
cephallenicus Pace, 1983a: 456 (Allotyphlus; subgenus Moreotyphlus; Type locality: Cephallinia, M. te Rudi).
corcyranus Coiffait, 1959: 401 (Allotyphlus; subgenus Allotyphlus; Type locality: Corfou: Dassia).
corcyricus Scheerpeltz, 1958c: 390 (Tropidotyphlus; Type locality: Val de Ropa auf der Insel Korfu).
— Coiffait, 1972: 454 (Allotyphlus; characters; Corfu).
— Pace, 1983a: 452 (allotyphlus; characters; Corfu).
▴ Distribution: Greece.
dexter Coiffait, 1959: 402 (Allotyphlus; subgenus Allotyphlus; Type locality: Corfou: Doukades).
euganeus Pace, 1975, see: pacei Coiffait, 1973.
garganicus Pace, 1975: 144 (Allotyphlus; subgenus Moreotyphlus; Type locality: Puglia sul Promontorio del Gargano, ai margini della Foresta Umbra, lungo la strada che scende a Vico Garganico, a circa 400 metri d’altitudine).
— Pace, 1977: 175, 198 (Allotyphlus; characters; Italy).
— Pace, 1985: 62 (Allotyphlus; characters).
— Zanetti, 1995: 14 (Allotyphlus; Italy).
— Pace, 1996: 137 (Allotyphlus; subgenus Moreotyphlus; characters; Italy).
▴ Distribution: Italy.
loebli Coiffait, 1973b: 220 (Allotyphlus; subgenus Moreotyphlus; Type locality: Grèce, Epire, Philiates).
lessiniensis Coiffait, 1973, see: pacei Coiffait, 1973.
marosticaensis Pace, 1978, see: pacei Coiffait, 1973.
moczarskii Pace, 1983a: 457 (Allotyphlus; subgenus Moreotyphlus; Type locality: Cephalonia, Argostoli).
monsfortensis Pace, 1978, see: pacei Coiffait, 1973.
montanus Coiffait, 1957: 67 (Allotyphlus; Type locality: Turquie d’Asie: Près d’une source au point culminant de la route d’Antalya à Burdur dans la traversée du Darim Dag).
— Coiffait, 1959: 313 (Allotyphlus; characters; Turkey).
— Coiffait, 1972: 453 (Allotyphlus; characters; Turkey).
▴ Distribution: Turkey.
naupliensis Coiffait, 1959: 404 (Allotyphlus; subgenus Moreotyphlus; Type locality: Pélopon-nèse: Yannouleika près de Nauplion).
— Coiffait, 1972: 456 (Allotyphlus; subgenus Moreotyphlus; characters; Greece).
▴ Distribution: Greece.
pacei Coiffait, 1973: 110 (Allotyphlus; subgenus Moreotyphlus; Type locality: Villaga Toara, Monti Berici, Italie).
— Coiffait, 1973b: 224 (Allotyphlus; characters).
— Pace, 1975b: 4, 10 (Allotyphlus; characters; notes).
— Pace, 1978a: 199 (Allotyphlus; characters; notes; Italy).
— Zanetti, 1995: 14 (Allotyphlus; Italy).
— Pace, 1996: 132 (Allotyphlus; subgenus Moreotyphlus; characters; key to subspecies; Italy).
▴ Distribution: Italy.
lessiniensis Coiffait, 1973: 111 (Mesotyphlus; Type locality: Grezzana, 400 m, Lessini, Italie).
— Pace, 1975b: 7, 10 (Allotyphlus; subspecies of pacei; characters; notes; Italy).
— Zanetti, 1995: 14 (Allotyphlus; subspecies of pacei; Italy).
— Pace, 1996: 132, 136 (Allotyphlus; subgenus Moreotyphlus; subspecies of pacei; characters; Italy).
euganeus Pace, 1975b: 8, 10 (Allotyphlus; subspecies of pacei; Type locality: Colli Euganei, versante settentrionale del Monte Fasolo, 250 metri nel Comune di Lozzo Atestino; versante settentrionale del monte Mottolone a 180 metri d’altitudine nel Comune di Arquà Petrarca).
— Zanetti, 1995: 14 (Allotyphlus; subspecies of pacei; Italy).
— Pace, 1996: 132, 136 (Allotyphlus; subgenus Moreotyphlus; subspecies of pacei; characters; Italy).
baldensis Pace, 1978a: 199 (Allotyphlus; subspecies of pacei; Type locality: Loppio, TN, Dossi della Barchessa, versante settentrionale del Dos Remit, del gruppo del Monte Baldo).
— Zanetti, 1995: 14 (Allotyphlus; subspecies of pacei; Italy).
— Pace, 1996: 132, 136 (Allotyphlus; subgenus Moreotyphlus; subspecies of pacei; characters; Italy).
monsfortensis Pace, 1978a: 200 (Allotyphlus; subspecies of pacei; Type locality: Monteforte d’Alpone, VR, Valle Ponsara; Le Volpare; Soave, VR, Tebaldi; Cazzano di Tramigna, VR; Montecchia di Crosara, VR, M. Bastia, Montebello Vicentio, VI).
— Zanetti, 1995: 14 (Allotyphlus; subspecies of pacei; Italy).
— Pace, 1996: 132, 136 (Allotyphlus; subgenus Moreotyphlus; subspecies of pacei; characters; Italy).
marosticaensis Pace, 1978a: 202 (Allotyphlus; subspecies of pacei; Type locality: Marostica).
patrensis Coiffait, 1959: 404 (Allotyphlus; subgenus Moreotyphlus; Type locality: Pélopon-nèse: Kamarès près de Patras).
— Coiffait, 1972: 456 (Allotyphlus; subgenus Moreotyphlus; characters; Greece).
▴ Distribution: Greece.
pisidicus Coiffait, 1957: 67 (Allotyphlus; Type locality: Turquie d’Asie: Dinar, à 40 km au nord du Lac de Burdur).
— Coiffait, 1959: 313 (Allotyphlus; characters; Turkey).
— Coiffait, 1972: 454 (Allotyphlus; characters; Turkey).
▴ Distribution: Turkey.
sinester Coiffait, 1959: 402 (Allotyphlus; subgenus Allotyphlus; Type locality: Corfou Palaeokastritsa).
sipontinus Pace, 1985: 60 (Allotyphlus; Type locality: Puglia Gargano dintorni di Manfre-donia).
— Zanetti, 1995: 14 (Allotyphlus; Italy).
— Pace, 1996: 130 (Allotyphlus; subgenus Allotyphlus; characters; Italy).
▴ Distribution: Italy.
spartensis Coiffait, 1959: 404 (Allotyphlus; subgenus Moreotyphlus; Type locality: Pélopon-nèse: Tripi près de Sparte, au pied du versant Est du Taygète).
— Coiffait, 1972: 456 (Allotyphlus; subgenus Moreotyphlus; characters; Greece).
▴ Distribution: Greece.
strinatii Coiffait, 1957: 67 (Allotyphlus; Type locality: Thrace grecque: Mesti).
— Coiffait, 1959: 313 (Allotyphlus; characters; Greece).
— Coiffait, 1972: 453 (Allotyphlus; characters; Greece).
▴ Distribution: Greece.
transhadriaticus Pace, 1977: 171 (Allotyphlus; Type locality: Promontorio del Gargano, Puglia, a circa 350 metri d’altitudine, in una valle presso la strada provinciale che scende a Vico Garganico, in localita Bosco Manatecco).
— Pace, 1985: 62 (Allotyphlus; characters).
— Zanetti, 1995: 14 (Allotyphlus; Italy).
— Pace, 1996: 128 (Allotyphlus; subgenus Allotyphlus; characters; Italy).
▴ Distribution: Italy.
turcicus Coiffait, 1957: 67 (Allotyphlus; Type locality: Turquie d’Asie: Celtiçi, à 30 km au sud du lac de Burdur).
Cyrtotyphlus
[4 species; Palaearctic Region]
Cyrtotyphlus Dodero, 1900: 401 (species included: convexus). Type species: Cyrtotyphlus convexus Dodero, fixed by monotypy.
— Heyden, Reitter, and Weise, 1906: 145 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1911: 147 (world catalog; 1 species).
— Winkler, 1925: 348 (catalog for Palaearctic region).
— Porta, 1926: 52 (characters; species of Italy).
— Scheerpeltz, 1933: 1138 (world catalog supplement).
— Blackwelder, 1952: 116 (type species: convexus).
— Scheerpeltz, 1955b: 463 (characters in key; type species: convexus; list of species).
— Coiffait, 1957: 61, 62 (characters).
— Coiffait, 1959: 276 (characters; key to species; type species: convexus).
— Horion, 1963: 296 (list of central European species).
— Coiffait, 1972: 386 (characters; type species: convexus; key to species of western Palaearctic region; references).
— Pace, 1976b: 53 (characters).
— Pace, 1977: 196 (key to species of Apennines).
— Zanetti, 1995: 14 (list of species of Italy).
— Pace, 1996: 75 (characters; key to species of Italy).
apulus Pace, 1975, see: Neocyrtotyphlus.
bonadonai Jarrige, 1949, see: convexus Dodero, 1900
convexus Dodero, 1900: 402 (Cyrtotyphlus; Type locality: Ruta, Liguria orient.).
— Bernhauer and Schubert, 1911: 147 (Cyrtotyphlus; catalog).
— Scheerpeltz, 1933: 1138 (Cyrtotyphlus; catalog).
— Coiffait, 1959: 278 (Cyrtotyphlus; characters; France).
— Horion, 1963: 296 (Cyrtotyphlus; Europe).
— Coiffait, 1972: 389 (Cyrtotyphlus; characters; Italy).
— Pace, 1977: 196 (Cyrtotyphlus; characters; Italy).
— Zanetti, 1995: 14 (Cyrtotyphlus; Italy).
— Pace, 1996: 78 (Cyrtotyphlus; characters; Italy).
▴ Distribution: France, Italy.
bonadonai Jarrige, 1949: 63 (Cyrtotyphlus; Type locality: Pont des Veyans, Vallée de la Siagne, Alpes-Mar.).
— Scheerpeltz, 1955b: 463 (Cyrtotyphlus).
— Coiffait, 1972: 389 (Cyrtotyphlus; synonym of convexus).
— Pace, 1996: 78 (Cyrtotyphlus; synonym of convexus).
kaiseri Pace, 1983a: 451 (Cyrtotyphlus; Type locality: Begovina bei Castelnuovo, Süd-Dalmatien).
thracicus Coiffait, 1957: 62 (Cyrtotyphlus; Type locality: Trace grecque: Sapé).
— Coiffait, 1959: 278 (Cyrtotyphlus; characters; Greece). — Coiffait, 1972: 389 (Cyrtotyphlus; characters; Greece).
▴ Distribution: Greece.
winkleri Breit, 1913: 235 (Cyrtotyphlus; Type locality: Nanos in Krain).
— Scheerpeltz, 1933: 1138 (Cyrtotyphlus; variety of convexus).
— Scheerpeltz, 1955b: 463 (Cyrtotyphlus).
— Coiffait, 1959: 278 (Cyrtotyphlus; subspecies of convexus; characters; Carniola).
— Coiffait, 1962d: 159 (Cyrotyphlus; valid species; aedeagus; notes; Slovenia).
— Coiffait, 1972: 389 (Cyrtotyphlus; characters; Carniola; Italy; Istria).
— Pace, 1975b: 9 (Cyrtotyphlus; characters).
— Pace, 1976b: 48, 49 (Cyrtotyphlus; characters; discussion of distribution).
— Zanetti, 1995: 14 (Cyrtotyphlus; Italy).
— Pace, 1996: 80 (Cyrtotyphlus; characters; Italy).
▴ Distribution: Italy, Slovenia.
Entomoculia
[121 species; Palaearctic Region]
Entomoculia Croissandeau, 1891: 150 (species included: sublaevis, grouvellei; [Note: According to Coiffait, 1959: 279 and 1972: 390 both species included in Entomoculia by Croissandeau were misidentified]). Type species: Entomoculia jeanneli Coiffait (= E. grouvellei sensu Croissandeau), fixed by application of Article 70.3.2 (see, 2001).
— Bernhauer and Schubert, 1911: 146 (synonym of Leptotyphlus).
— Normand, 1939: 490 (key to Tunisian species).
— Coiffait, 1955a: 66 (characters; type species jeanneli).
— Scheerpeltz, 1955b: 463 (characters in key; type species: grouvellei Fauvel; list of species).
— Coiffait, 1957: 61, 62 (characters; key to subgenera and species).
— Coiffait, 1959: 278 (characters; key to species; type species: grouvellei Croissandeau).
— Hervé, 1971b (notes on interpretation of aedeagus).
— Coiffait, 1972: 390 (characters; type species: jeanneli [= grouvellei]; key to species of western Palaearctic region; references).
— Pace, 1977: 197 (key to species of Apennines).
— Thelot, 1983: 311 (key to species from limestone region of Var, France).
— Orousset, 1984: 300 (list of species of Corsica).
— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).
— Zanetti, 1995: 14 (list of species of Italy).
— Pace, 1996: 80 (characters; key to species of Italy).
— Herman, 2001: 5 (type species; discussion; fixed by application of article 70.3).
Stenotyphlus Coiffait, 1955a: 67 (subgenus of Entomoculia; species included: bonadonai, tanneronensis, siagnensis, affinis, esterelensis, berberus, kabylianus). Type species: Entomoculia (Stenotyphlus) bonadonai Coiffait, fixed by original designation.
— Coiffait, 1957: 63, 65 (subgenus of Entomoculia; characters; key to species).
— Coiffait, 1959: 279, 298 (subgenus of Entomoculia; type species: bonadonai Coiffait; key to species).
— Coiffait, 1972: 390, 393, 421 (subgenus of Entomoculia; type species: bonadonai; characters; references).
— Pace, 1996: 121 (subgenus of Entomoculia; key to species of Italy).
Parentomoculia Scheerpeltz, 1955b: 464 (species included: argus, veneriata, angusticeps, capito, cyclops, laetifica, siccana, jupiter, dapsilis, exsculpta, mendica, orion, perpensa, pauxilla, saturnius, simplaria). Type species: Leptotyphlus argus Normand, fixed by original designation.
— Coiffait, 1959: 278 (synomym of Entomoculia; type species: argus Normand).
— Coiffait, 1972: 390 (synonym of Entomoculia).
aberrans Coiffait, 1962d: 168 (Entomoculia; subgenus Stenotyphlus; Type locality: Espagne: Province de Gerona, Palafrugel, Tamariu).
— Coiffait, 1972: 426, 435 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
adjacia Coiffait, 1957: 65 (Entomoculia; subgenus Entomoculia; Type locality: Corse: Ajaccio).
— Coiffait, 1959: 287, 294 (Entomoculia; characters; Corsica).
— Coiffait, 1972: 408, 419 (Entomoculia; characters; Corsica).
— Pace, 1996: 103 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica. affinis Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Cannes).
— Coiffait, 1959: 300, 304 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 426, 435 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
alticola Coiffait, 1959: 396 (Entomoculia; subgenus Entomoculia; Type locality: Alpes-Maritimes: vers 1000 mètres à Coursegoules).
ampurdana Coiffait, 1959: 290, 293 (Entomoculia; subgenus Entomoculia; Type locality: Prov. de Gerona, environs de Figueras).
angulata Coiffait, 1959: 400 (Entomoculia; subgenus Stenotyphlus; Type locality: Alpes-Maritimes: la basse vallée de l’Esteron à Gilette).
— Coiffait, 1972: 426, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
angusticeps Normand, 1939: 499 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Le Kef).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 308 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
— Coiffait, 1972: 438 (Entomoculia; subgenus Stenotyphlus; characters).
▴ Distribution: Tunisia.
antheorensis Coiffait, 1955a: 67 (Entomoculia; Type locality: Var: Antheor).
— Coiffait, 1959: 288 (Entomoculia; characters; France).
— Coiffait, 1972: 408 (Entomoculia; characters; France).
— Hervé, 1977: 168 (Entomoculia; characters).
▴ Distribution: France.
cavalairensis Coiffait, 1959: 398 (Entomoculia; subgenus Entomoculia; Type locality: Var: Cavalaire).
— Coiffait, 1972: 410, 415 (Entomoculia; characters; France).
— Hervé, 1977: 169, 176 (Entomoculia; race of antheorensis; characters).
opulenta Coiffait, 1962d: 162 (Entomoculia; subgenus Entomoculia; Type locality: Var: Forêt de Pierrefeu, Les Montauts).
— Coiffait, 1972: 410, 415 (Entomoculia; characters; France).
— Hervé, 1977: 170, 176 (Entomoculia; race of antheorensis; characters).
aprutiana Pace, 1977, see: montisconeri Pace, 1977.
apuana Pace, 1977: 166 (Entomoculia; Type locality: Alpi Apuane a Stazzema).
— Poggi, 1985: 176 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 104 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
arcsensis Coiffait, 1959: 396 (Entomoculia; subgenus Entomoculia; Type locality: Var: Vidauban).
— Coiffait, 1972: 405, 416 (Entomoculia; characters; France).
— Thelot, 1983: 304 (Entomoculia; characters; France).
▴ Distribution: France.
argensis Coiffait, 1959: 394 (Entomoculia; subgenus Entomoculia; Type locality: Var: Le Muy).
— Coiffait, 1972: 402, 416 (Entomoculia; characters; France).
— Thelot, 1983: 305 (Entomoculia; synonym of arcsensis).
argensis Coiffait, 1959, see: arcsensis Coiffait, 1959.
argus Normand, 1939: 491 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisia: Aïn-Draham).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 282 (Entomoculia; characters; Tunisia).
— Coiffait, 1972: 396, 421 (Entomoculia; characters; Tunisia).
▴ Distribution: Tunisia.
aspensis Coiffait, 1957: 64 (Entomoculia; subgenus Entomoculia; Type locality: Pyr. Orientales: Bouleternère).
— Coiffait, 1959: 286, 293 (Entomoculia; cited as asprensis; characters; France).
— Coiffait, 1972: 402, 417 (Entomoculia; cited as asprensis; characters; Pyrenees).
▴ Distribution: France.
atoma Saulcy, 1878: 118 (Mayetia; Type locality: Sardaigne, près de Bonnari).
— Coiffait, 1959: 306 (Entomoculia; subgenus Stenotyphlus; characters; Sardinia).
— Coiffait, 1972: 436 (Entomoculia; subgenus Stenotyphlus; characters).
— Pace, 1996: 124 (Entomoculia; subgenus Stenotyphlus; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
▴ Distribution: Italy.
barcelonensis Coiffait, 1962d: 165 (Entomoculia; subgenus Stenotyphlus; Type locality: Espagne: Barcelone, colline de Tibidabo, versant nord, près du sommet).
— Coiffait, 1972: 432, 436 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
barensis Coiffait, 1972: 430, 434, 447 (Entomoculia; subgenus Stenotyphlus; Type locality: Alpes Maritimes: Le Bar-sur-Loup).
berbera Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Algérie: Yakouren).
— Coiffait, 1959: 300, 308 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
— Coiffait, 1972: 426, 437 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
bonadonai Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Cannes, Le Cannet, Valbonne).
— Coiffait, 1959: 302, 304 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 430, 435 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
bonifacia Coiffait, 1959: 398 (Entomoculia; subgenus Entomoculia; Type locality: Corse: Santa Manza près de Bonifacio).
— Coiffait, 1972: 409, 419 (Entomoculia; characters; Corsica).
— Pace, 1996: 101 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica.
brignolensis Coiffait, 1959: 400 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: le versant Nord de la montagne de la Loube à Brignoles).
— Coiffait, 1972: 432, 435 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
canariensis Outerelo, 1982: 323 (Entomoculia; subgenus Stenotyphlus; replacement name for viti Outerelo).
viti Outerelo, 1980: 73 (Entomoculia; subgenus Stenotyphlus; [preoccupied]; Type locality: Islas Canarias: isla de La Gomera, El Cedro, 600 m).
capito Normand, 1939: 501 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Le Kef).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 300, 310 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
— Coiffait, 1972: 422, 439 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
caprai Poggi, 1977a: 154 (Entomoculia; subgenus Stenotyphlus; Type locality: Ficuzza, Sicilia, prov. Palermo).
— Pace, 1996: 123 (Entomoculia; subgenus Stenotyphlus; characters; Italy).
— Sparacio, 1995: 160 (Entomoculia; characters; notes; Sicily).
— Zanetti, 1995: 14 (Entomoculia; Italy).
▴ Distribution: Italy.
catalonica Coiffait, 1955a: 67 (Entomoculia; Type locality: Prov. de Gerona: San Pedro Pescador).
— Coiffait, 1959: 288, 293 (Entomoculia; characters; Spain).
— Coiffait, 1972: 413, 417 (Entomoculia; characters; Spain).
▴ Distribution: Spain.
cavalairensis Coiffait, 1959, see: antheorensis Coiffait, 1955.
cerberus Coiffait, 1962d: 167 (Entomoculia; subgenus Stenotyphlus; Type locality: Pyrénées-Orientale: Cerbère, route de Banyuls, près d’une petite source).
— Coiffait, 1972: 428, 435 (Entomoculia; subgenus Stenotyphlus; characters; Pyrenees).
▴ Distribution: France.
coiffaiti Hervé, 1967b: 328 (Entomoculia; subgenus Stenotyphlus; Type locality: Hyères, Var, au pied nord du col du Serre, à l’hubac du Mont des Oiseaux).
— Coiffait, 1972: 430, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
maurettensis Hervé, 1968: 77 (Entomoculia; subgenus Stenotyphlus; subspecies of coiffaiti; Type locality: ravin à l’hubac du Fenouillet, massif des Maurettes, commune de La Crau, Var).
colasi Hervé, 1969b: 74 (Entomoculia; subgenus Stenotyphlus; Type locality: 2 km au Nord-Est du village de Cuers, Var).
— Hervé, 1971b (Entomoculia; notes on interpretation of aedeagus).
— Coiffait, 1972: 430, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
comellinii Coiffait, 1965: 137 (Entomoculia; Type locality: Espagne: Catalogne, Province de Barcelone, Vallirana).
— Coiffait, 1972: 398, 418 (Entomoculia; cited as commellinii; characters; Spain).
▴ Distribution: Spain.
confusa Coiffait, 1959: 294 (Entomoculia; subgenus Entomoculia; Type locality: Corse: Porto Vecchio).
— Coiffait, 1972: 418 (Entomoculia; characters).
— Pace, 1996: 104 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica.
consiglioi Pace, 1991: 107 (Entomoculia; Type locality: Italia: Lazio: Isola di Palmarola, Fosso delle Radiche).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 112 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
convexa Coiffait, 1959: 292 (Entomoculia; subgenus Entomoculia; Type locality: Var: Lentier).
corbarensis Coiffait, 1959: 397 (Entomoculia; subgenus Entomoculia; Type locality: Corse: Pietracorbara sur la côte Est du Cap Corse).
— Coiffait, 1972: 405, 419 (Entomoculia; characters; Corsica).
— Pace, 1996: 119 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica.
cyclops Normand, 1939: 501 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Le Kef).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 282 (Entomoculia; characters; Tunisia).
— Coiffait, 1972: 396 (Entomoculia; characters; Tunisia).
▴ Distribution: Tunisia.
dapsilis Normand, 1950: 105 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Bizerte).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1972: 450 (Entomoculia).
▴ Distribution: Tunisia.
dentata Coiffait, 1959: 397 (Entomoculia; subgenus Entomoculia; Type locality: au pied du Monte Minervio à Pino, Corse).
— Coiffait, 1972: 405, 419 (Entomoculia; characters; Corsica).
— Pace, 1996: 118 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica.
dispersa Coiffait, 1955a: 67 (Entomoculia; Type locality: Drôme: Nyons).
— Coiffait, 1959: 286, 291 (Entomoculia; characters; France).
— Coiffait, 1972: 399, 416 (Entomoculia; characters; France).
— Thelot, 1983: 302 (Entomoculia; characters; France).
▴ Distribution: France.
lorguensis Coiffait, 1959: 396 (Entomoculia; subgenus Entomoculia; Type locality: Var: Lorgues).
— Coiffait, 1972: 404, 417 (Entomoculia; characters; France).
— Thelot, 1983: 302 (Entomoculia; synonym of dispersa).
dissimilis Coiffait, 1959: 310 (Entomoculia; subgenus Stenotyphlus; Type locality: Tunisie: Le Kef).
doderoi Coiffait, 1955a: 66 (Entomoculia; Type locality: Emilie: Finale).
— Coiffait, 1959: 287 (Entomoculia; characters; Italy).
— Coiffait, 1972: 405, 415 (Entomoculia; characters; Italy).
— Pace, 1977: 197 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 93 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
domensis Coiffait, 1962, see: henryi Coiffait, 1959.
ebusica Coiffait, 1973b: 219 (Entomoculia; Type locality: Iles Baléares, Ibiza, Cala San Vicente).
— Pons and Palmer, 1996: 163 (Entomoculia; Balearic endemic; notes; habitat; Eivissa).
▴ Distribution: Balearic Islands.
enigmatica Coiffait, 1962d: 161 (Entomoculia; subgenus Entomoculia; Type locality: Pyrénées-Orientales: Valleberme).
espanoli Coiffait, 1955a: 67 (Entomoculia; Type locality: Barcelone).
— Coiffait, 1959: 284, 293 (Entomoculia; characters; Spain).
— Coiffait, 1972: 398, 418 (Entomoculia; characters; Spain).
▴ Distribution: Spain.
esterelensis Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: St-Jean-de-l’Estérel).
— Coiffait, 1959: 301 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 428 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
etrurica Coiffait, 1957: 65 (Entomoculia; subgenus Entomoculia; Type locality: Italie: Environs de Grosseto).
— Coiffait, 1959: 288 (Entomoculia; characters; Italy).
— Coiffait, 1972: 410 (Entomoculia; characters; Italy).
— Pace, 1977: 197 (Entomoculia; characters; Italy).
— Poggi, 1985: 167 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 97 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
exsculpta Normand, 1939: 502 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: La Calle).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 284, 294 (Entomoculia; characters; Algeria).
— Coiffait, 1972: 396, 420 (Entomoculia; characters; Algeria).
▴ Distribution: Algeria.
gardinii Pace, 1994: 235 (Entomoculia; Type locality: Sardegna: dintorni di Cuglieri, Oristano).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 318 (Entomoculia; subgenus Entomoculia; characters; Sardinia).
▴ Distribution: Sardinia.
gavarrensis Coiffait, 1962d: 161 (Entomoculia; subgenus Entomoculia; Type locality: Espagne: Province de Gerona, Santa Pelaya).
grouvellei sensu Croissandeau, 1891 [Note: Nomen nudum; misidentification], see: jeanneli Coiffait, 1955.
guixolensis Coiffait, 1962d: 165 (Entomoculia; subgenus Stenotyphlus; Type locality: Espagne: Province de Gerona, San Feliu de Guixols).
— Coiffait, 1972: 422, 436 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
hadriatica Pace, 1975: 140 (Entomoculia; Type locality: circa km 2 di distanza, ad est, dal centro abitato di Genga, provincia di Ancona, nel versante adriatico dell’Appennino Marchigiano, a circa 300 metri d’altitudine, presso un torrentello affluente di sinistra del torrente Sentino).
— Pace, 1977: 197 (Entomoculia; characters; Italy).
— Poggi, 1985: 176 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 109 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
henryi Coiffait, 1959: 398 (Entomoculia; subgenus Entomoculia; Type locality: Var: Port Cros).
— Coiffait, 1972: 408l, 415 (Entomoculia; characters; France).
— Hervé, 1977: 171 (Entomoculia; characters; France).
▴ Distribution: France.
domensis Coiffait, 1962d: 164 (Entomoculia; subgenus Entomoculia; Type locality: Var: Le Dom des Baumes).
— Coiffait, 1972: 410 (Entomoculia; characters; France).
— Hervé, 1977: 171, 172 (Entomoculia; synonym of henryi; characters).
hervei Coiffait, 1957: 64 (Entomoculia; subgenus Entomoculia; Type locality: Var: Dardenne près de Toulon).
— Coiffait, 1959: 286, 292 (Entomoculia; characters; France).
— Coiffait, 1972: 402, 415 (Entomoculia; characters; France).
— Hervé, 1974: 55 (Entomoculia; characters; microscopic preparation of aedeagus).
— Hervé, 1976 (Entomoculia; aedeagal structure).
— Thelot, 1983: 300 (Entomoculia; characters; France).
▴ Distribution: France.
hoffmanni Coiffait, 1957: 66 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Tanneron, N.D. de Peygros).
— Coiffait, 1959: 301, 304 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Hervé, 1968a: 198 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 430, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
iknusa Coiffait, 1959: 294 (Entomoculia; subgenus Entomoculia; Type locality: Sardaigne: San Sperate près de Cagliari).
— Coiffait, 1972: 419 (Entomoculia; characters).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 117 (Entomoculia; subgenus Entomoculia; characters; Sardinia).
▴ Distribution: Sardinia.
inferna Coiffait, 1959: 304 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Fréjus).
jeanneli Coiffait, 1955a: 66 (Entomoculia; proposed for the misidentification of grouvellei by Croissandeau; Type locality: Environs de Nice).
— Coiffait, 1959: 286, 292 (Entomoculia; characters; France).
— Coiffait, 1972: 405, 416 (Entomoculia; characters; France).
▴ Distribution: France.
grouvellei sensu Croissandeau, 1891: 151 (Entomoculia; [Note: Nomen nudum, misidentification (Article 49); Croissandeau attributed the species to Fauvel but according to Coiffait, 1955a: 66, 1959: 279, and 1972: 390 the species was misidentified]; locality cited: Nice).
jeannetensis Coiffait, 1959: 400 (Entomoculia; subgenus Stenotyphlus; Type locality: Alpes-Maritimes: St-Jeannet).
— Coiffait, 1972: 426, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
julii Poggi, 1985: 168 (Entomoculia; Type locality: Isola di Montecristo, Arcipelago toscano, prov. Livorno, Valle dei Lecci, 150 m).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 95 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
jupiter Normand, 1939: 497 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Fernana).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 308 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
— Coiffait, 1972: 438 (Entomoculia; subgenus Stenotyphlus; characters).
▴ Distribution: Tunisia.
kabyliana Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Algérie: Adekar).
— Coiffait, 1959: 301 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
— Coiffait, 1972: 428 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
keltiensis Coiffait, 1970b: 146 (Entomoculia; subgenus Stenotyphlus; Type locality: Souk el Arba des Beni Hassan, Rif occidental, Maroc).
— Coiffait, 1972: 428 (Entomoculia; subgenus Stenotyphlus; characters; Morocco).
— Coiffait, 1973a: 274 (Entomoculia; Morocco).
▴ Distribution: Morocco.
labronica Poggi, 1985: 179 (Entomoculia; Type locality: Dintorni di Livorno, Toscana).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 107 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
laetifica Normand, 1939: 499 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Le Kef).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 310 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
— Coiffait, 1972: 439 (Entomoculia; subgenus Stenotyphlus; characters).
▴ Distribution: Tunisia.
laneyriei Coiffait, 1955a: 66 (Entomoculia; Type locality: Alpes Maritimes: Vallée de l’Esteron).
— Coiffait, 1959: 284, 292 (Entomoculia; characters; France).
— Coiffait, 1972: 399, 416 (Entomoculia; characters; Maritime Alpes).
▴ Distribution: France.
lauricola Outerelo and Hernández, 1989: 515 (Entomoculia; subgenus Stenotyphlus; Type locality: Islas Canarias: Tenerife: Vueltas de Taganana).
lepinensis Pace, 1991: 107 (Entomoculia; Type locality: Italia: Lazio, Monti Lepini, Norma, zona boschiva a Km 2 a Nord delle rovine dell’antica Norba).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 114 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
ligurica Coiffait, 1957: 65 (Entomoculia; subgenus Entomoculia; Type locality: Environs de Gênes, Belvédère).
— Coiffait, 1959: 288 (Entomoculia; characters; Italy).
— Coiffait, 1972: 412 (Entomoculia; characters; Italy).
— Pace, 1977: 197 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 91 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
lobata Coiffait, 1959: 397 (Entomoculia; subgenus Entomoculia; Type locality: Corse: Ile Rousse).
— Coiffait, 1972: 405, 419 (Entomoculia; characters; Corsica).
— Pace, 1996: 120 (Entomoculia; subgenus Entomoculia; characters; Corsica).
▴ Distribution: Corsica.
lorguensis Coiffait, 1959, see: dispersa Coiffait, 1955.
lucensis Coiffait, 1962d: 162 (Entomoculia; subgenus Entomoculia; Type locality: Var, le Luc).
— Coiffait, 1972: 402 (Entomoculia; characters; France).
— Hervé, 1976a (Entomoculia; aedeagal characters).
— Thelot, 1983: 307 (Entomoculia; characters; France).
▴ Distribution: France.
rougoni Coiffait, 1971a: 175 (Entomoculia; Type locality: France: Bois de Cadarache, Bouches-du-Rhône).
— Coiffait, 1972: 400 (Entomoculia; characters; France).
— Hervé, 1976a (Entomoculia; aedeagal characters).
— Thelot, 1983: 307 (Entomoculia; synonym of lucensis).
magdalenae Coiffait, 1957: 66 (Entomoculia; subgenus Stenotyphlus; Type locality: Levante espagnol, Province de Casteillon: Santa Magdalena).
— Coiffait, 1959: 300 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
— Coiffait, 1972: 423 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
magrinii Pace, 1994: 232 (Entomoculia; Type locality: Italia: Toscana, Monte Pisano, S. Giuliano Terme, Pisa).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 317 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
malierensis Hervé, 1968a: 199 (Entomoculia; subgenus Stenotyphlus; Type locality: ravin secondaire du vallon de La Malière à Collobrières, dans les Maures, Var).
— Coiffait, 1972: 423, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
ultima Hervé, 1969: 253 (Entomoculia; subgenus Stenotyphlus; subspecies of malierensis; Type locality: colline de Peigros, commune de Pierrefeu, Var).
mancinii Briganti, 1979, see: toscanensis Coiffait, 1957.
maremmana Pace, 1979: 172 (Entomoculia; Type locality: Monti dell’Uccellina, Poggio Lecci, presso S. Rabano, 250 m).
— Poggi, 1985: 176 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 109 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
marinae Pace, 1991: 107 (Entomoculia; Type locality: Italia: Lazio: Isola di Zannone, dolina del Monte Pellegrino).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 114 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
maritima Coiffait, 1959: 292 (Entomoculia; subgenus Entomoculia; Type locality: Alpes-Maritimes: Mandelieu).
mars Coiffait, 1959: 296 (Entomoculia; subgenus Entomoculia; Type locality: Tunisie: Bordj Messaoudi).
maurettensis Hervé, 1968, see: coiffaiti Hervé, 1967.
mayeti Normand, 1907: 122 (Leptotyphlus; Type locality: Port-Vendres, près du fort Bear).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; synonym of grouvellei).
— Portevin, 1929: 392 (Leptotyphlus; characters; France).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; synonym of grouvellei).
— Scheerpeltz, 1955b: 463 (Entomoculia).
— Coiffait, 1959: 293 (Entomoculia; characters; France).
— Coiffait, 1972: 417 (Entomoculia; characters).
▴ Distribution: France.
melendoi Outerelo, 1980a: 57, 62 (Entomoculia; subgenus Stenotyphlus; Type locality: Arroyo Frio, 950 m, la Sierra de Cazorla).
mendica Normand, 1939: 494 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Philippeville).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 294 (Entomoculia; characters; Algeria).
— Coiffait, 1972: 419 (Entomoculia; characters; Algeria).
▴ Distribution: Algeria.
meridionalis Coiffait, 1959: 296 (Entomoculia; subgenus Entomoculia; Type locality: Tunisie: Sousse).
montisconeri Pace, 1977: 156 (Entomoculia; Type locality: Marche sul Monte Conero, a sud della città di Ancona, intorno i 400 metri d’altitudine prevalentemente sulle pendici meridionali, al di sotto della Badia di S. Pietro).
— Poggi, 1985: 173 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 98 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
aprutiana Pace, 1977: 163 (Entomoculia; Type locality: Abruzzo, in um bosco presso la riva destra del fiume Sangro, a circa 500 metri dalla sua foce, nel territorio del comune di Torino di Sangro, a circa 8 metri d’altitudine).
— Poggi, 1985: 173 (Entomoculia; synonym of montisconeri).
— Pace, 1996: 98 (Entomoculia; synonym of montisconeri).
nerviensis Coiffait, 1962d: 164 (Entomoculia; subgenus Entomoculia; Type locality: Alpes-Maritimes italiennes, Province d’Imperia, Isolabona, vallée de la Nervia, au nord de Vintimille).
— Coiffait, 1972: 411 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 90 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
normandi Coiffait, 1955a: 66 (Entomoculia; Type locality: Tunisie: Le Kef).
— Coiffait, 1959: 282 (Entomoculia; characters; Tunisia).
— Coiffait, 1972: 395 (Entomoculia; characters; Tunisia).
▴ Distribution: Tunisia.
numidica Coiffait, 1959: 308 (Entomoculia; subgenus Stenotyphlus; Type locality: Algérie: Miliana).
— Coiffait, 1972: 437 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
occidentalis Coiffait, 1957: 64 (Entomoculia; subgenus Entomoculia; Type locality: Pyr. Orientales: Maury).
— Coiffait, 1959: 285, 293 (Entomoculia; characters; France).
— Coiffait, 1972: 399, 417 (Entomoculia; characters; Pyrenees).
▴ Distribution: France.
opulenta Coiffait, 1962, see: antheorensis Coiffait, 1955.
opulentissima Coiffait, 1962d: 164 (Entomoculia; subgenus Entomoculia; Type locality: Var: La Garde-Freinet).
— Coiffait, 1972: 410, 415 (Entomoculia; characters; France).
— Hervé, 1977: 172, 173 (Entomoculia; characters; France).
▴ Distribution: France.
orion Normand, 1939: 492 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Philippeville).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1972: 449 (Entomoculia).
▴ Distribution: Algeria.
orpheus Coiffait, 1959: 304 (Entomoculia; subgenus Stenotyphlus; Type locality: Alpes-Maritimes: Les Courmettes).
— Coiffait, 1972: 435 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
pardii Pace, 1979: 183 (Entomoculia; Type locality: Monte Calvana presso Firenze).
— Poggi, 1985: 178 (Entomoculia; notes; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 105 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
pauxilla Normand, 1939: 500 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Tlemcen).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 307 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
— Coiffait, 1972: 437 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
perpensa Normand, 1939: 495 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Philippeville).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 296 (Entomoculia; characters; Algeria).
— Coiffait, 1972: 420 (Entomoculia; characters; Algeria).
▴ Distribution: Algeria.
pretiosa Pace, 1977: 168 (Entomoculia; subgenus Stenotyphlus; Type locality: Marche sul Monte Conero, presso Ancona a circa 450 metri d’altitudine).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 122 (Entomoculia; subgenus Stenotyphlus; characters; Italy).
▴ Distribution: Italy.
prodigiosa Coiffait, 1962d: 168 (Entomoculia; subgenus Stenotyphlus; Type locality: Espagne: Province de Gerona, Bagur, Fornells, au pied d’oliviers).
— Coiffait, 1972: 426, 436 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
quartensis Coiffait, 1965: 138 (Entomoculia; Type locality: Espagne: Catalogne, Prov. de Gerona, Quart).
robusta Coiffait, 1959: 306 (Entomoculia; subgenus Stenotyphlus; Type locality: Algérie: Bugeaud).
— Coiffait, 1972: 437 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
rosai Pace, 1978: 38 (Entomoculia; Type locality: Isola di Capraia, Il Piano).
— Poggi, 1985: 173 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 108 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
rougoni Coiffait, 1971, see: lucensis Coiffait, 1962.
sardoa Coiffait, 1959: 294 (Entomoculia; subgenus Entomoculia; Type locality: Sardaigne: Ozieri).
— Coiffait, 1972: 419 (Entomoculia; characters).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 117 (Entomoculia; subgenus Entomoculia; characters; Sardinia).
▴ Distribution: Sardinia.
sassariensis Pace, 1978b: 191 (Entomoculia; Type locality: dintorni di Sassari).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 116 (Entomoculia; subgenus Entomoculia; characters; Sardinia).
▴ Distribution: Sardinia.
satanus Coiffait, 1959: 296 (Entomoculia; subgenus Entomoculia; Type locality: Algérie: Philippeville).
saturnia Normand, 1939: 493 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Bône).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 308 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
— Coiffait, 1972: 437 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
senyensis Coiffait, 1957: 66 (Entomoculia; subgenus Stenotyphlus; Type locality: Catalogne: Figaro au pied de la Sierra de Montseny).
— Coiffait, 1959: 302, 304 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
— Coiffait, 1972: 432, 436 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
septentrionis Thelot, 1983: 308 (Entomoculia; Type locality: Montmeyan, Var, lieu-dit les Vinatiers).
serratensis Coiffait, 1957: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Catalogne: Monistrol au pied de la Sierra de Montserrat).
— Coiffait, 1959: 302, 305 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
— Coiffait, 1972: 432, 436 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
siagnensis Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Pont des Veyans).
— Coiffait, 1959: 302 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 430 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
sicana Pace, 1993a: 31 (Entomoculia; Type locality: Sicilia: Castelbuono, Pintorna).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 319 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
siccana Normand, 1939: 498 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisie: Le Kef).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 300, 310 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
— Coiffait, 1972: 426, 438 (Entomoculia; subgenus Stenotyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
simplaria Normand, 1939: 496 (Leptotyphlus; subgenus Entomoculia; Type locality: Algérie: Bône).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 287 (Entomoculia; characters; Algeria).
— Coiffait, 1972: 408 (Entomoculia; characters; Algeria).
▴ Distribution: Algeria.
solarii Coiffait, 1957: 65 (Entomoculia; subgenus Entomoculia; Type locality: Ligurie Occidentale: Laigueglia).
— Coiffait, 1959: 288, 290 (Entomoculia; characters; Italy).
— Coiffait, 1972: 410, 413 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 95 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
strangulata Coiffait, 1957: 64 (Entomoculia; subgenus Entomoculia; Type locality: Algérie: Bugeaud).
— Coiffait, 1959: 284 (Entomoculia; characters; Algeria).
— Coiffait, 1972: 396 (Entomoculia; characters; Algeria).
▴ Distribution: Algeria.
subhervei Coiffait, 1959: 394 (Entomoculia; subgenus Entomoculia; Type locality: Besse-sur-Issole, Var).
— Coiffait, 1972: 402, 416 (Entomoculia; characters; France).
— Thelot, 1983: 305 (Entomoculia; characters; France).
▴ Distribution: France.
subtilis Coiffait, 1959: 304 (Entomoculia; subgenus Stenotyphlus; Type locality: Español: Masnou, près de Barcelone).
— Coiffait, 1972: 435 (Entomoculia; subgenus Stenotyphlus; characters; Spain).
▴ Distribution: Spain.
sylvatica Coiffait, 1965: 137 (Entomoculia; Type locality: France: Var: Bagnols en Forêt).
tanneronensis Coiffait, 1955a: 67 (Entomoculia; subgenus Stenotyphlus; Type locality: Var: Tanneron).
— Coiffait, 1959: 301, 304 (Entomoculia; subgenus Stenotyphlus; characters; France).
— Coiffait, 1972: 428, 434 (Entomoculia; subgenus Stenotyphlus; characters; France).
▴ Distribution: France.
theloti Hervé, 1977: 174, 177 (Entomoculia; Type locality: France: Var: La Garde-Freinet).
thoracica Coiffait, 1959: 306 (Entomoculia; subgenus Stenotyphlus; Type locality: Bône).
— Coiffait, 1972: 436 (Entomoculia; subgenus Stenotyphlus; characters; Algeria).
▴ Distribution: Algeria.
toscanensis Coiffait, 1957: 65 (Entomoculia; subgenus Entomoculia; Type locality: Toscane: Monte Argentaro).
— Coiffait, 1959: 287, 290 (Entomoculia; characters; Italy).
— Coiffait, 1972: 408, 415 (Entomoculia; characters; Italy).
— Pace, 1977: 197 (Entomoculia; characters; Italy).
— Poggi, 1985: 165 (Entomoculia; characters; Italy).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 98 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
mancinii Briganti, 1979: 93 (Entomoculia; subgenus Entomoculia; Type locality: dondo di una dolina presso Pian Fagiano nell’Isola di Giannutri).
— Poggi, 1985: 165 (Entomoculia; synonym of toscanensis).
— Pace, 1996: 98 (Entomoculia; synonym of toscanensis).
tossaensis Coiffait, 1962d: 159 (Entomoculia; subgenus Entomoculia; Type locality: Espagne: province de Gerona, Tossa de Mar).
ultima Hervé, 1969, see: malierensis Hervé, 1968.
uranus Coiffait, 1959: 308 (Entomoculia; subgenus Stenotyphlus; Type locality: Tunisia).
veneriata Normand, 1939: 494 (Leptotyphlus; subgenus Entomoculia; Type locality: Tunisia: Camp de la Santé, près d’Aïn-Draham).
— Scheerpeltz, 1955b: 464 (Parentomoculia).
— Coiffait, 1959: 287, 296 (Entomoculia; characters; Tunisia).
— Coiffait, 1972: 406, 420 (Entomoculia; characters; Tunisia).
▴ Distribution: Tunisia.
vicina Coiffait, 1962d: 162 (Entomoculia; subgenus Entomoculia; Type locality: Espagne: Province de Gerona, Callonge, plage Saint-Georges).
vignai Pace, 1991: 107 (Entomoculia; Type locality: Italia: Lazio, Isola di Ponza, versante NO Monte Guardia).
— Zanetti, 1995: 14 (Entomoculia; Italy).
— Pace, 1996: 113 (Entomoculia; subgenus Entomoculia; characters; Italy).
▴ Distribution: Italy.
viti Coiffait, 1973b: 219 (Entomoculia; Type locality: Corse: Olmeto).
viti Outerelo, 1980, see: canariensis Outerelo, 1982.
zariquieyi Coiffait, 1955a: 66 (Entomoculia; Type locality: Prov. de Gérona: Cadaques).
— Coiffait, 1959: 284, 293 (Entomoculia; characters; Spain).
— Coiffait, 1972: 399, 418 (Entomoculia; characters; Spain).
▴ Distribution: Spain.
zegzeiensis Coiffait, 1970b: 145 (Entomoculia; subgenus Stenotyphlus; Type locality: Berkane, Rif oriental, Maroc).
Mesotyphlus
[29 species; Palaearctic Region]
Mesotyphlus Coiffait, 1957: 62, 68 (species included: grouvellei, regalis, solarii). Type species: Leptotyphlus grouvellei Fauvel, fixed by original designation.
— Coiffait, 1959: 313 (characters; key to species; type species: grouvellei).
— Coiffait, 1965b: 679 (characters; key to subgenera and species).
— Coiffait, 1972: 458 (characters; type species: grouvellei; key to species of western Palaearctic region; references).
— Pace, 1977: 197 (key to species of Apennines).
— Pace, 1979c: 104 (key to species of Italy).
— Orousset, 1984: 300 (list of species of Corsica).
— Zanetti, 1995: 15 (list of species of Italy).
— Pace, 1996: 138 (characters; key to species of Italy).
Teleotyphlus Coiffait, 1965b: 680, 682, 687 (subgenus of Mesotyphlus; species included: rivularis). Type species: Mesotyphlus (Teleotyphlus) rivularis Coiffait, fixed by monotypy.
— Coiffait, 1972: 458, 460 (subgenus of Mesotyphlus; [Note: Coiffait cited M. maritimus as the type species of Teleotyphlus but it was not an originally included species. In 1972: 463, 471 he included M. maritimus in the nominate subgenus, not in Teleotyphlus. Moreover, the type species of Teleotyphlus had already been fixed by monotypy]; characters; type species: maritimus; key to species of western Palaearctic region).
Acanthotyphlus Coiffait, 1972: 458, 461, 465 (cited as Acantotyphlus and Acanthotyphlus; subgenus of Mesotyphlus; [Note: This name was described earlier (Coiffait, 1965b: 680) but the name was unavailable because a type species was not designated (Article 13.3). Cited as Acanthotyphlus (pp. 458, 460, 465, 468, 471, 645) and Acantotyphlus (pp. 461); similar errors were made when the name was originally published (see Coiffait, 1965b). I have accepted Acanthotyphlus as the correct spelling]; species included: inversus, furcatus, albufeirensis, simplex, affinis, bordigheirensis, paeonius, campus, montanus, siliquus, silvensis, rupestris). Type species: Mesotyphlus (Acantotyphlus) simplex Coiffait, fixed by original designation.
— Coiffait, 1965b: 680, 683, 687 (cited as Acanthotyphlus and Acantotyphlus; subgenus of Mesotyphlus; [Note: In this article Acantotyphlus was first published with characters and species but no type species was designated so the name was unavailable (Article 13.3). Cited as Acantotyphlus (pp. 680, 683) and Acanthotyphlus (pp. 684, 687 and 688 where this spelling was used nine times)]; characters; key to species).
— Pace, 1979c: 105 (cited as Acanthotyphlus; subgenus of Mesotyphlus; key to species of Italy).
Dexiotyphlus Coiffait, 1972: 458, 461, 463 ([Note: This name was described earlier (Coiffait, 1965b: 680) but was an unavailable name because a type species was not designated (Article 13.3). The name was also used again before 1972 (Coiffait, 1970b: 148) but is available only if the characters for the species are considered to apply also to the subgeneric name]; subgenus of Mesotyphlus; species included: brevis, pervincus, vicinus). Type species: Mesotyphlus (Dexiotyphlus) pervincus Coiffait, fixed by original designation.
— Coiffait, 1965b: 680, 683, 687 (subgenus of Mesotyphlus; [Note: In this article the name Dexiotyphlus was first published with characters and species but no type species was designated so it is an unavailable name (Article 13.3)]; characters; key to species).
— Pace, 1979c: 105 (subgenus of Mesotyphlus; key to species of Italy).
— Pace, 1996: 140, 145 (cited as Dexyotyphlus; subgenus of Mesotyphlus; characters; key to species of Italy).
affinis Coiffait, 1965b: 683, 687 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Loulé).
— Coiffait, 1972: 466 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
albufeirensis Coiffait, 1970b: 148 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Albufeira, Algarve, Portugal).
— Coiffait, 1972: 466 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
alutaceus Dodero, 1908, see: grouvellei Fauvel, 1890.
andreinii Pace, 1979c: 100 (Mesotyphlus; subgenus Dexiotyphlus; Type locality: Toscana: Pergine, Arezzo).
— Pace, 1996: 150 (Mesotyphlus; subgenus Dexiotyphlus, cited as Dexyotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
▴ Distribution: Italy.
bipunctatus Coiffait, 1973, see: Allotyphlus.
bordigheirensis Coiffait, 1970b: 150 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Bordiheira, Algarve, Portugal).
— Coiffait, 1972: 467 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
bordonii Pace, 1979c: 102 (Mesotyphlus; subgenus Acanthotyphlus; Type locality: Toscana: Giuncarico, Grosseto).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
— Pace, 1996: 147 (Mesotyphlus; subgenus Dexiotyphlus, cited as Dexyotyphlus; characters; Italy).
▴ Distribution: Italy.
brevis Coiffait, 1970b: 148 (Mesotyphlus; subgenus Dexiotyphlus; Type locality: Bordigheira, Algarve, Portugal).
— Coiffait, 1972: 463 (Mesotyphlus; subgenus Dexiotyphlus; [Note: Cited in Teleotyphlus on page 471]; characters; Portugal).
▴ Distribution: Portugal.
calaber Pace, 1975: 147 (Mesotyphlus; subgenus Acanthotyphlus; Type locality: Calabria, nei pressi di Sambiase, provincia di Catanzaro, a 4 chilometri a nord del centro abitato, vicino al villaggio di Gabella, a circa 560 metri d’altitudine, in una valle presso il torrente Cantagalli).
— Pace, 1977: 197 (Mesotyphlus; characters; Italy).
— Pace, 1979c: 98 (Mesotyphlus; subgenus Dexiotyphlus; characters).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
— Pace, 1996: 151 (Mesotyphlus; subgenus Dexiotyphlus, cited as Dexyotyphlus; characters; Italy).
▴ Distribution: Italy.
campus Coiffait, 1965b: 685, 688 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Boliquième près de Albufeira).
— Coiffait, 1972: 467 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
corsicus Coiffait, 1959: 406 (Mesotyphlus; Type locality: Corse: au pied du Mont Minervio à Pino).
— Coiffait, 1972: 461 (Mesotyphlus; characters; Corsica).
— Pace, 1996: 145 (Mesotyphlus; subgenus Moreotyphlus; characters; Corsica).
▴ Distribution: Corsica.
etruscus Pace, 1979: 176 (Mesotyphlus; Type locality: Monti dell’Uccellina, Poggio Lecci, dintorni di S. Rabano, 200–300 m).
— Pace, 1979c: 95 (Mesotyphlus; subgenus Mesotyphlus; Italy).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
— Pace, 1996: 143 (Mesotyphlus; subgenus Moreotyphlus; characters; Italy).
▴ Distribution: Italy.
furcatus Coiffait, 1965b: 683, 688 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Porto de Lagos près de Portimao).
— Coiffait, 1972: 466 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
grossetensis Pace, 1979c: 98 (Mesotyphlus; subgenus Dexiotyphlus; Type locality: Toscana: Poggio Cavallol, Grosseto).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
— Pace, 1996: 149 (Mesotyphlus; subgenus Dexiotyphlus, cited as Dexyotyphlus; characters; Italy).
▴ Distribution: Italy.
grouvellei Fauvel, 1890: 356 (Leptotyphlus; Type locality: Not cited but presumably Nice).
— Ganglbauer, 1895: 607 (Leptotyphlus; characters; France).
— Bernhauer, 1908e: 332 (Leptotyphlus; characters).
— Dodero, 1908: 634, 637 (Leptotyphlus; characters).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Dodero, 1916: 341 (Leptotyphlus; Sicily).
— Dodero, 1918: 106 (Leptotyphlus; notes; Catalonia).
— Portevin, 1929: 392 (Leptotyphlus; characters; France).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1957: 68 (Mesotyphlus).
— Coiffait, 1959: 314 (Mesotyphlus; characters; France).
— Coiffait, 1972: 461 (Mesotyphlus; characters; France).
— Pace, 1996: 141 (Mesotyphlus; subgenus Mesotyphlus; characters; Italy).
▴ Distribution: France, Italy, Spain.
alutaceus Dodero, 1908: 637 (Leptotyphlus; variety of grouvellei; Type locality: Nizza Marittima).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; variety of grouvellei).
— Scheerpeltz, 1955b: 463 (Entomoculia).
inversus Coiffait, 1970b: 150 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Sta. Catharina, Algarve, Portugal).
— Coiffait, 1972: 466 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
lessiniensis Coiffait, 1973, see: Allotyphlus.
lusitanicus Coiffait, 1965b: 680, 685 (Mesotyphlus; subgenus Mesotyphlus; Type locality: Portugal: Estremadure, Mafra).
maritimus Coiffait, 1965b: 680, 685 (Mesotyphlus; subgenus Mesotyphlus; Type locality: Portugal: Estremadure, Amoreira).
montanus Coiffait, 1965b: 685, 688 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Querença, près de Loulè).
— Coiffait, 1972: 467 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
munitius Coiffait, 1965b: 680, 687 (Mesotyphlus; subgenus Mesotyphlus; Type locality: Portugal: Estremadure, Obidos).
paeonius Coiffait, 1965b: 683, 688 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Amoideras près de Loulé).
— Coiffait, 1972: 467 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
pervincus Coiffait, 1965b: 683, 687 (Mesotyphlus; subgenus Dexiotyphlus; Type locality: Portugal: Algarve, Estoï près de Faro).
— Coiffait, 1972: 465 (Mesotyphlus; subgenus Dexiotyphlus; characters; Portugal).
▴ Distribution: Portugal.
regalis Coiffait, 1955a: 68 (Paratyphlus; Type locality: Toulouse).
— Coiffait, 1957: 68 (Mesotyphlus).
— Coiffait, 1959: 314 (Mesotyphlus; characters; France).
— Coiffait, 1972: 462 (Mesotyphlus; characters; France).
▴ Distribution: France.
rivularis Coiffait, 1965b: 682, 687 (Mesotyphlus; subgenus Teleotyphlus; Type locality: Portugal: Algarve, Porto de Lagos, près de Portimao).
— Coiffait, 1972: 463 (Mesotyphlus; subgenus Teleotyphlus; characters; Portugal).
▴ Distribution: Portugal.
rupestris Coiffait, 1965b: 685, 687 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Vilharinos près de São Braz d’Alportel).
— Coiffait, 1972: 469 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
siliquus Coiffait, 1965b: 685, 687 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Portugal: Algarve, São Romao près de São Braz d’Alportel).
— Coiffait, 1972: 467 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
silvensis Coiffait, 1965b: 685, 688 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, Silves).
— Coiffait, 1972: 468 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
simplex Coiffait, 1965b: 683, 687 (Mesotyphlus; subgenus Acantotyphlus; Type locality: Portugal: Algarve, São Romao près de São Braz d’Alportel).
— Coiffait, 1972: 466 (Mesotyphlus; subgenus Acantotyphlus; characters; Portugal).
▴ Distribution: Portugal.
solarii Coiffait, 1957: 68 (Mesotyphlus; Type locality: Italie: environs de Gênes: Belvédère).
— Coiffait, 1959: 314 (Mesotyphlus; characters; Italy).
— Coiffait, 1972: 462 (Mesotyphlus; characters; Italy).
— Pace, 1977: 198 (Mesotyphlus; characters; Italy).
— Pace, 1979c: 96 (Mesotyphlus; subgenus Dexiotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Mesotyphlus; Italy).
— Pace, 1996: 149 (Mesotyphlus; subgenus Dexiotyphlus, cited as Dexyotyphlus; characters; Italy).
▴ Distribution: Italy.
temperei Coiffait, 1979b: 313 (Mesotyphlus; Type locality: Corse: Omessa).
vicinus Coiffait, 1965b: 683, 687 (Mesotyphlus; subgenus Dexiotyphlus; Type locality: Portugal: Algarve, Santa Barbara de Nexe, près de Loulé).
Neocyrtotyphlus
[1 species; Palaearctic Region]
Neocyrtotyphlus Pace, 1976b: 52 (subgenus of Cyrtotyphlus; species included: apulus). Type species: Cyrtotyphlus (Neocyrtotyphlus) apulus Pace, fixed by original designation.
apulus Pace, 1975: 137 (Cyrtotyphlus; Type locality: Promontorio del Gargano in Puglia, sul versante settentrionale del Monte Calena a circa 400 metri d’altitudine, ai margini della Foresta Umbra).
Paratyphlus
[28 species; Palaearctic Region]
Paratyphlus Blackwelder, 1952: 294 ([Note: This name was first described by Normand, 1939: 487 who included three species, kabylianus, doderoi, and hipponensis, but failed to designate a type species thereby making Paratyphlus unavailable. With Blackwelder's designation and reference to Normand's description the name became available with Blackwelder, 1952, as the author and date]; subgenus of Leptotyphlus). Type species: Leptotyphlus doderoi Normand, fixed by subsequent designation by Blackwelder, 1952: 294.
— Normand, 1939: 487 ([Note: This is the original description of Paratyphlus but without a type species designation by Normand Paratyphlus is unavailable (Article 13.3). It became available when Blackwelder, 1952: 294 fixed the type species]; subgenus of Leptotyphlus; characters; species included: kabylianus, doderoi, hipponensis).
— Coiffait, 1955a: 68 (valid genus).
— Scheerpeltz, 1955b: 460 (characters in key; type species: doderoi; list of species).
— Coiffait, 1957: 62, 68 (characters; key to subgenera).
— Coiffait, 1959: 314 (characters; key to species; type species: kabylianus, cited as kabilianus; [Note: The type species was already fixed by Blackwelder, 1952]).
— Coiffait, 1965b: 688 (notes; characters; key to species of Portugal).
— Coiffait, 1972: 473 (characters; type species: kabylianus [Note: The type species was already fixed by Blackwelder, 1952]; key to species of western Palaearctic region; references).
Ischnotyphlus Coiffait, 1957: 68 (subgenus of Paratyphlus; species included: cephalotes, serratensis, mateui). Type species: Paratyphlus mateui Coiffait, fixed by original designation.
— Coiffait, 1959: 316, 318 (subgenus of Paratyphlus; cited as a new subgenus; characters; key to species; type species: mateui).
— Coiffait, 1972: 473, 475, 481 (subgenus of Paratyphlus; characters; type species: mateui; key to species of western Palaearctic region; references).
algarvensis Coiffait, 1965b: 689 (Paratyphlus; subgenus Paratyphlus; Type locality: Portugal: Algarve, Almançil).
atlasicus Coiffait, 1970b: 146 (Paratyphlus; Type locality: Ouaoumana, au pied du Moyen Atlas, Maroc).
— Coiffait, 1972: 477 (Paratyphlus; characters; Morocco).
— Coiffait, 1973a: 274 (Paratyphlus; Morocco).
▴ Distribution: Morocco.
barcelonensis Coiffait, 1962d: 169 (Paratyphlus; subgenus Paratyphlus; Type locality: Espagne: Barcelone, coteau de Tibidabo, sur le versant nord, près du sommet).
berberus Coiffait, 1957: 69 (Paratyphlus; subgenus Paratyphlus; Type locality: Algérie: Yakouren).
— Coiffait, 1959: 318 (Paratyphlus; characters; Algeria).
— Coiffait, 1972: 479 (Paratyphlus; characters; Algeria).
▴ Distribution: Algeria.
besucheti Coiffait, 1970b: 148 (Paratyphlus; Type locality: San Morell, près Arta, Majorque).
— Coiffait, 1972: 479 (Paratyphlus; characters; Balearic Islands).
— Pons and Palmer, 1996: 160 (Paratyphlus; Balearic endemic; notes; habitat; Mallorca).
▴ Distribution: Balearic Islands.
boiteli Normand, 1950: 105 (Leptotyphlus; subgenus Paratyphlus; Type locality: Tunisie: Bizerte).
brazensis Coiffait, 1965b: 689 (Paratyphlus; subgenus Paratyphlus; Type locality: Portugal: Algarve, São Romao, près São Braz d’Alportel).
carvoeirensis Coiffait, 1970b: 146 (Paratyphlus; Type locality: Carvoeiro, Algarve, Portugal).
cephalotes Coiffait, 1957: 69 (Paratyphlus; subgenus Ischnotyphlus; Type locality: Algérie: Cherchel).
— Coiffait, 1959: 322 (Paratyphlus; subgenus Ischnotyphlus; characters; Algeria).
— Coiffait, 1972: 483 (Paratyphlus; subgenus Ischnotyphlus; characters; Algeria).
▴ Distribution: Algeria.
comarrugi Coiffait, 1959: 410 (Paratyphlus; subgenus Ischnotyphlus; Type locality: Comarruga, localité située au bord de la mer, près et au Sud de Vendrell, Province de Tarragone).
— Coiffait, 1972: 486 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
cristobali Coiffait, 1959: 406 (Paratyphlus; subgenus Paratyphlus; Type locality: Baléares: Ile de Minorque: prélèvement de terre fait sous la lucarne dans la Grotte Na Pulida, près de San Agusti au Sud de San Cristobal).
— Coiffait, 1972: 479 (Paratyphlus; characters; Balearic Islands).
— Pons and Palmer, 1996: 159 (Paratyphlus; Balearic endemic; notes; habitat; Menorca).
▴ Distribution: Balearic Islands.
delicatulus Coiffait, 1965b: 688, 689 (Paratyphlus; subgenus Paratyphlus; Type locality: Portugal: Algarve, São Romao, près São Braz d’Alportel).
doderoi Normand, 1910: 88 (Leptotyphlus; variety of grouvellei; Type locality: Le Kef).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; variety of grouvellei).
— Scheerpeltz, 1955b: 460 (Paratyphlus; type species of Paratyphlus).
— Coiffait, 1959: 318 (Paratyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
episcopus Coiffait, 1965: 138 (Paratyphlus; subgenus Ischnotyphlus; Type locality: Espagne: Prov. de Cadix, San Roque).
— Coiffait, 1972: 482 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
fourhalensis Coiffait, 1970b: 146 (Paratyphlus; Type locality: Berkane, Rif oriental, Maroc).
— Coiffait, 1972: 477 (Paratyphlus; characters; Morocco).
— Coiffait, 1973a: 274 (Paratyphlus; Morocco).
▴ Distribution: Morocco.
ginestensis Coiffait, 1959: 408 (Paratyphlus; subgenus Ischnotyphlus; Type locality: la région de la Punta Ginesta au Sud de Casteldefells, Province de Barcelone).
— Coiffait, 1972: 483 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
hipponensis Normand, 1939: 503 (Leptotyphlus; subgenus Paratyphlus; Type locality: Algérie: Bône).
— Coiffait, 1959: 318 (Paratyphlus; characters; Algeria).
— Coiffait, 1972: 477 (Paratyphlus; characters; Algeria).
▴ Distribution: Algeria.
kabylianus Normand, 1939: 504 (Leptotyphlus; subgenus Paratyphlus; Type locality: Algérie: Station forestière d’Adekar, entre Yakouren et Bougie).
mallorcensis Coiffait, 1955a: 68 (Paratyphlus; Type locality: Majorque: Puerto Andraitac).
— Coiffait, 1959: 318 (Paratyphlus; characters; Majorca).
— Coiffait, 1972: 478 (Paratyphlus; characters; Balearic Islands).
— Pons and Palmer, 1996: 160 (Paratyphlus; Balearic endemic; notes; habitat; Mallorca).
▴ Distribution: Balearic Islands.
mateui Coiffait, 1955a: 68 (Paratyphlus; Type locality: Prov. de Malaga: Benaojan).
— Coiffait, 1957: 69 (Paratyphlus; subgenus Ischnotyphlus).
— Coiffait, 1959: 322 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
— Coiffait, 1972: 483 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
menorquensis Coiffait, 1959: 406 (Paratyphlus; subgenus Paratyphlus; Type locality:
Baléares: Alcaufar près de San Luis dans la pointe Sud Est de l’Ile de Minorque).
— Coiffait, 1972: 479 (Paratyphlus; characters; Balearic Islands).
— Pons and Palmer, 1996: 160 (Paratyphlus; Balearic endemic; notes; habitat; Menorca).
▴ Distribution: Balearic Islands.
morandi Coiffait, 1957: 69 (Paratyphlus; subgenus Paratyphlus; Type locality: Espagne: Heredad San Juan à Pego).
— Coiffait, 1959: 320 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain). — Coiffait, 1972: 482 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
necropolis Coiffait, 1959: 410 (Paratyphlus; subgenus Ischnotyphlus; Type locality: au pied
d’un gros caroubier près de l’entrée du cimetière de Vendrell, Province de Tarragone).
— Coiffait, 1972: 483 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
pedroi Coiffait, 1957: 69 (Paratyphlus; subgenus Paratyphlus; Type locality: Espagne: Province de Tarragone: environs de Vendrell).
— Coiffait, 1959: 318 (Paratyphlus; characters; Spain). — Coiffait, 1972: 475 (Paratyphlus; characters; Spain).
▴ Distribution: Spain. regalis Coiffait, 1955, see: Mesotyphlus.
serratensis Coiffait, 1957: 69 (Paratyphlus; subgenus Ischnotyphlus; Type locality: Catalogne: Montserrat, Font dels Monjos).
— Coiffait, 1959: 322 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain). — Coiffait, 1972: 483 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
sitgensis Coiffait, 1959: 408 (Paratyphlus; subgenus Ischnotyphlus; Type locality: au pied de
figuiers dans la montagne à l’Est et près de Sitges, Province de Barcelone).
— Coiffait, 1972: 485 (Paratyphlus; subgenus Ischnotyphlus; characters; Spain).
▴ Distribution: Spain.
spiniphallus Coiffait, 1957: 69 (Paratyphlus; subgenus Paratyphlus; Type locality: Espagne: Province de Tarragone: Parello).
— Coiffait, 1959: 318 (Paratyphlus; characters; Spain). — Coiffait, 1972: 477 (Paratyphlus; characters; Spain).
▴ Distribution: Spain.
torressalai Coiffait, 1957: 68 (Paratyphlus; subgenus Paratyphlus; Type locality: Espagne: Vergel).
Tribe Leptotyphlini
Leptotyphlini Fauvel, 1874.
— Jakobson, 1908: 446, 477 (characters; list of species of Russia and western Europe).
— Saiz, 1973b: 289 (distribution).
— Outerelo and Gamarra, 1985: 16 (key to genera of Iberian Peninsula).
— Newton and Thayer, 1992: 56 (tribe of Leptotyphlinae).
— Pace, 1996: 59, 60 (characters).
Leptotyphli Fauvel, 1874: 329 [= 1874b: 35] (genera included: Leptotyphlus). Type genus:
Leptotyphlus (see Newton and Thayer, 1992: 56).
Leptotyphlina Fauvel, 1874.
Leptotyphlini Coiffait, 1957: 61, 70 (cited as new tribe).
— Coiffait, 1959: 274, 324 (characters; distribution; key to genera).
— Coiffait, 1963d: 380, 381 (characters; biogeographic notes).
— Scheerpeltz, 1968a: 34 (catalog of Austrian species).
— Coiffait, 1972: 383, 493 (characters; key to genera and species of western Palaearctic region).
— Orousset, 1984: 301 (list of species of Corsica).
— Newton and Thayer, 1992: 56 (synonym of Leptotyphlinae).
Leptotyphlitae Coiffait, 1957.
Afrotyphlus
[2 species; Ethiopian Region]
Afrotyphlus Fagel, 1954: 2 (species included: leleupi). Type species: Afrotyphlus leleupi Fagel, fixed by original designation and monotypy.
— Coiffait, 1957: 70 (characters).
— Coiffait, 1959: 386 (characters; type species: leleupi).
— Coiffait, 1972: 493 (characters).
leleupi Fagel, 1954: 3 (Afrotyphlus; Type locality: Congo belge: Kivu: territorie de Kabare, Nyakasiba, 1800 m).
— Coiffait, 1959: 386 (Afrotyphlus; characters; Belgian Congo).
— Coiffait, 1967b: 3 (Afrotyphlus; notes).
— Coiffait, 1977e: 660 (Afrotyphlus; notes; may be parthengenetic species; Rwanda).
▴ Distribution: Congo.
troglophilus Coiffait, 1967b: 4 (Afrotyphlus; Type locality: grotte Tala-Tala, mont Hoyo, 1170 m).
Eotyphlus
[4 species; Palaearctic Region]
Eotyphlus Coiffait, 1955a: 72 (species included: orientalis, olivieri, renouardi). Type species: Leptotyphlus orientalis Coiffait, fixed by original designation.
— Coiffait, 1957: 70 (characters).
— Coiffait, 1959: 382 (characters; key to species; type species: orientalis).
— Coiffait, 1972: 604 (characters; type species: orientalis; key to species of western Palaearctic region; references).
olivieri Coiffait, 1955a: 72 (Eotyphlus; Type locality: Liban: Beyrouth).
orientalis Coiffait, 1954a: 158 (Leptotyphlus; Type locality: Liban: Bater ech Chouf).
— Coiffait, 1955b: 72 (Eotyphlus; [Note: Coiffait did not cite the original reference nor the original genus; I assume that the orientalis he meant is the one he described in Leptotyphlus]; cited as type species of Eotyphlus).
— Coiffait, 1959: 384 (Eotyphlus; characters; Lebanon).
— Coiffait, 1972: 607 (Eotyphlus; characters; Lebanon).
▴ Distribution: Lebanon.
palaestinus Thelot, 1985: 681 (Eotyphlus; Type locality: Israël-Galilée: Ginosar, 200 m).
renouardi Coiffait, 1955a: 72 (Eotyphlus; Type locality: Liban: Kahalé).
Epalxotyphlus
[1 species; Palaearctic Region]
Epalxotyphlus Coiffait, 1955a: 68 (species included: lavangnei). Type species: Leptotyphlus lavagnei Normand, fixed by original designation and monotypy.
— Coiffait, 1957: 70 (characters).
— Coiffait, 1959: 330 (characters; type species: lavagnei).
— Coiffait, 1972: 495 (characters; type species: lavagnei; species of western Palaearctic region).
lavagnei Normand, 1916: 137 (Leptotyphlus; Type locality: environs de Montpellier).
Hesperotyphlus
[20 species; Palaearctic Region]
Herperotyphlus Coiffait, 1955a: 72 (subgenus of Leptotyphlus; species included: jeanneli).
Type species: Leptotyphlus jeanneli Jarrige, fixed by original designation.
— Coiffait, 1959: 332, 334, 378 (subgenus of Leptotyphlus; characters; key to species; type species: jeanneli).
— Coiffait, 1965b: 690 (characters; notes; key to species of Portugal).
— Coiffait, 1972: 497 (characters; valid genus; type species: jeanneli; key to species of western Palaearctic region; references).
— Luna de Carvalho, 1982: 201 (key to males [Portugal]).
albercanus Outerelo, 1977a: 26 (Hesperotyphlus; Type locality: El Cabaco, Salamanca a unos 1100 m, el la base de La Peña de Francia).
algarvensis Coiffait, 1965b: 690, 696 (Hesperotyphlus; Type locality: Portugal: Algarve, Porto de Lagos près de Portimao).
— Coiffait, 1972: 499 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 201 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
amoenus Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Minho, Amarante).
— Coiffait, 1972: 500 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 201 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
baetanevesi Luna de Carvalho, 1983: 200 (Hesperotyphlus; Type locality: Portugal: Viseu, Parque do Fontelo, 40.39N, 7.55W).
batuequensis Outerelo, 1977a: 24 (Hesperotyphlus; Type locality: el valle del rio Batuecas, Salamanca, 590 m).
beirensis Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Beira Litoral, parc de Buçaco).
— Coiffait, 1972: 501 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 202 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
besucheti Coiffait, 1973b: 222 (Hesperotyphlus; Type locality: Portugal: district de Coïmbra, Serra de Lousa, Candal, 700 m).
carballalae Outerelo, 1977: 70 (Hesperotyphlus; Type locality: Moreda del Caurel, Sierra del Caurel, Lugo, 720 m).
estrelensis Coiffait, 1965b: 694, 696 (Hesperotyphlus; Type locality: Portugal: Serra de Estrela, Manteigas).
— Coiffait, 1972: 502 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 202 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
faramellus Outerelo, 1978: 23 (Hesperotyphlus; Type locality: España: el valle del rio Tinto, cuenca del Sar, lugar de Faramello, cerca de Santiago de Compostela, La Coruña, 60 m).
hurdanus Outerelo, 1977a: 26 (Hesperotyphlus; Type locality: Nuñomoral Cáceres en el valle del rio Hurdano, comarca de las Hurdes a unos 480 m).
jeanneli Jarrige, 1949: 61 (Leptotyphlus; Type locality: Pena-Maïor, N. Portugal).
— Coiffait, 1955a: 72 (Leptotyphlus; subgenus Hesperotyphlus).
— Coiffait, 1959: 379 (Leptotyphlus; subgenus Hesperotyphlus; characters; Portugal).
— Coiffait, 1965b: 692, 695 (Hesperotyphlus; characters; Portugal).
— Coiffait, 1972: 500 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 201 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
jucundus Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Minho, Amarante).
— Coiffait, 1972: 500 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 201 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
montanus Coiffait, 1965b: 694, 695 (Hesperotyphlus; Type locality: Portugal: Serra de Estrela, Manteigas).
novoai Outerelo, 1977: 65 (Hesperotyphlus; Type locality: Moscoso, Pontevedra, 300 m).
realensis Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Tras os Montes, environs de Vila Real).
— Coiffait, 1972: 500 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 201 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
seminarius Coiffait, 1979b: 314 (Hesperotyphlus; Type locality: Porto, jardin du séminaire de Vilar).
socius Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Serra de Estrela, Manteigas).
validus Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Tras os Montes, Serra de Marao, Campea).
— Coiffait, 1972: 500 (Hesperotyphlus; characters; Portugal).
— Luna de Carvalho, 1982: 202 (Hesperotyphlus; characters).
▴ Distribution: Portugal.
vicinus Coiffait, 1965b: 692, 695 (Hesperotyphlus; Type locality: Portugal: Beira Litoral, Buçaco).
Kenotyphlus
[4 species; Palaearctic Region]
Kenotyphlus Coiffait, 1957: 71, 80 (subgenus of Leptotyphlus; species included: turcicus, pisidicus, pamphylicus). Type species: Kenotyphlus turcicus Coiffait, fixed by original designation.
— Coiffait, 1959: 380 (characters; key to species; type species: turcicus).
— Coiffait, 1972: 602 (characters; type species: turcicus; key to species of western Palaearctic region; references).
pamphylicus Coiffait, 1957: 80 (Kenotyphlus; Type locality: Turquie: d’Asie: Environs d’Antalya).
— Coiffait, 1959: 382 (Kenotyphlus; characters; Turkey).
— Coiffait, 1972: 604 (Kenotyphlus; characters; Turkey).
▴ Distribution: Turkey.
pisidicus Coiffait, 1957: 80 (Kenotyphlus; Type locality: Turquie d’Asie: Dinar à 50 km au nord de Burdur).
— Coiffait, 1959: 382 (Kenotyphlus; characters; Turkey).
— Coiffait, 1972: 604 (Kenotyphlus; characters; Turkey).
▴ Distribution: Turkey.
rhodiensis Coiffait, 1973b: 222 (Kenotyphlus; Type locality: Ile de Rhodes, Petaloudes).
turcicus Coiffait, 1957: 80 (Kenotyphlus; Type locality: Turquie d’Asie: près d’une source au point culminant de la route Burdur-Antalya, dans la traversée du Darim Dag).
Kilimatyphlus
[1 species; Ethiopian Region]
Kilimatyphlus Coiffait, 1959: 384 (species included: babas). Type species: Leptotyphlus babas Tottenham, fixed by original designation and monotypy.
babas Tottenham, 1953b: 487 (Leptotyphlus; Type locality: Kilimanjaro: Lyamungu, 4300 ft.).
Leptotyphlus
[201 species; Palaearctic Region]
Leptotyphlus Fauvel, 1874: 330 [= 1874b: 36] (species included: sublaevis). Type species: Leptotyphlus sublaevis Fauvel, fixed by monotypy.
— Fauvel, 1875a: XIV [= 1875b: 216] (catalog).
— Mulsant and Rey, 1878a: 328 (characters).
— Duvivier, 1883: 178 (catalog).
— Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 606 (characters).
— Dodero, 1908: 634 (key to species).
— Heyden, Reitter, and Weise, 1906: 145 (list of species of Europe and Caucasus).
— Bernhauer and Schubert, 1911: 146 (world catalog; 11 species).
— Winkler, 1925: 347 (catalog for Palaearctic region).
— Porta, 1926: 49 (characters; key to species of Italy).
— Portevin, 1929: 391 (characters; key to French species).
— Scheerpeltz, 1933: 1138 (world catalog supplement).
— Normand, 1935: 363 (list of species of Tunisia).
— Normand, 1939: 486 (notes; characters).
— Blackwelder, 1952: 217 (type species: sublaevis).
— Scheerpeltz, 1955b: 458 (characters in key; type species: sublaevis; list of species).
— Coiffait, 1957: 71 (characters; key to subgenera).
— Coiffait, 1959: 332, 340 (characters; key to species; type species: sublaevis).
— Székessy, 1963: 34 (characters).
— Hervé, 1965: 85 (notes).
— Hervé, 1966a (composition of angelicus and furcatus species groups).
— Coiffait, 1972: 505 (characters; type species: sublaevis; key to species of western Palaearctic region; references).
— Pace, 1977: 198 (key to species of Apennines).
— Orousset, 1987c: 127 (notes, characters, and list of species for tyrrhenius group of species).
— Zanetti, 1995: 15 (list of species of Italy).
— Pace, 1996: 152 (characters; key to subgenera; key to species of Italy).
Stigmotyphlus Coiffait, 1955a: 68 (subgenus of Leptotyphlus; species included: cribratus, grannus, sylvanus, belenus, sucellus, esus, camulus, bormanus, impressus). Type species: Leptotyphlus cribratus Fauvel, fixed by original designation.
— Coiffait, 1957: 72 (characters; key to species groups).
— Coiffait, 1959: 332, 334 (subgenus of Leptotyphlus; characters; key to species; type species: cribratus).
— Coiffait, 1972: 505, 509, 553 (subgenus Leptotyphlus; characters; type species: cribratus; key to species of western Palaearctic region; references).
— Pace, 1996: 155, 240 (subgenus of Leptotyphlus; characters; key to species of Italy).
Anomotyphlus Coiffait, 1972: 505, 508, 583 (subgenus of Leptotyphlus; species included: minos, hades). Type species: Leptotyphlus hades Coiffait, fixed by original designation.
— Pace, 1995: 30, 31 (subgenus of Leptotyphlus; key to species).
— Pace, 1996: 155, 275 (subgenus of Leptotyphlus; characters; species of Italy).
Odontotyphlus Coiffait, 1972: 505, 509, 563 (subgenus of Leptotyphlus; species included: brevicornis, dentatus, priapus, londensis, sassii, lucensis, asinus, transensis, schuleri, joannis, collensis, domensis, maurettensis, laminatus, janasensis, dujardini, dispersus, angelicus, balachovskyi, phoenix, strobinoi, crassiphallus, langastouensis, gardensis, laneyriei, compitalis, garonnettensis, dispar, sublaneyriei, fissuralis, henryi, arcsensis, monachus, lobatus, furcatus, argensis, trifurcatus, tressensi, calenzanus, omessae, kyrnosus, coiffaiti, occidentalis). Type species: Leptotyphlus angelicus Coiffait, fixed by original designation.
Subhesperotyphlus Coiffait, 1972: 505, 509, 510 (subgenus of Leptotyphlus; species included: exilis, solarii). Type species: Cylindrogaster exilis Mulsant and Rey, fixed by original designation.
Porotyphlus Pace, 1991: 110 (subgenus of Leptotyphlus; species included: calabricus, circaeus, brevicornis, viti, finalensis, apenninus). Type species: Leptotyphlus calabricus Coiffait, fixed by original designation.
agrestis Orousset, 1984b: 195 (Leptotyphlus; subgenus Odontotyphlus; Type locality: Forêt d’Aitone, Est d’Evisa, Corse-du-sud, alt. 1000 m).
— Orousset, 1988a: 91 (Leptotyphlus; subgenus Odontotyphlus; Corsica).
— Pace, 1996: 268 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
aithaliensis Orousset, 1983a: 293 (Leptotyphlus; subgenus Leptotyphlus; Type locality: San Felo, île d’Elbe).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 181 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
albareensis Coiffait, 1957: 77 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Alpes Mar.: Sospel, Albarea, Mt. Razet).
— Coiffait, 1959: 368 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 545 (Leptotyphlus; subgenus Leptotyphlus; cited as albarensis; characters; Maritime Alps).
— Pace, 1996: 218 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
albingaunicus Pace, 1977b: 296 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: presso Berioli, prov. di Savona, a circa 400 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 245 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
alutaceus Dodero, 1908, see: Mesotyphlus.
anachoreta Pace, 1975: 154 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Grisolia, provincia di Cosenza, nella Calabria settentrionale a circa 550 metri d’altitudine, in linea d’aria a Km 8.5 di distanza dalla costa tirrenica, ai piedi del Monte Carpinoso, presso il torrente Vaccuta).
— Pace, 1977: 199 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 232 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
anchorifer Coiffait, 1957: 76 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Arige, Vallèe du Garbet: Ercé; Haute Garonne: Castelbiague; Arbas, entrée de la grotte de Gourgue).
— Coiffait, 1959: 358 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 530, 532 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
angelicus Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Var: Notre-Dame-des-Anges).
— Coiffait, 1959: 377 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Hervé, 1967c: 230 (Leptotyphlus; characters).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
angusticeps Normand, 1939, see: Entomoculia.
antonius Coiffait, 1959: 420 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Saint-Antonin [Note: See Orousset, 1985, for correction of type locality]).
— Coiffait, 1972: 547 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1985: 177 (Leptotyphlus; correction of type locality: Saint Antoine; characters; Corsica).
— Pace, 1996: 215 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
antrorum Coiffait, 1959: 414 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Aude: la grotte du Roc de l’Homme mort en forêt de Bélesta à Puivert).
— Coiffait, 1972: 519 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
apenninus Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus; Type locality: Italia: provincia di Bologna: Emilia, Pian del Voglio).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 279 (Leptotyphlus; subgenus Porotyphlus; characters; Italy).
▴ Distribution: Italy.
arcsensis Coiffait, 1959: 425 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Vidauban).
— Coiffait, 1972: 577 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
argensis Coiffait, 1957: 79 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: rive droite de l’Argens entre Roquebrune et Le Mury).
— Coiffait, 1959: 378 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 579 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France. argodi Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Alpes-Mar.: Les Baumettes).
— Coiffait, 1959: 368 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 545 (Leptotyphlus; subgenus Leptotyphlus; characters; Maritime Alps).
▴ Distribution: France.
argus Normand, 1939, see: Entomoculia.
arizanus Coiffait, 1959: 412 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariège: la vallée boisée de l’Artillac, versant Nord du Massif de l’Arize, commune de Sentenac-de Sérou).
— Coiffait, 1972: 530, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
asinus Coiffait, 1962d: 181 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Forêt communale de Gonfaron, Canton de Plaines).
— Hervé, 1967c: 226, 230 (Leptotyphlus; subspecies of angelicus; characters).
— Coiffait, 1972: 569 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
aspericeps Scheerpeltz, 1931, see: Allotyphlus.
asprensis Coiffait, 1957: 78 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyr. orientales: Bouleternère).
— Coiffait, 1959: 366 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 543 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France. attis Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Pyr.-Orientales: Banyuls).
— Coiffait, 1959: 352 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 523 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
audrasi Jarrige, 1949: 63 (Leptotyphlus; Type locality: Lyon, Rhône).
— Coiffait, 1959: 346 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 517 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
aurensis Coiffait, 1957: 76 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Hautes-Pyr., Vallée d’Aure: Heches; Vallée d’Aure à Bagnères de Bigorre).
— Coiffait, 1959: 356 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 529, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
babas Tottenham, 1953, see: Kilimatyphlus.
baillauriensis Coiffait, 1962d: 177 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Banyuls, vallée de la Baillaurie, rive droite).
— Coiffait, 1972: 543 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
balachowskyi Hervé, 1966b: 109 (Leptotyphlus; Type locality: un peu à l’ouest du village du Plan de la Tour sur le chemin de Sigalas, Var).
— Hervé, 1967c: 230, 232 (Leptotyphlus; characters).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; cited as balachovskyi; characters; France).
▴ Distribution: France.
bastiensis Coiffait, 1973b: 220 (Leptotyphlus; Type locality: Corse: environs de Bastia).
— Orousset, 1987c: 130 (Leptotyphlus; synonym of devillei).
— Pace, 1996: 183 (Leptotyphlus; subgenus Leptotyphlus; cited as valid species; characters; Corsica).
▴ Distribution: Corsica.
beieri Scheerpeltz, 1931, see: Rhopalotyphlus.
belenus Coiffait, 1955a: 68 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Mar.: Caussols).
— Coiffait, 1959: 338 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 559 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
bicorniger Orousset, 1985: 179 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Rocchiu-Pinzutu, entre la Bocca di Larone et Solenzara, commune di Quenza, alt. 400 m, Corse du sud).
— Pace, 1996: 216 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
bifurcus Orousset, 1984: 306 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pont de Truggia, vallée du Liamone, alt. 75 m, Corse du sud).
— Pace, 1996: 229 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
bigerricus Coiffait, 1959: 358 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Hautes-Pyrénées: Ste-Marie-de-Campan, rive droite de l’Adour).
— Coiffait, 1972: 532, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
boiteli Normand, 1950, see: Paratyphlus.
bonadonai Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Grasse à l’Estérel).
— Coiffait, 1959: 362 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 536 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; characters; France).
▴ Distribution: France.
tellus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Var: Tanneron).
— Coiffait, 1959: 364 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 538 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; synonym of bonadonai).
bordei Coiffait, 1957: 78 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Bonifacio).
— Coiffait, 1959: 370 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 547 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 191 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
bormanus Coiffait, 1955a: 69 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Mar. Vence).
— Coiffait, 1959: 338 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 559 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
bouilloni Coiffait, 1959: 344 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariège: St-Lary, près des anciennes ardoisières).
— Coiffait, 1972: 515 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
boulouensis Coiffait, 1962d: 176 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Le Boulou, près de l’établissement thermal).
— Coiffait, 1972: 542 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: Pyrenees.
brescius Thelot, 1980: 52 (Leptotyphlus; subgenus Odontotyphlus; Type locality: France: petit ravin secondaire perpendiculaire à la Bresque, Commune de Salernes, Var).
brevicornis Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Sardaigne: Aritzo).
— Coiffait, 1959: 355 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Coiffait, 1972: 565 (Leptotyphlus; subgenus Odontotyphlus; characters; Sardinia).
— Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 283 (Leptotyphlus; subgenus Porotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
calabricus Coiffait, 1970a: 141 (Leptotyphlus; Type locality: Sambiase, Calabre).
— Coiffait, 1972: 529 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Pace, 1975: 164 (Leptotyphlus; characters of female; Italy).
— Pace, 1977: 200 (Leptotyphlus; characters; Italy).
— Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus; type species of Porotyphlus).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 281 (Leptotyphlus; subgenus Porotyphlus; characters; Italy).
▴ Distribution: Italy.
calcatogius Coiffait, 1959: 416 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Calcatogio près Sari d’Orcino).
— Coiffait, 1972: 549 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 130 (Leptotyphlus; subgenus Leptotyphlus; Corsica).
— Pace, 1996: 209 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
calenzanus Coiffait, 1959: 421 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Calenzana).
— Coiffait, 1972: 581 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Orousset, 1984b: 190 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Pace, 1996: 272 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
campanus Pace, 1977: 183 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: pendici occidentali dei Monti Picentini tra Montecorvino e Acerno, provincia di Salerno, sulla detra del fiume Tusciano a circa 600 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 261 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
camulus Coiffait, 1955a: 69 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Mar. Grasse).
— Coiffait, 1959: 338 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 559 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
capito Normand, 1939, see: Entomoculia.
cargeseensis Orousset, 1984: 303 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Sud de la Forêt d’Esigna, près de Cargèse, alt. 150 m, Corse du sud).
— Pace, 1996: 232 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
cariniventris Coiffait, 1959: 411 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Maritimes: dans la basse vallée de l’Estéron à Gilette).
— Coiffait, 1962d: 172 (Leptotyphlus; subgenus Stigmotyphlus; aedeagus).
— Coiffait, 1972: 561 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
carniolicus Bernhauer, 1908, see: Megatyphlus.
castillonensis Coiffait, 1957: 75 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Alpes Maritimes: Forêt de Menton, commune de Castillon).
— Coiffait, 1959: 352 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 523 (Leptotyphlus; subgenus Leptotyphlus; characters; Maritime Alps).
— Pace, 1996: 221 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: France, Italy.
chappuisi Coiffait, 1959: 412 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariège: au col du Saraillé, commune d’Ercé).
— Coiffait, 1972: 527 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
charon Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Tarn: Castres).
— Coiffait, 1959: 342 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 512 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
cheironensis Thelot, 1984: 236, 240 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Conségudes - Alpes Maritimes, ubac du bois de Cheiron dans la clue de la Bouisse).
chiavariensis Orousset, 1984: 307 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Forêt de Chiavari, au sud d’Ajaccio, alt. 200 m, Corse du sud).
— Pace, 1996: 196 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
circaeus Pace, 1977: 194 (Leptotyphlus; Type locality: versante settentrionale del Monte Circeo, Lazio, presso Torre Paola, entro i confini del Parco Nazionale).
— Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 280 (Leptotyphlus; subgenus Porotyphlus; characters; Italy).
▴ Distribution: Italy.
coiffaiti Hervé, 1964: 295 (Leptotyphlus; Type locality: versant nord des Pradels, dans les Maures, Var).
— Coiffait, 1972: 581 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
solitarius Hervé, 1964: 300 (Leptotyphlus; subspecies of coiffaiti; Type locality: Var, Ramatuelle).
occidentalis Hervé, 1964a: 598 (Leptotyphlus; subspecies of coiffaiti; Type locality: Cap Bénat, Var).
collensis Coiffait, 1959: 374 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Le Pradet, Forêt de la Colle Noire).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
compitalis Hervé, 1965: 90 (Leptotyphlus; Type locality: France: au crosement des routes Ste-Maxime, le Muy, Plan de la Tour, Var).
— Coiffait, 1972: 575 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
garonnettensis Hervé, 1965: 90 (Leptotyphlus; subspecies of compitalis; Type locality: France: vallon de la Garonnette, au Nord-Est de Sainte-Maxime, Var).
confrater Normand, 1947a: 6 (Leptotyphlus; variety of tuniseus; Type locality: Tunisie: Aïn-Draham; Fernana).
— Coiffait, 1959: 340 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
— Coiffait, 1972: 563 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
contractus Hervé, 1978, see: priapus Coiffait, 1955.
corsicus Coiffait, 1955, see: omessae Dodero, 1908.
couseranus Coiffait, 1959: 350 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariège: environs de St-Girons, Lacourt).
— Coiffait, 1972: 523 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
crassiphallus Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Var: Pont des Veyans).
— Coiffait, 1959: 376 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
cribratus Fauvel, 1890: 356 (Leptotyphlus; Type locality: Nice).
— Croissandeau, 1891: 150 (Leptotyphlus; cited as cribrosus Fauvel; characters; France).
— Ganglbauer, 1895: 607 (Leptotyphlus; characters; France).
— Bernhauer, 1908e: 332 (Leptotyphlus; characters).
— Dodero, 1908: 636, 640 (Leptotyphlus; characters; Italy).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Dodero, 1916: 341 (Leptotyphlus; Italy).
— Portevin, 1929: 392 (Leptotyphlus; characters; France).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1959: 338 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 559 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Pace, 1996: 244 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: France, Italy.
cribriceps Bernhauer, 1908e: 331 (Leptotyphlus; Type locality: Corsica).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Scheerpeltz, 1955b: 461 (Cyclotyphlus).
— Coiffait, 1972: 555 (Leptotyphlus; subgenus Stigmotyphlus; cited as “Bernh. in litt.” characters; Corsica).
— Pace, 1996: 248 (Leptotyphlus; subgenus Stigmotyphlus; characters; Corsica).
▴ Distribution: Corsica.
curtii Breit, 1911, see: Megatyphlus.
curtulus Jarrige, 1949, see: Gynotyphlus.
cyclops Normand, 1939, see: Entomoculia.
cyrneus Orousset, 1984: 304 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pastric-ciola, est de Vico, alt. 550 m, Corse du sud).
— Pace, 1996: 227 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
dapsilis Normand, 1950, see: Entomoculia.
demeter Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Pyr.-Orientales: La Preste).
— Coiffait, 1959: 350 (Leptotyphlus; subgenus Leptotyphlus; characters; France; Spain).
— Coiffait, 1972: 521, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain; France).
▴ Distribution: France, Spain.
dentatus Coiffait, 1957: 76 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Ajaccio).
— Coiffait, 1959: 354 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 565 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Orousset, 1984b: 188 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Pace, 1996: 268 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
denticulatus Coiffait, 1957: 73 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Sardaigne: Arizto).
— Coiffait, 1959: 336 (Leptotyphlus; subgenus Stigmotyphlus; characters; Sardinia).
— Coiffait, 1972: 557 (Leptotyphlus; subgenus Stigmotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 239 (Leptotyphlus; subgenus Subhesperotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
devillei Coiffait, 1957: 77 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: ? Omessa).
— Coiffait, 1959: 366 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 540 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 130 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 177 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
nonzius Coiffait, 1959: 416 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Nonza).
— Coiffait, 1972: 553 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 130 (Leptotyphlus; synonym of devillei).
— Pace, 1996: 177 (Leptotyphlus; synonym of devillei).
dispar Hervé, 1965: 88 (Leptotyphlus; Type locality: France: Beauvallon, entre Sainte-Maxime et Grimaud, Var).
— Coiffait, 1972: 577 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
dispersus Hervé, 1967c: 228 (Leptotyphlus; Type locality: versant nord des Pradels, La Môle, Var).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
vignesi Thelot, 1975: 47 (Leptotyphlus; subspecies of dispersus; Type locality: France: Var: massif des Maures, forêt du Dom de Bormes).
distinctus Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Hérault: Castelnau).
— Coiffait, 1959: 366 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 543 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
doderoi Normand, 1910, see: Paratyphlus.
doderoi Coiffait, 1957, see: sardiniensis Herman, 2001.
domensis Coiffait, 1957: 79 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Auberge du Dom).
— Coiffait, 1959: 374 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Hervé, 1966: 116 (Leptotyphlus; characters).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
dubius Coiffait, 1957: 75 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse).
— Coiffait, 1959: 348 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 519 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 134 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 183 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
dujardini Hervé, 1967a: 250 (Leptotyphlus; Type locality: ravin en haut du vallon du Fournel, château de la Mère, Sainte-Maxime, Var).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
eaqus Coiffait, 1957: 74 (Leptotyphlus; subgenus Leptotyphlus; [Note: Cited as eaqus in a paragraph preceding the description but the name with the description was caqus. Coiffait used eaqus in two subsequent publications so I accept that as the proper spelling and treat caqus as a lapsus]; Type locality: Hérault: Clermont l’Hérault).
— Coiffait, 1959: 344 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 513 (Leptotyphlus; subgenus Leptotyphlus; characters).
▴ Distribution: France.
elbanus Pace, 1980a: 159 (Leptotyphlus; Type locality: Isola d’Elba, M.te Capanne, versante nord e a metà valle sopra Poggio, 400–500 m).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 187 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
esus Coiffait, 1955a: 69 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Mar. Pegomas).
— Coiffait, 1959: 340 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 561 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
▴ Distribution: France.
insularis Coiffait, 1972: 563 (Leptotyphlus; subgenus Stigmotyphlus; subspecies of esus; [Note: Cited in a key and with Hervé as author however I have not found this name described by Hervé and Coiffait did not give a reference to it]; Type locality: Var, Ile de Port Cros).
eurydice Coiffait, 1962d: 172 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: St-Martin-d’Albère).
— Coiffait, 1972: 521 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
exilis Mulsant and Rey, 1875: 131 (Cylindrogaster; Type locality: environs de Massane, Pyrénées-Orientales).
— Mulsant and Rey, 1876: 12 (Cylindrogaster; characters; France).
— Fauvel, 1876a: 61 [= 1876: 246] (Leptotyphlus; synonym of sublaevis Fauvel).
— Saulcy, 1878: 121 (Leptotyphlus; characters).
— Mulsant and Rey, 1878a: 330 (Leptotyphlus; [Note: Misidentification: Bernhauer and Schubert, 1911: 147 cited sublaevis Mulsant and Rey as a synonym of exilis; Mulsant and Rey did not describe the species as new, they attributed it to Fauvel]; exilis cited as synonym of sublaevis; characters; France).
— Croissandeau, 1891: 150 (Leptotyphlus; characters; Pyrenees).
— Dodero, 1908: 636, 639 (Leptotyphlus; characters; Italy).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Coiffait, 1959: 379 (Leptotyphlus; subgenus Hesperotyphlus; characters; France; Spain).
— Coiffait, 1972: 510 (Leptotyphlus; subgenus Subhesperotyphlus; characters; France; Spain).
▴ Distribution: France, Italy, Spain.
eximius Orousset, 1988b: 329 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse-du-Sud: Calalonga, commune de Bonifacio, alt. 20 m).
— Pace, 1996: 230 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
exsculptus Normand, 1939, see: Entomoculia.
ezeensis Coiffait, 1957: 77 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Alpes Mar.: Eze-sur-Mer).
— Coiffait, 1959: 368 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 543 (Leptotyphlus; subgenus Leptotyphlus; characters; Maritime Alps).
— Pace, 1996: 220 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
fallax Orousset, 1985: 179 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Rocchiu-Pinzutu, entre la Bocca di Larone et Solenzara, commune de Quenza, alt. 400 m, Corse du sud).
— Pace, 1996: 212 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
farinolensis Orousset, 1985d: 287 (Leptotyphlus; subgenus Leptotyphlus; Type locality: France: Corse: Sparagaggio, commune de Farinole, altitude 150 m, Haute-Corse).
— Orousset, 1987c: 132 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 196 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
faunus Coiffait, 1955, see: hervei Jarrige, 1949.
finalensis Pace, 1977b: 291 (Leptotyphlus; Type locality: presso Calice Ligure, lungo la strada per Eze).
— Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 279 (Leptotyphlus; subgenus Porotyphlus; characters; Italy).
▴ Distribution: Italy.
fissuralis Coiffait, 1959: 376 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Pignans, Vallon des Martels).
— Coiffait, 1972: 577 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
fonfrigidus Coiffait, 1962d: 173 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Céret, route de Fonfrède).
— Coiffait, 1972: 519, 525, 585 (Leptotyphlus; subgenus Leptotyphlus; cited as fontfrigidus and fontrigidus but in the index as fonfrigidus; characters; Pyrenees).
▴ Distribution: France.
foroiuliensis Pace, 1976: 112 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: versante settentrionale del M. Jof a circa 410 m; versante settentrionale del Monte S. Lorenzo a circa 600 m).
— Pace, 1978a: 199 (Leptotyphlus; subgenus Stigmotyphlus; notes; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 256 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
foveiceps Dodero, 1908: 636, 639 (Leptotyphlus; Type locality: Voltaggio, Borzoli, Ruta e Rapallo, tutte localitá dell’Apennino genovese).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Coiffait, 1959: 335 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
— Coiffait, 1972: 556 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
— Pace, 1977: 198 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 248 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
furcatus Coiffait, 1957: 79 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Collobrière des Maures).
— Coiffait, 1959: 378 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Hervé, 1964: 295 (Leptotyphlus; notes; France).
— Hervé, 1965: 92 (Leptotyphlus; characters; notes).
— Coiffait, 1972: 579 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
galiberti Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Tarn: Castres).
— Coiffait, 1959: 358 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 532 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
garbetensis Coiffait, 1957: 76 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariege, Vallée du Garbet: Aulus).
— Coiffait, 1959: 358 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 530, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
gardensis Coiffait, 1959: 425 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Garde Freinet).
— Coiffait, 1972: 575 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
garonnettensis Hervé, 1965, see: compitalis Hervé, 1965.
giglionicus Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Toscane: Ile de Giglio).
— Coiffait, 1959: 348 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Coiffait, 1972: 519 (Leptotyphlus; subgenus Leptotyphlus; characters; Giglio Island).
— Coiffait, 1972: 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Giglio Island).
— Pace, 1977: 200 (Leptotyphlus; characters; Italy).
— Pace, 1980e: 161 (Leptotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 185 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
gracilis Coiffait, 1957: 74 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyr. Orientales: Montbollo près Amélie-les-Bains).
— Coiffait, 1959: 346 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 517 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
grannus Coiffait, 1955a: 68 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Vaucluse: Bedoin).
— Coiffait, 1959: 335 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 555 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
▴ Distribution: France.
grouvellei Fauvel, 1890, see: Mesotyphlus.
grouvellei Croissandeau, 1891 [Note: Misidentified name], see: Entomoculia.
hades Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Var: Notre-Dame-des Anges).
— Coiffait, 1959: 372 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 583 (Leptotyphlus; subgenus Anomotyphlus; characters; France).
— Pace, 1995: 31 (Leptotyphlus; subgenus Anomotyphlus; characters).
▴ Distribution: France.
henryi Coiffait, 1959: 423 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Ile de Port Cros).
— Coiffait, 1972: 577 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
hervei Jarrige, 1949: 62 (Leptotyphlus; Type locality: l’Estérel).
— Coiffait, 1959: 362 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 536 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; characters; France).
▴ Distribution: France.
faunus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Var: Auberge des Adrets).
— Coiffait, 1959: 362 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 538 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; synonym of hervei).
vulcanus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Var: Auberge des Adrets).
— Coiffait, 1959: 362 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 536 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; synonym of hervei).
subhervei Coiffait, 1959: 420 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: l’Estérel à Agay).
— Coiffait, 1972: 538 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Thelot, 1984: 240 (Leptotyphlus; synonym of hervei).
hipponensis Normand, 1939, see: Paratyphlus.
ilvensis Orousset, 1983a: 291 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Procchio, île d’Elbe).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 173 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
impressus Coiffait, 1955a: 69 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Sardaigne: Teulada).
— Coiffait, 1959: 336 (Leptotyphlus; subgenus Stigmotyphlus; characters; Sardinia).
— Coiffait, 1972: 515 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 223 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
inopinatus Pace, 1975: 157 (Leptotyphlus; Type locality: presso Sambiase, Catanzaro, a Km 4 a nord dal abitato, vicino al villaggio di Gabella a circa 560 metri d’altitudine, in una valletta presso il torrente Cantagalli).
— Pace, 1977: 200 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 206 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
insularis Coiffait, 1972, see: esus Coiffait, 1955.
irenae Coiffait, 1952: 16 (Leptotyphlus; Type locality: Toulouse, Jardin des Plantes, France).
— Scheerpeltz, 1955b: 460 (Oligotyphlus; cited as ireneae).
— Coiffait, 1959: 358 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 532, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
janasensis Hervé, 1966: 117 (Leptotyphlus; Type locality: France: forêt de Janas, cap Sicié, Var).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
jeanneli Jarrige, 1949, see: Hesperotyphlus.
joannis Hervé, 1971: 119 (Leptotyphlus; Type locality: France: Var: ravin secondaire du vallon de la Foux, au nord de Cuers).
— Coiffait, 1972: 569 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Thelot, 1979: 20 (Leptotyphlus; subgenus Odontotyphlus; notes).
▴ Distribution: France.
jupiter Normand, 1939, see: Entomoculia.
kabylianus Normand, 1939, see: Paratyphlus.
kahleni Pace, 1991: 110 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Italia: provincia di Vercelli: Piemonte, Pecco Ripa, Valchiusella, am fuss Eiche).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 251 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
kovaci Šustek, 2000: 151 (Leptotyphlus; Type locality: E. Slovakia, at entrance of the Ardovská jaskyň cave, 48°31′ N, 20°25′ E, 317 m, near Ardovo village).
kyrnosus Coiffait, 1957, see: omessae Dodero, 1908.
laetificus Normand, 1939, see: Entomoculia.
lamberti Thelot, 1979: 17 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: vallon des Veys, commune de Cuers, Var).
laminatus Coiffait, 1959: 422 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: sur le plateau de Grand Cap à Morière-le-Cap, au Nord de Toulon).
— Hervé, 1971: 116, 118 (Leptotyphlus; characters of female; France).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Thelot, 1979: 20 (Leptotyphlus; subgenus Odontotyphlus; notes).
▴ Distribution: France.
laneyriei Hervé, 1965: 86 (Leptotyphlus; Type locality: France: le Plan de la Tour, dans les Maures, Var).
— Coiffait, 1972: 575 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
langastouensis Hervé, 1969: 251 (Leptotyphlus; Type locality: vallon de Langastoua, massif des Maures, commune de Vidauban, Var).
— Coiffait, 1972: 575 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
laticeps Dodero, 1908: 636, 639 (Leptotyphlus; Type locality: Omessa; Corsica).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; synonym of cribriceps).
— Coiffait, 1959: 335 (Leptotyphlus; subgenus Stigmotyphlus; cited as valid species; characters; Corsica).
— Coiffait, 1972: 555 (Leptotyphlus; subgenus Stigmotyphlus; characters; Corsica).
— Pace, 1996: 250 (Leptotyphlus; subgenus Stigmotyphlus; characters; Corsica).
▴ Distribution: Corsica.
lavagnei Normand, 1916, see: Epalxotyphlus.
lepinensis Pace, 1977: 185 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Monti Lepini, presso Carpineto Romano, tra i 600 e i 700 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 203 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
lessinicus Pace, 1973: 313 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Monti Lessini: Valle d’Alpone, Castelvero).
— Pace, 1976: 114 (Leptotyphlus; characters).
— Pace, 1978a: 197 (Leptotyphlus; subgenus Stigmotyphlus; notes; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 253 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
levasseuri Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Nice: Vallon des Fleurs).
— Coiffait, 1959: 368 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 545 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Pace, 1996: 216 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: France, Italy.
liguricus Coiffait, 1957: 78 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Italie: Ligurie: Arenzano).
— Coiffait, 1959: 370 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Coiffait, 1972: 549 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Pace, 1977: 198 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 199 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
lobatus Coiffait, 1959: 424 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Maures orientales à Beauvallon).
— Coiffait, 1972: 579 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
londensis Coiffait, 1959: 422 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Pas du Cerf dan le vallon du Pansard au Nord de La Londe-les-Maures).
— Hervé, 1964a: 598 (Leptotyphlus; notes; France).
— Coiffait, 1972: 567 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Hervé, 1978: 138 (Leptotyphlus; notes; characters; France).
▴ Distribution: France.
lucensis Coiffait, 1962d: 183 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: le Luc, sur la route du Lac aux Mayons au bord septentrional des Maures).
— Coiffait, 1972: 567 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
luci Hervé, 1972: 92 (Leptotyphlus; Type locality: ravin secondaire à 2 km au nord de Châteauvert, Var).
magrinii Pace, 1995: 28 (Leptotyphlus; subgenus Anomotyphlus; Type locality: Sardegna: Dorgali, prov. di Nuoro).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 320 (Leptotyphlus; subgenus Anomotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
mandibularis Coiffait, 1957: 73 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Tunisie: Tamera).
— Coiffait, 1959: 340 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
— Coiffait, 1972: 563 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
maritimus Coiffait, 1957: 73 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes Maritimes: Eze-sur-Mer).
— Coiffait, 1959: 336 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 556 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
martellii Pace, 1974: 503 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Toscana-Arezzo).
— Pace, 1977: 200 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 193 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
massatensis Coiffait, 1959: 412 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariége: entre Massat et le col des Caougnous).
— Coiffait, 1972: 523 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
maurettensis Hervé, 1968: 79 (Leptotyphlus; Type locality: ravin à l’hubac du Fenouillet, massif des Maurettes, commune de La Crau, Var).
— Coiffait, 1972: 571 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
maureillensis Coiffait, 1962d: 174 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Maureillas, route de Las Illas près d’un suintement).
— Coiffait, 1972: 542 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
mauryensis Coiffait, 1962d: 174 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Maury, près d’une petite source au sud du bourg).
— Coiffait, 1972: 513 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
mayeti Normand, 1907, see: Entomoculia.
mendicus Normand, 1939, see: Entomoculia.
meriensis Coiffait, 1959: 414 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Meria).
— Coiffait, 1972: 553 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 132 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 172 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
micros Scheerpeltz, 1931, see: Gynotyphlus.
mimus Pace, 1977: 188 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Monti Lepini, Lazio, a circa 700 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 235 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
minervius Coiffait, 1959: 414 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Monte Minervio).
— Coiffait, 1972: 553 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 130 (Leptotyphlus; synonym of devillei).
— Pace, 1996: 179 (Leptotyphlus; subgenus Leptotyphlus; valid species; characters; Corsica).
▴ Distribution: Corsica.
minos Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Catalogne: Alella).
— Coiffait, 1959: 372 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
— Coiffait, 1972: 583 (Leptotyphlus; subgenus Anomotyphlus; characters; Spain).
— Pace, 1995: 30, 31 (Leptotyphlus; subgenus Anomotyphlus; characters).
▴ Distribution: Spain.
molisensis Pace, 1977: 180 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: versante settentrionale della Montagna del Matese tra i 750 e gli 800 metri d’altitudine presso il centro abitato di S. Massimo, Campobasso).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 259 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
monachus Hervé, 1967b: 527 (Leptotyphlus; Type locality: forêt domainale de la Verne, ancien chemin de la Chartreuse, dans les Maures, Var).
— Coiffait, 1972: 579 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
muricatus Orousset, 1988a: 89 (Leptotyphlus; subgenus Odontotyphlus; Type locality: Corse-du-Sud: Zonza, 750 m).
— Pace, 1996: 267 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
mutilus Orousset, 1984, see: omessae Dodero, 1908.
necropolis Coiffait, 1962d: 177 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Villeneuve-des-Corbières, au pied d’un olivier près du cimetière).
— Coiffait, 1972: 549 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
nerviensis Coiffait, 1962d: 179 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Alpes-Maritimes italiennes: Province d’Imperia, Isolabona, Vallée de la Nervia, au nord de Vintimille).
— Coiffait, 1972: 540 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Pace, 1977b: 291 (Leptotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 210 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
nistosensis Coiffait, 1965: 138 (Leptotyphlus; Type locality: Hautes-Pyrénées, Bas Nistos).
— Coiffait, 1972: 530, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
nonzius Coiffait, 1959, see: devillei Coiffait, 1957.
normandi Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Ariège: Foix).
— Coiffait, 1959: 346 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 517 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
occidentalis Hervé, 1964, see: coiffaiti, Hervé, 1964.
ochsi Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Alpes-Mar: Vence).
— Coiffait, 1959: 371 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 551 (Leptotyphlus; subgenus Leptotyphlus; characters; Maritime Alps).
▴ Distribution: France.
oglasensis Poggi, 1976: 614 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Isola Montecristo, Arcipelago toscano, prov. Livorno, Collo dei Lecci, 500 m).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 195 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
olotensis Coiffait, 1959: 370 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Prov. de Gerona: Olot).
— Coiffait, 1972: 545 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
▴ Distribution: Spain.
omessae Dodero, 1908: 636, 639 (Leptotyphlus; Type locality: Corsica; Omessa; [Note: See lectotype designation by Orousset, 1984b]).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Coiffait, 1959: 348 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 581 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Orousset, 1984b: 192 (Leptotyphlus; subgenus Odontotyphlus; lectotype designation; characters; Corsica).
— Pace, 1996: 272 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
corsicus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Corse).
— Coiffait, 1959: 360 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 534 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1984b: 192 (Leptotyphlus; synonym of omessae).
kyrnosus Coiffait, 1957: 79 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: ? Omessa).
— Coiffait, 1959: 374 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 581 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
— Orousset, 1984b: 192 (Leptotyphlus; synonym of omessae).
mutilus Orousset, 1984b: 194 (Leptotyphlus; subgenus Odontotyphlus; subspecies of omessae; Type locality: Grotte Sapara, près de Castiglione, Haute-Corse, alt. 525 m).
— Pace, 1996: 272 (Leptotyphlus; subgenus Odontotyphlus; subspecies of omessae; characters; Corsica).
orbensis Coiffait, 1959: 418 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: St-Antonin [Note: See Orousset, 1985, for correction of type locality]).
— Coiffait, 1972: 551 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1985: 177 (Leptotyphlus; correction of type locality: Saint Antoine).
— Pace, 1996: 191 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
orcus Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Tarn: Castres).
— Coiffait, 1959: 346 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 517 (Leptotyphlus; subgenus Leptotyphlus; characters).
▴ Distribution: France.
orientalis Coiffait, 1954, see: Eotyphlus.
orion Normand, 1939, see: Entomoculia.
orpheus Coiffait, 1957: 75 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyr. Orientales: Le Perthus).
— Coiffait, 1959: 352 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 525 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
osellai Pace, 1973: 318 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Prealpi Bresciane nel Comune di Vallio).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 202 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
pauxillus Normand, 1939, see: Entomoculia.
peillensis Coiffait, 1957: 74 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Alpes Maritimes: Peille, dans la vallée du Paillon).
— Coiffait, 1959: 346 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 515 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Pace, 1996: 208 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
perpensus Normand, 1939, see: Entomoculia.
perpusillus Dodero, 1900, see: Gynotyphlus.
perroti Coiffait, 1957: 78 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Hérault: St-Pons-de-Mauchiens).
— Coiffait, 1959: 372 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 551 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
pertusatus Coiffait, 1959: 418 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: près du phare de Pertusato, Bonifacio).
— Coiffait, 1972: 530, 536 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 191 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
petretensis Coiffait, 1959: 418 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Petreto).
— Coiffait, 1972: 549 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 188 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
phami Orousset, 1987c: 134 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Sainte-Lucie de Tallano, alt. 350 m, Corse-du-Sud).
— Pace, 1996: 173 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
phoenix Coiffait, 1962d: 180 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Cavalaire, le Dattier).
— Hervé, 1964: 294 (Leptotyphlus; notes; France).
— Hervé, 1967c: 230, 232 (Leptotyphlus; characters).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
picardi Lavagne, 1917: 138 (Leptotyphlus; variety of lavagnei; Type locality: Castelnau, Agde, Marsillargues, Colombières-sur-Orb, Avèze).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; variety of lavagnei).
— Coiffait, 1959: 364 (Leptotyphlus; subgenus Leptotyphlus; cited as valid species; characters; France).
— Coiffait, 1972: 54 (Leptotyphlus; subgenus Leptotyphlus; characters).
▴ Distribution: France.
picenus Pace, 1977: 176 (Leptotyphlus; subgenus Subhesperotyphlus; Type locality: Appennino Marchigiano tra i 350 e i 500 metri d’altitudine, ai piedi dei monti situati presso il corso superiore del fiume Cesano, presso le località Bellisio Solfare e Fonte Avellana, ai piedi del Monte Catria).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 264 (Leptotyphlus; subgenus Odontotyphlus; characters; Italy).
▴ Distribution: Italy.
pluto Coiffait, 1955a: 69 (Leptotyphlus; Type locality: Alpes-Mar. Vaugrenier).
— Coiffait, 1959: 343 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 512 (Leptotyphlus; subgenus Leptotyphlus; characters; Maritime Alps).
▴ Distribution: France.
poirioni Hervé, 1971a: 206 (Leptotyphlus; Type locality: France: Var: Esterel, vallon descendant du col de l’Evêque vers la mer, entre l’hubac du Cap Roux et l’adret du Pic d’Aurelle).
pontelecciensis Orousset, 1987c: 134 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ponte-Leccia, route de Piedicroce, alt. 350 m, Haute-Corse).
— Pace, 1996: 183 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
priapus Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Var: Fréjus).
— Coiffait, 1959: 374 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 567 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
contractus Hervé, 1978: 139 (Leptotyphlus; subspecies of priapus; Type locality: Vauloube, dans le haut vallon du Reyran sur le territoire de Bagnols-en-Forêt, Var).
proprianensis Coiffait, 1959: 420 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Propriano).
— Coiffait, 1972: 547 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Pace, 1996: 189 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
pyrenaeus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Ariège: Salsein).
— Coiffait, 1959: 356 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 530, 536 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
queinneci Orousset, 1984: 302 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Croce, nord-est de Petreto, chênai verte, alt. 750 m, Corse du sud).
— Pace, 1996: 223 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
remensis Pace, 1977b: 288 (Leptotyphlus; Type locality: San Remo, presso Vignai, nel vallone del torrente Oxentina, a circa 500 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 211 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
reticulatus Coiffait, 1959: 411 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Maritimes: le vallon de Laghet à une dizaine de kilomètres au Nord Ouest de Nice).
— Coiffait, 1972: 561 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
revelierei Saulcy, 1878, see: Cephalotyphlus.
revelierianus Orousset, 1988b: 331 (Leptotyphlus; subgenus Odontotyphlus; Type locality: Corse-du-Sud: Ponte de Criviscia, commune de Zonza, alt. 650 m).
— Pace, 1996: 227 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
rhadamanthus Coiffait, 1957: 78 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Cendo).
— Coiffait, 1959: 370 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 549 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 130 (Leptotyphlus; subgenus Leptotyphlus; notes; Corsica).
— Pace, 1996: 219 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
ribasensis Coiffait, 1957: 75 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Espagne: Prov. de Gérona: Ribas de Freser).
— Coiffait, 1959: 348 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
— Coiffait, 1972: 521 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
▴ Distribution: Spain.
rivularis Coiffait, 1962d: 179 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Las Illas, au bord d’un petit torrent que franchit la route de Maureillas à Las Illas).
— Coiffait, 1972: 551 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
romanus Pace, 1975: 150 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: provincia di Roma, nel comune di Carpineto Romano, sui Monti Lepini tra i 560 e i 700 metri d’altitudine).
— Pace, 1977: 198 (Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 259 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
ruffoi Pace, 1975: 161 (Leptotyphlus; subgenus Anomotyphlus; Type locality: Calabria settentrionale, presso il paese di Grisolia a circa 550 metri d’altitudine ai piedi del Monte Carpinoso, presso il torrente Vaccuta).
— Pace, 1977: 198 (Leptotyphlus; characters; Italy).
— Pace, 1995: 30, 31 (Leptotyphlus; subgenus Anomotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 275 (Leptotyphlus; subgenus Anomotyphlus; characters; Italy).
▴ Distribution: Italy.
samniticus Pace, 1977: 192 (Leptotyphlus; Type locality: versante settentrionale della Montagna del Matese, presso il centro abitato di S. Massimo, intorno i 710 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 236 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
sardiniensis Herman, 2001: 36 (Leptotyphlus; replacement name for doderoi Coiffait).
doderoi Coiffait, 1957: 77 (Leptotyphlus; subgenus Leptotyphlus; [preoccupied]; Type locality: Sardaigne: Seui).
— Coiffait, 1959: 364 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Coiffait, 1972: 540 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 238 (Leptotyphlus; subgenus Subhesperotyphlus; characters; Sardinia).
— Herman, 2001: 36 (Leptotyphlus; synonym of sardiniensis).
sardous Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Sardaigne: Banari).
— Coiffait, 1959: 350 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Coiffait, 1972: 521 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 223 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
sassii Hervé, 1966a: 335 (Leptotyphlus; Type locality: France: La Garde-Freinet, près du col de Vignon dans les Maures, Var).
— Hervé, 1967c: 230 (Leptotyphlus; characters).
— Coiffait, 1972: 567 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
saturnius Normand, 1939, see: Entomoculia.
saulcyi Croissandeau, 1891: 150 (Leptotyphlus; Type locality: Corse, Omessa).
savonensis Pace, 1977b: 294 (Leptotyphlus; Type locality: presso il villaggio di Berioli, prov. di Savona, a circa 500 metri d’altitudine).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 200 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
sbordonii Pace, 1991: 110 (Leptotyphlus; Type locality: Italia: Lazio: Isola di Ponza).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 202 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
schuleri Hervé, 1967: 245 (Leptotyphlus; Type locality: Cogolin, Var, hubac du ravin de la rivière de Grimaud à l’ouest de la ferme de l’Amirauté).
— Hervé, 1967c: 230 (Leptotyphlus; characters).
— Coiffait, 1972: 569 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
senensis Pace, 1979b: 432 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Montagnola Senese, sul versante settentrionale del Monte Maggio, a circa 400 metri d’altitudine).
— Pace, 1980e: 161 (Leptotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 168 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
senyensis Coiffait, 1957: 77 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Catalogne: Figaro, au pied de la Sierra de Montseny).
— Coiffait, 1959: 364 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
— Coiffait, 1972: 540 (Leptotyphlus; subgenus Leptotyphlus; characters; Spain).
▴ Distribution: Spain.
serianensis Coiffait, 1957: 76 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Italie: Piario, dans le Val Seriana).
— Coiffait, 1959: 354 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Coiffait, 1972: 527 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 201 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
siccanus Normand, 1939, see: Entomoculia.
simplarius Normand, 1939, see: Entomoculia.
solarii Coiffait, 1957: 80 (Leptotyphlus; subgenus Hesperotyphlus; Type locality: Sardaigne: Aritzo).
— Coiffait, 1959: 380 (Leptotyphlus; subgenus Hesperotyphlus; characters; Sardinia).
— Coiffait, 1972: 510 (Leptotyphlus; subgenus Subhesperotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 238 (Leptotyphlus; subgenus Subhesperotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
solitarius Hervé, 1964, see: coiffaiti Hervé, 1964.
sospelensis Coiffait, 1957: 73 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes Maritimes: Sospel, Mt. Razet).
— Coiffait, 1959: 338 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 559 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
sourniensis Coiffait, 1962d: 173 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Pyrénées-Orientales: Sournia).
— Coiffait, 1972: 513 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
strobinoi Hervé, 1967c: 226 (Leptotyphlus; Type locality: hubac du massif des Pradels, commune de la Môle, Var).
— Coiffait, 1972: 573 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
subhervei Coiffait, 1959, see: hervei Jarrige, 1949.
sublaevis Fauvel, 1874: 330 [= 1874b: 36] (Leptotyphlus; Type locality: Omessa, Corse; [Note: See lectotype designation by Orousset, 1985]. Originally cited from Bastia, Corte, Omessa; ? cap Corse).
— Saulcy, 1878: 119 (Leptotyphlus; characters; Corsica).
— Sainte-Claire Deville, 1906: 78 (Leptotyphlus; Corsica).
— Bernhauer, 1908e: 332 (Leptotyphlus; characters).
— Dodero, 1908: 635, 638 (Leptotyphlus; characters; Italy).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Portevin, 1929: 392 (Leptotyphlus; characters; France).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1959: 370 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Coiffait, 1972: 547 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1985: 174 (Leptotyphlus; lectotype designation: Omessa, Corse; characters; Corsica).
— Pace, 1996: 212 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica, France, Italy.
sublaevis: Mulsant and Rey, 1878, see: exilis Mulsant and Rey, 1875.
sublaneyriei Hervé, 1965: 88 (Leptotyphlus; Type locality: France: Beauvallon, entre Sainte-Maxime et Grimaud, Var).
— Coiffait, 1972: 577 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
▴ Distribution: France.
subpyrenaeus Coiffait, 1959: 351 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Ariège: Aulus).
— Coiffait, 1972: 523, 534 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
sucellus Coiffait, 1955a: 69 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: S. Alpes-Mar. Les Veyans).
— Coiffait, 1959: 339 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 561 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
sylvanus Coiffait, 1955a: 68 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Var: St-Cassien-des-Bois).
— Coiffait, 1959: 336 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
— Coiffait, 1972: 557 (Leptotyphlus; subgenus Stigmotyphlus; characters; France).
▴ Distribution: France.
taggiensis Coiffait, 1962d: 170 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Maritimes italiennes: Taggia, ravine d’Ossentina).
— Coiffait, 1972: 557 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 246 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
tellus Coiffait, 1955, see: bonadonai Coiffait, 1955.
thaumasios Orousset, 1987c: 132 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Poggio, commune de Barbaggio, alt. 325 m, Haute-Corse).
— Pace, 1996: 176 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
transensis Coiffait, 1959: 376 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: Trans-en-Provence).
— Coiffait, 1972: 569 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Hervé, 1972: 89 (Leptotyphlus; characters; France).
▴ Distribution: France.
tressensi Coiffait, 1965: 138 (Leptotyphlus; Type locality: France: Bouche-du-Rhône, Gémenos, Parc de St-Pons).
— Coiffait, 1972: 581 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Hervé, 1978: 134 (Leptotyphlus; notes; characters; France).
▴ Distribution: France.
trifurcatus Coiffait, 1959: 424 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Var: sur le versant Nord de la montagne de la Loube à Brignoles).
— Coiffait, 1972: 579 (Leptotyphlus; subgenus Odontotyphlus; characters; France).
— Hervé, 1978: 134 (Leptotyphlus; notes; characters; France).
— Thelot, 1979: 20 (Leptotyphlus; subgenus Odontotyphlus; notes).
▴ Distribution: France.
troglophilus Coiffait, 1957: 75 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Hautes-Pyr.: Sarrancolin, à l’entrée de la grotte d’Ilhet).
— Coiffait, 1959: 353 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 525 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
truggiensis Orousset, 1984b: 190 (Leptotyphlus; subgenus Odontotyphlus; Type locality: Pont de Truggia, vallée de Liamone, Corse-du-sud, alt. 60 m).
— Orousset, 1988a: 91 (Leptotyphlus; subgenus Odontotyphlus; Corsica).
— Pace, 1996: 270 (Leptotyphlus; subgenus Odontotyphlus; characters; Corsica).
▴ Distribution: Corsica.
tuniseus Normand, 1910: 87 (Leptotyphlus; Type locality: Ain Draham et au Kef, Tunisia).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1959: 340 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
— Coiffait, 1972: 563 (Leptotyphlus; subgenus Stigmotyphlus; characters; Tunisia).
▴ Distribution: Tunisia.
tyrrhenius Dodero, 1908: 638 (Leptotyphlus; Type locality: Monte Argentaro, Isola de Giglio e dintorni di Grosseto, tutte nella Toscana meridionale).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1959: 368 (Leptotyphlus; subgenus Leptotyphlus; cited as tyrrhenicus; characters; Italy).
— Coiffait, 1972: 545 (Leptotyphlus; subgenus Leptotyphlus; cited as tyrrhenicus; characters; Italy).
— Pace, 1977: 199 (Leptotyphlus; characters; cited as tyrrhenicus; Italy).
— Pace, 1979b: 434 (Leptotyphlus; characters).
— Pace, 1980e: 161 (Leptotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 174 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
uccellinensis Pace, 1979: 179 (Leptotyphlus; Type locality: Monti dell’Uccellina, Poggio Lecci, presso S. Rabano, 280 m).
— Pace, 1980e: 161 (Leptotyphlus; characters).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 170 (Leptotyphlus; subgenus Leptotyphlus; characters; Italy).
▴ Distribution: Italy.
vandeli Coiffait, 1959: 354 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Kercabanac au confluent de l’Arac et du Salat).
— Coiffait, 1972: 527, 532 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
▴ Distribution: France.
varagensis Thelot, 1980: 50 (Leptotyphlus; subgenus Leptotyphlus; Type locality: France: berge du Grand Vallat, commune de Varages, Var).
velatus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Sardaigne: Campeda).
— Coiffait, 1959: 354 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Coiffait, 1972: 527 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 207 (Leptotyphlus; subgenus Leptotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
veneriatus Normand, 1939, see: Entomoculia.
ventralis Coiffait, 1962d: 171 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Alpes-Maritimes, La Colle sur Loup).
— Coiffait, 1972: 561 (Leptotyphlus; subgenus Stigmotyphlus; characters; Maritime Alps).
▴ Distribution: France.
venus Coiffait, 1955a: 70 (Leptotyphlus; Type locality: Pyr.-Orientales: Port-Vendres).
— Coiffait, 1959: 352 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1962d: 177 (Leptotyphlus; subgenus Leptotyphlus; characters; France).
— Coiffait, 1972: 525 (Leptotyphlus; subgenus Leptotyphlus; characters; Pyrenees).
▴ Distribution: France.
veronensis Pace, 1978a: 194 (Leptotyphlus; subgenus Stigmotyphlus; Type locality: Grezzana, VR).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 254 (Leptotyphlus; subgenus Stigmotyphlus; characters; Italy).
▴ Distribution: Italy.
vicus Coiffait, 1959: 416 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Vico).
— Coiffait, 1972: 553 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
— Orousset, 1987c: 136 (Leptotyphlus; subgenus Leptotyphlus; notes; Corsica).
— Pace, 1996: 177 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
vignesi Thelot, 1975, see: dispersus Hervé, 1966.
villiersi Orousset, 1984e: 64 (Leptotyphlus; subgenus Leptotyphlus; Type locality: Corse: Omessa).
— Pace, 1996: 226 (Leptotyphlus; subgenus Leptotyphlus; characters; Corsica).
▴ Distribution: Corsica.
viti Pace, 1978b: 189 (Leptotyphlus; Type locality: dintorni di Sassari).
— Pace, 1991: 110 (Leptotyphlus; subgenus Porotyphlus).
— Zanetti, 1995: 15 (Leptotyphlus; Italy).
— Pace, 1996: 284 (Leptotyphlus; subgenus Porotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
vivesi Coiffait, 1976c: 55 (Leptotyphlus; subgenus Subhesperotyphlus; Type locality: Prov. de Gerona, Olot, Santigosa).
vulcanus Coiffait, 1955, see: hervei Jarrige, 1949.
zariquieyi Coiffait, 1955a: 71 (Leptotyphlus; Type locality: Prov. de Gérona: Cadaques).
Newtonius
[1 species; Australian Region]
Newtonius Naomi, 1996c: 235 (species included: placidus). Type species: Newtonius placidus Naomi, fixed by original designation and monotypy.
placidus Naomi, 1996c: 239 (Newtonius; Type locality: Indonesia: Irian Jaya, Jayapura).
Portotyphlus
[1 species; Palaearctic Region]
Portotyphlus Coiffait, 1979b: 315 (species included: ramiroi). Type species: Portotyphlus ramiroi Coiffait, fixed by original designation.
ramiroi Coiffait, 1979b: 315 (Portotyphlus; Type locality: Porto “jardin de Signora M. de Silva”).
Sekotyphlus
[1 species; Ethiopian Region]
Sekotyphlus Coiffait, 1967b: 4 (species included: alberti). Type species: Sekotyphlus alberti Coiffait, fixed by original designation and monotypy.
alberti Coiffait, 1967b: 4 (Sekotyphlus; Type locality: Congo ex-belge, Parc Albert, secteur Tshiaberimu, mont Musimba, 2450 m, près de la rivière Musabaki, affl. Talya Nord).
Tribe Metrotyphlini
Metrotyphlini Coiffait, 1963d: 381 (genera included: Metrotyphlus). Type genus: Metrotyphlus (see Newton and Thayer, 1992: 57).
— Coiffait, 1972: 383, 608 (characters; key to genera and species of western Palaearctic region).
— Saiz, 1973b: 288 (distribution).
— Frank and Thomas, 1984a: 1413 (checklist of New World taxa).
— Newton, 1985: 206 (notes on austral distribution).
— Newton and Thayer, 1992: 57 (tribe of Leptotyphlinae).
— Pace, 1996: 59, 60, 285 (characters; key to genera).
Apotyphlus
[2 species; Neotropical Region]
Apotyphlus Coiffait and Saiz, 1965a: 129 (species included: relictus). Type species: Apotyphlus relictus Coiffait and Saiz, fixed by original designation and monotypy.
— Coiffait and Saiz, 1968: 346 (list of Chilean species).
— Coiffait and Decou, 1972: 136 (tribal assignment; notes).
— Decou, 1973: 360 (tribal assignment; notes).
— Saiz, 1974: 47, 48 (characters; key to species of Chile).
proximus Saiz, 1974: 48 (Apotyphlus; Type locality: Chile: Concón, Valpso., Cord. de la Costa).
relictus Coiffait and Saiz, 1965a: 131 (Apotyphlus; Type locality: Chili: Cordillera de la Costa, Maitencillo).
Banatotyphlus
[1 species; Palaearctic Region]
Banatotyphlus Decu, 1982: 63 (species included: racovitzai). Type species: Banatotyphlus racovitzai Decu, fixed by monotypy.
racovitzai Decu, 1982: 64 (Banatotyphlus; Type locality: vallée d’Ogasu Gåurii, Monts Locvei).
Egeotyphlus
[3 species; Palaearctic Region]
Egeotyphlus Coiffait, 1957: 71, 80 (species included: thracicus, cited as thoracicus). TYPE
SPECIES: Egeotyphlus thracicus Coiffait, fixed by original designation and monotypy.
— Coiffait, 1959: 388 (characters; type species: thracicus).
— Coiffait, 1972: 614 (characters; type species: thracicus; key to species of western Palaearctic region; references).
— Pace, 1983a: 460 (characters).
— Pace, 1984: 17 (key to species).
Epirotyphlus Pace, 1995a: 29 (subgenus of Egeotyphlus; species included: leucadiensis, zecevici). Type species: Egeotyphlus (Epirotyphlus) leucadiensis Pace, fixed by original designation.
leucadiensis Pace, 1995a: 27 (Egeotyphlus; subgenus Epirotyphlus; Type locality: Grecia: Isola di Leukas, Aplopena).
thracicus Coiffait, 1957: 81 (Egeotyphlus; Type locality: Thrace grecque: Makri).
— Coiffait, 1959: 389 (Egeotyphlus; characters; Greece).
— Coiffait, 1972: 614 (Egeotyphlus; characters; Greece).
— Pace, 1984: 17 (Egeotyphlus; characters).
▴ Distribution: Greece.
zecevici Pace, 1984: 17 (Egeotyphlus; Type locality: Serbia: Stol planina, 800 m).
Gynotyphlus
[3 species; Palaearctic Region]
Gynotyphlus Coiffait, 1955a: 72 (species included: perpusillus). Type species: Leptotyphlus perpusillus Dodero, fixed by original designation and monotypy.
— Coiffait, 1959: 390 (characters; key to species; type species: perpusillus).
— Coiffait, 1972: 617 (characters; type species: perpusillus; key to species of western Palaearctic region; references).
— Boháč, 1993: 44 (in list of species of Czech Republic and Slovakia).
— Zanetti, 1995: 16 (list of species of Italy).
— Pace, 1996: 302 (characters; key to species of Italy).
Oligotyphlus Scheerpeltz, 1955b: 460 (species included: perpusillus, ireneae, curtulus, micros, along with 17 unavailable names). Type species: Leptotyphlus perpusillus Dodero, fixed by original designation.
— Scheerpeltz, 1940: 28 ([published with characters but without species]).
— Coiffait, 1959: 390 (synomym of Gynotyphlus; type species: perpusillus).
— Coiffait, 1972: 617 (synonym of Gynotyphlus).
— Pace, 1996: 302 (synonym of Gynotyphlus).
Austriacotyphlus Scheerpeltz, 1959: 4 (species included: piffli). Type species: Austriacotyphlus piffli Scheerpeltz, fixed by original designation and monotypy.
— Horion, 1963: 296 (list of central European species).
— Székessy, 1963: 34 (characters).
— Lohse, 1964: 107 (characters; key to central European species).
— Scheerpeltz, 1968a: 34 (catalog of Austrian species).
— Coiffait, 1972: 616 (characters; type species: piffli; key to species of western Palaearctic region; references).
— Tikhomirova, 1973a: 171 (checklist of species of USSR).
— Pace, 1983b: 44 (synonym of Gynotyphlus).
— Lohse and Lucht, 1989: 139 (synonym of Gynotyphlus).
— Pace, 1996: 302 (synonym of Gynotyphlus).
beierianus Scheerpeltz, 1958c: 396 (Oligotyphlus; Type locality: im südlichen Epirus in der Umgebung von Nisista, Xerovuni, in etwa 700 bis 800 m Seehöhe).
corcyrensis Scheerpeltz, 1958a: 394 (Oligotyphlus; Type locality: Val de Ropa auf Korfu).
curtulus Jarrige, 1949, see: perpusillus Dodero, 1900.
perpusillus Dodero, 1900a: 573 (Leptotyphlus; Type locality: Genova).
— Dodero, 1908: 635, 638 (Leptotyphlus; characters; Italy).
— Bernhauer and Schubert, 1911: 147 (Leptotyphlus; catalog).
— Coiffait, 1955a: 72 (Gynotyphlus).
— Coiffait, 1957: 81 (Gynotyphlus; subspecies).
— Coiffait, 1959: 392 (Gynotyphlus; key to races; characters; Italy).
— Besucht and Comellini, 1964: 313 (Gynotyphlus; Switzerland).
— Coiffait, 1965b: 696 (Gynotyphlus; notes).
— Coiffait, 1972: 618 (Gynotyphlus; characters; key to subspecies; Italy).
— Pace, 1983b: 46 (Gynotyphlus; characters).
— Coiffait, 1984a: 117 (Gynotyphlus; probably parthenogenetic; France; Turkey; Mediterranean region; Corsica; Sardinia; Sicily; Morocco; Portugal; Austria).
— Zanetti, 1995: 16 (Gynotyphlus; Italy).
— Pace, 1996: 304 (Gynotyphlus; characters; Italy).
▴ Distribution: Portugal, France, Corsica, Italy, Sardinia, Sicily, Switzerland, Austria, Greece, Ionian Islands, Croatia, Yugoslavia, Turkey, Morocco.
micros Scheerpeltz, 1931: 399 (Leptotyphlus; Type locality: Tale Perivolakia bei Kaligoni auf Levkas).
— Scheerpeltz, 1933: 1138 (Leptotyphlus; catalog).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; Greece).
— Coiffait, 1968b: 93 (Gynotyphlus; race of perpusillus; notes; Greece).
— Coiffait, 1972: 620 (Gynotyphlus; subspecies of perpusillus; characters; Greece).
— Coiffait, 1973b: 223 (Gynotyphlus; subspecies of perpusillus; Greece).
curtulus Jarrige, 1949: 62 (Leptotyphlus; Type locality: Pont des Veyens).
— Scheerpeltz, 1955b: 460 (Oligotyphlus).
— Coiffait, 1959: 392 (Gynotyphlus; race of perpusillus; characters; France).
— Coiffait, 1972: 619 (Gynotyphlus; subspecies of perpusillus; characters; France).
— Pace, 1983b: 46 (Gynotyphlus; subspecies of perpusillus; characters).
rhodanicus Coiffait, 1955a: 72 (Gynotyphlus; subspecies of perpusillus; Type locality: Lyon).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; France).
— Coiffait, 1972: 619 (Gynotyphlus; subspecies of perpusillus; characters; France; Portugal; Switzerland).
garumnicus Coiffait, 1955a: 72 (Gynotyphlus; subspecies of perpusillus; Type locality: Vallée de la Garonne de Toulouse à Bordeaux).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; France).
— Coiffait, 1972: 619 (Gynotyphlus; subspecies of perpusillus; characters; France).
dalmaticus Coiffait, 1955a: 72 (Gynotyphlus; subspecies of perpusillus; Type locality: Dalmatie: Susurac).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; Dalmatia).
— Coiffait, 1972: 619 (Gynotyphlus; subspecies of perpusillus; characters; Dalmatia).
macedonicus Coiffait, 1957: 81 (Gynotyphlus; subspecies of perpusillus; Type locality: Macédoine Yougoslave: Katlanovo).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; Macedonian Yugoslavia).
— Coiffait, 1972: 620 (Gynotyphlus; subspecies of perpusillus; characters; Macedonia).
turcicus Coiffait, 1957: 81 (Gynotyphlus; subspecies of perpusillus; Type locality: Turquie d’Asie: Karamursel sur la rive sud de la mer de Marmara).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; Turkey).
— Coiffait, 1972: 620 (Gynotyphlus; subspecies of perpusillus; characters; Turkey).
— Coiffait, 1973b: 222 (Gynotyphlus; subspecies of perpusillus; Turkey).
meridionalis Coiffait, 1957: 81 (Gynotyphlus; subspecies of perpusillus; Type locality: Italie du Sud: Vallo Lucano, aux confins de la Calabre).
— Coiffait, 1959: 393 (Gynotyphlus; race of perpusillus; characters; Italy).
— Coiffait, 1972: 620 (Gynotyphlus; subspecies of perpusillus; characters; Italy).
— Zanetti, 1995: 16 (Gynotyphlus; subspecies of perpusillus; Italy).
— Pace, 1996: 304, 306 (Gynotyphlus; subspecies of perpusillus; characters; Italy).
piffli Scheerpeltz, 1959: 9 (Austriacotyphlus; Type locality: Südhang des Leopoldsberges, Wienerwald, nordwestlich von Wien).
— Horion, 1963: 296 (Austriacotyphlus; Austria).
— Lohse, 1964: 107 (Austriacotyphlus; characters; central Europe).
— Coiffait, 1972: 616 (Austriacotyphlus; characters; Austria).
— Pace, 1983b: 46 (Gynotyphlus; subspecies of perpusillus; characters).
— Lucht, 1987: 95 (Austriacotyphlus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).
— Lohse and Lucht, 1989: 139 (Gynotyphlus; subspecies of perpusillus).
— Boháč, 1993: 44 (Gynotyphlus; Slovakia ?; Czech Republic ?).
sequanus Miré, 1983: 248 (Gynotyphlus; subspecies of perpusillus; Type locality: Port-villez).
dalmaticus Coiffait, 1955, see: perpusillus Dodero, 1900.
garumnicus Coiffait, 1955, see: perpusillus Dodero, 1900.
macedonicus Coiffait, 1957, see: perpusillus Dodero, 1900.
meridionalis Coiffait, 1957, see: perpusillus Dodero, 1900.
micros Scheerpeltz, 1931, see: perpusillus Dodero, 1900.
piffli Scheerpeltz, 1959, see: perpusillus Dodero, 1900.
rhodanicus Coiffait, 1955, see: perpusillus Dodero, 1900.
sequanus Miré, 1983, see: perpusillus Dodero, 1900.
turcicus Coiffait, 1957, see: perpusillus Dodero, 1900.
Metrotyphlus
[12 species; Palaearctic Region]
Metrotyphlus Coiffait, 1959: 327, 328 (subgenus of Megatyphlus; species included: hervei, serianensis). Type species: Megatyphlus hervei Coiffait, fixed by original designation.
— Coiffait, 1972: 609 (characters; type species: hervei; key to species of western Palaearctic region; references).
— Pace, 1977a: 13 (key to species).
— Zanetti, 1995: 15 (list of species of Italy).
— Pace, 1996: 286 (characters; key to species of Italy).
Agnotyphlus Coiffait, 1972: 609, 611, 613 (subgenus of Metrotyphlus; species included: istriensis). Type species: Metrotyphlus (Agnotyphlus) istriensis Coiffait, fixed by original designation and monotypy.
besucheti Pace, 1976, see: serianensis Coiffait, 1957.
brixianus Pace, 1973: 320 (Metrotyphlus; subgenus Agnotyphlus; Type locality: dintorni di Vallio, in provincia di Brescia).
— Pace, 1976a: 682 (Metrotyphlus; characters).
— Pace, 1976: 111 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Zanetti, 1995: 15 (Metrotyphlus; Italy).
— Pace, 1996: 294 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
castrii Coiffait, 1965: 140 (Metrotyphlus; Type locality: Italie: Province de Vicenza, Recoaro).
— Coiffait, 1972: 612 (Metrotyphlus; characters; Italy).
— Pace, 1973: 323 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
— Pace, 1976: 111 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 297 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
esteronensis Coiffait, 1959: 410 (Megatyphlus; subgenus Metrotyphlus; Type locality: Alpes-Maritimes: dans la basse vallée de l’Estéron à Gilette).
goritianus Pace, 1977a: 8 (Metrotyphlus; subgenus Agnotyphlus; Type locality: monti del Carso di Monfalcone, presso il centro abitato di Devetachi, a circa 215 metri d’altitudine).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 299 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
hervei Coiffait, 1957: 71 (Megatyphlus; Type locality: Alpes Mar.: Eze-sur-Mer).
— Coiffait, 1959: 330 (Megatyphlus; subgenus Metrotyphlus; characters; France).
— Coiffait, 1972: 612 (Metrotyphlus; characters; Maritime Alps).
▴ Distribution: France.
istriensis Coiffait, 1972: 613 (Metrotyphlus; subgenus Agnotyphlus; Type locality: Istrie, Monte Maggiore).
— Pace, 1976: 111 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Pace, 1996: 299 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Croatia.
pauper Pace, 1976: 108 (Metrotyphlus; subgenus Agnotyphlus; Type locality: sul versante settentrionale del Monte Jof a circa 410 metri, presso Maniago; sul versante settentrionale del Col Cravest a circa 250 m, presso Travesio).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 300 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
sbordonii Pace, 1979e: 107 (Metrotyphlus; subgenus Metrotyphlus; Type locality: Sardegna: presso Laconi, Nuoro, nel parco Aymerich).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 288 (Metrotyphlus; subgenus Metrotyphlus; characters; Sardinia).
▴ Distribution: Sardinia.
serianensis Coiffait, 1957: 71 (Megatyphlus; Type locality: Italie: Piario dans le Val Seriana).
— Coiffait, 1959: 330 (Megatyphlus; subgenus Metrotyphlus; characters; Italy).
— Coiffait, 1972: 612 (Metrotyphlus; characters; Italy).
— Pace, 1976: 111 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 292 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy, Switzerland.
besucheti Pace, 1976a: 679 (Metrotyphlus; subgenus Agnotyphlus; Type locality: dintorni di Chiasso, Canto Ticino).
— Pace, 1976: 111 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
— Pace, 1977a: 13 (Metrotyphlus; subgenus Agnotyphlus; characters).
— Pace, 1996: 292 (Metrotyphlus; subgenus Agnotyphlus; subspecies of serianensis; characters; Switzerland).
tarvisianus Pace, 1991: 111 (Metrotyphlus; subgenus Agnotyphlus; Type locality: Italia: Veneto, soligo, boschi a N del centro abiatato).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 302 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
viti Pace, 1985: 62 (Metrotyphlus; Type locality: Liguria, Uscio, M. Fascie).
— Zanetti, 1995: 16 (Metrotyphlus; Italy).
— Pace, 1996: 295 (Metrotyphlus; subgenus Agnotyphlus; characters; Italy).
▴ Distribution: Italy.
zoiai Pace, 1991: 111 (Metrotyphlus; subgenus Agnotyphlus; Type locality: Italia: Toscana, Ostria, (dintorni di Varano, provincia di Massa C.), 240 m).
Rhopalotyphlus
[1 species; Palaearctic Region]
Rhopalotyphlus Scheerpeltz, 1955b: 462 (species included: beieri). Type species: Rhopalotyphlus beieri Scheerpeltz, fixed by original designation and monotypy.
beieri Scheerpeltz, 1931: 394 (Leptotyphlus; Type locality: Kaligoni auf Levkas).
Venezillotyphlus
[1 species; Neotropical Region]
Venezillotyphlus Decu, 1990: 37 (species included: pecki). Type species: Venezillotyphlus pecki Decu, fixed by original designation and monotypy.
pecki Decu, 1990: 38 (Venezillotyphlus; Type locality: Carupano playa, Estado Sucre, NV Venezuela).
Tribe Neotyphlini
Neotyphlini Coiffait, 1963d: 381, 382 (genera included: Megatyphlus, Prototyphlus, Chiliotyphlus; Coiffait indicated that other African and North and South American genera were included but failed to list them by name). Type genus: Neotyphlus (see Newton and Thayer, 1992: 57).
— Coiffait, 1964b: 16 (notes; distribution).
— Coiffait, 1972: 383, 622 (characters; key to genera and species of western Palaearctic region).
— Saiz, 1973b: 289 (distribution).
— Frank and Thomas, 1984a: 1412, 1413, 1414 (characters; checklist of New World taxa; comparative table of characters for New World genera).
— Newton, 1985: 206 (notes on austral distribution).
— Newton and Thayer, 1992: 57 (tribe of Leptotyphlinae).
— Pace, 1996: 59, 60, 306 (characters; key to genera).
— Pace, 1999: 80 (key to Palaearctic genera).
— Newton, Thayer, Ashe, and Chandler, 2000: 382 (characters; notes).
Apheliotyphlus
[1 species; Palaearctic Region]
Apheliotyphlus Pace, 1976: 105 (subgenus of Megatyphlus; species included: nobilis). Type species: Megatyphlus (Apheliotyphlus) nobilis Pace, fixed by original designation.
— Pace, 1979d: 472 (subgenus of Megatyphlus; key to species).
— Zanetti, 1995: 16 (list of species of Italy).
— Pace, 1996: 307 (valid genus; characters; species of Italy).
— Pace, 1999: 80 (characters).
nobilis Pace, 1976: 105 (Megatyphlus; subgenus Apheliotyphlus; Type locality: Friuli, presso Maniago, sul versante settentrionale del M. Jof a circa 410–450 m, presso il torrente Colvera di Jof).
Cafrotyphlus
[2 species; Ethiopian Region]
Cafrotyphlus Coiffait, 1964b: 13 (species included: leleupi). Type species: Cafrotyphlus leleupi Coiffait, fixed by original designation.
— Coiffait and Decou, 1972: 136 (tribal assignment; notes).
— Decou, 1973: 360 (tribal assignment; notes).
leleupi Coiffait, 1964b: 13 (Cafrotyphlus; Type locality: Natal: Ngomi forest, Nongoma district, 700 m).
strinatii Coiffait, 1977e: 659 (Cafrotyphlus; Type locality: Afrique du Sud: Montagne de la Table, Wynberg Cave).
Cainotyphlus
[2 species; Nearctic Region]
Cainotyphlus Coiffait, 1962: 156, 161 (species included: pacificus, walkeri). Type species: Cainotyphlus pacificus Coiffait, fixed by original designation.
— Coiffait and Decou, 1972: 136 (tribal assignment; notes).
— Decou, 1973: 360 (tribal assignment; notes).
— Moore and Legner, 1975: 265 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 250 (characters; notes).
— Newton, Thayer, Ashe, and Chandler, 2000: 383 (2 Nearctic species; notes; characters in key).
pacificus Coiffait, 1962: 162 (Cainotyphlus; Type locality: Corte Madera, Marin County, California).
walkeri Coiffait, 1962: 162 (Cainotyphlus; Type locality: Nelder Grove, Madera County, California).
Chiliotyphlus
[3 species; Neotropical Region]
Chiliotyphlus Coiffait, 1963d: 378 (species included: aculeensis, pacificus). Type species: Chiliotyphlus aculeensis Coiffait, fixed by original designation.
— Coiffait and Saiz, 1965a: 136 (characters).
— Coiffait and Saiz, 1968: 346 (list of Chilean species).
— Coiffait and Decou, 1972: 136 (tribal assignment; notes).
— Decou, 1973: 360 (tribal assignment; notes).
— Saiz, 1974: 47, 48 (characters; key to species of Chile).
aculeensis Coiffait, 1963d: 378 (Chiliotyphlus; Type locality: Chili: Cordillera de la Costa, Aculeo, 50 km environ au sud-sud-ouest de Santiago du Chili, 300 m).
pacificus Coiffait, 1963d: 378 (Chiliotyphlus; Type locality: Chili: Cordillera de la Costa, Zapallar, 60 km environ au nord de Valparaiso, 900 m).
spinosus Saiz, 1974: 48, 49 (Chiliotyphlus; Type locality: Chile: Casablanca, Valparaiso, Cordillera de la Costa).