Open Access
How to translate text using browser tools
18 July 2001 Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium.
VI. Staphylinine Group (Part 3)
Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini
Staphylinidae Incertae Sedis
Fossils, Protactinae
Lee H. Herman
Author Affiliations +
Abstract

This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies.

Subtribe Quediina

Quediina Kraatz, 1857.

  • — Jakobson, 1908: 447 (characters).

  • — Jakobson, 1909: 514 (key to genera; list of species of Russia and western Europe).

  • — Johansen, 1914: 341 (characters; key to Danish genera).

  • — Hatch, 1957: 172, 214 (characters; key to species of Pacific Northwest).

  • — Arnett, 1963: 279 (characters; list of North American genera).

  • — Newton and Thayer, 1992: 65 (subtribe of Staphylinini).

  • — Smetana, 1995e: 48 (characters).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (list of New Zealand genera; number of New Zealand species).

  • — Smetana and Davies, 2000: 12, 14 (characters).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 331 (key to Nearctic genera).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 331, 392 (key to Nearctic genera; characters; notes).

Platycnemidiformes Nordmann, 1837: 6 (genera included: Platycnemus). Type genus: Platycnemus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

  • — Newton, 1995: 48, 49, 51 (name emended to Platycnemini).

Platycnemini Nordmann, 1837.

  • — Newton, 1995: 51 (name emended from Platycnemidiformes; requested that the International Commission of Zoological Nomenclature Platycnemini not be given priority over junior family-group names in current general usage in the Staphylininae and that it be placed on the Official List of Family-Group Names in Zoology).

  • — ICZN, 1996: 215, 216 (Platycnemini placed on Official List of Family Group Names in Zoolgy; it and other family group names based on Platycnemus not given priority over junior family-group names in general current usage in the Staphylininae).

Quediiformes Kraatz, 1857d: 473 (genera included: Tanygnathus, Acylophorus, Euryporus, Heterothops, Quedius, Astrapaeus, Cordylaspis, Homorocerus, Haematodes, Scariphaeus, and with question Caranistes and Palaestrinus). Type genus: Quedius (see Newton and Thayer, 1992: 65).

Quediides Thomson, 1859: 25 (genera included: Velleius, Quedius, Microsaurus, Raphirus, Euryporus, Heterothops, Acylophorus, Tanygnathus). Type genus: Quedius (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Quedii Kraatz, 1857.

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1944: 143 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Seevers, 1965: 329 (myrmecophilus species).

  • — Coiffait, 1978: 7,9 (characters; cited as new subtribe).

Quediinae Kraatz, 1857.

  • — Casey, 1915: 396 (characters; discussion).

  • — Moore, 1964a: 89 (characters; notes).

  • — Moore and Legner, 1974g: 559 (key to genera of America north of Mexico).

  • — Moore and Legner, 1974h: 472 (catalog of the taxonomy, biology, and ecology of developmental stages of species of America north of Mexico).

  • — Moore and Legner, 1975: 48 (catalog; North America north of Mexico).

  • — Moore and Legner, 1979: 39 (characters; tabular key to genera of North America north of Mexico).

  • — E. Matthews, 1982: 7, 24 (note; key to genera of Australia).

  • — Lawrence and Newton, 1995: 830 (included in Staphylininae).

  • — Downie and Arnett, 1996: 387 (characters; key to genera of Northeastern North America).

Quediini Kraatz, 1857.

  • — Porta, 1907: 85 (characters; key to genera).

  • — Reitter, 1908: 101, 105, 106 (characters; key to genera).

  • — Eichelbaum, 1909: 184 (list of genera with references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

  • — Blatchley, 1910: 367 (characters).

  • — Casey, 1915: 397 (key to North American genera).

  • — Bernhauer and Schubert, 1916: 409 (world catalog).

  • — Porta, 1926: 108 (characters; species of Italy).

  • — Portevin, 1929: 334 (characters; key to French genera).

  • — Cameron, 1932: 2, 264 (characters; key to genera of British India).

  • — Joy, 1932: 106 (key to British genera).

  • — Scheerpeltz, 1933: 1418 (world catalog supplement).

  • — Scheerpeltz, 1940: 48 (key to Palaearctic genera).

  • — V. Hansen, 1952: 112 (characters; key to Danish genera).

  • — Smetana, 1958: 312 (characters; key to Czechoslovakian genera).

  • — Scheerpeltz, 1962b: 263 (key to Palaearctic genera).

  • — Palm, 1963b: 123 (characters; key to genera and species of Sweden).

  • — Lohse, 1964: 202 (characters; key to central European genera and species).

  • — Pototskaia, 1967: 75 (larval characters).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Kasule, 1970: 56 (larval characters; larval key to some genera).

  • — Saiz, 1971 (notes on Chilean taxa).

  • — Smetana, 1971 (revision of species and genera for America north of Mexico; key to genera and species; characters; collecting techniques).

  • — Smetana, 1977b: 178 (characters; notes on classification; key to genera of Holarctic region).

  • — Coiffait, 1978: 7 (characters; key to subtribes and genera of western Palaearctic region).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Smetana, 1981a: 354 (key to genera of Chile).

  • — Newton, 1985: 208 (notes on austral distribution and taxonomic notes).

  • — Outerelo and Gamarra, 1985: 61 (key to genera of Iberian Peninsula).

  • — Dettner, 1987: 30 (defensive glands).

  • — Smetana, 1988: 169, 178, 179 (characters of genital segments; characters; key to genera).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Gyrohypnini on the Official List of Family-Group Names in Zoology and that it and other family-group names based on Quedius be given precedence over those based on Platycnemus).

  • — Dettner, 1993a (summary and discussion of defensive secretions and exocrine glands and their bearing on phylogeny).

  • — Smetana, 1995: 14, 23 (characters; revision of species of Taiwan; key to genera and species of Taiwan; cladistic analysis).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature Quediini Kraatz be given precedence over Platycnemini Nordmann).

  • — Cho, 1996b: 108 (key to genera of Korea).

  • — ICZN, 1996: 216 (Quediini placed on Official List of Family Group Names in Zoology; it and other family group names based on Quedius have precedence over Platycnemini and other family group names based on Platycnemus).

Acylophorini Scheerpeltz, 1968a: 97 (catalog of Austrian species; genera included: Atangynathus). Type genus: Acylophorus (see Newton and Thayer, 1992: 65).

  • — Outerelo and Gamarra, 1985: 67 (genera of Iberian Peninsula).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Heterothopsi Coiffait, 1978: 7, 300 (genera included: Heterothops, Atanygnathus). Type genus: Heterothops (see Newton and Thayer, 1992: 65).

  • — Smetana, 1984c: 278 (characters; discussion; probably not valid).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Acylophorini Outerelo and Gamarra, 1985: 48, 67 (genera included: Acylophorus). Type genus: Acylophorus (see Newton and Thayer, 1992: 65).

  • — Newton and Thayer, 1992: 65 (synonym of Quediina).

Acylohsellus
[1 species; Oriental Region]

Acylohsellus Smetana, 1995d: 114 (species included: longiceps). Type species: Acylophorus longiceps Cameron, fixed by original designation and monotypy.

longiceps Cameron, 1918b: 172 (Acylophorus; Type locality: Borneo, Mt. Merinjak, Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1995d: 116 (Acylohsellus; characters; Borneo; Sumatra).

  • Distribution: Borneo, Indonesia.

Acylophorus [128 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, and Australian Regions]

Acylophorus Nordmann, 1837: 127 (species included: ahrensii, femoralis). Type species: Acylophorus ahrensii Nordmann, fixed by subsequent designation by Blackwelder, 1943: 466.

  • — Erichson, 1839a: 482 (characters).

  • — Erichson, 1840: 519 (characters).

  • — Heer, 1841: 584 (characters).

  • — Lacordaire, 1854: 84 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 534 (characters).

  • — Jacquelin du Val, 1856: 38 (characters).

  • — Kraatz, 1857d: 479 (characters).

  • — Redtenbacher, 1857: 199 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — Thomson, 1859: 26 (characters; type species: glabricollis).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 181 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Wencker and Silbermann, 1866: 27 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 566 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 316 [= 1874a: 542] (characters).

  • — Redtenbacher, 1874: 195 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 831 [= 1877: 687] (characters; France).

  • — Sharp, 1876: 97 (notes).

  • — LeConte, 1878a: 388 (key to North American species).

  • — Horn, 1878: 150 (characters; key to North American species).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1884: 324 (notes; Mexican and Central American species).

  • — Sharp, 1888: 222 (characters).

  • — Seidlitz, 1891: 400 (key to some European species).

  • — Seidlitz, 1891a: 427 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 98 (list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 380 (characters).

  • — Heyden, Reitter, and Weise, 1906: 177 (list of species of Europe and Caucasus).

  • — Porta, 1907: 88 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Blatchley, 1910: 368 (characters).

  • — Petri, 1912: 80 (list of species of Siebenbürgen).

  • — Johansen, 1914: 342 (characters; key to species).

  • — Casey, 1915: 426 (characters; key to species).

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 37 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Cameron, 1921a: 355, 405 (characters; key to Singapore species; catalog).

  • — Munster, 1923: 203 (key to Scandinavian species).

  • — Winkler, 1925: 393 (catalog for Palaearctic region).

  • — Cameron, 1925: 73 (catalog of Indian species).

  • — Porta, 1926: 119 (characters; species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 335 (characters; key to French species).

  • — Roubal, 1930: 397 (catalog; Slovakia).

  • — Cameron, 1932: 302 (characters; key to species of British India).

  • — Joy, 1932: 106 (key to British species).

  • — Scheerpeltz, 1933: 1466 (world catalog supplement).

  • — Normand, 1935: 373 (list of species of Tunisia).

  • — Bierig, 1938: 119–123 (characters; notes; illustration of variation of fourth segment of maxillary palpus and mandibles; type species).

  • — Blackwelder, 1943: 466 (characters; type species: ahrensii; key to species of West Indies).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 168 (list of British species).

  • — Tottenham, 1949: 377, 423 (type species: glaberrimus Herbst = ahrensii Nordmann; checklist of British species).

  • — Blackwelder, 1952: 38 (type species: ahrensii Nordmann).

  • — V. Hansen, 1952: 161 (characters; Danish species).

  • — Hatch, 1957: 231 (characters; species of Pacific Northwest).

  • — Kocher, 1958: 161 (checklist of species; Morocco).

  • — Smetana, 1958: 398 (characters; habitus of one species; type species: glaberrimus; key to Czechoslovakian species).

  • — Smetana, 1959a: 216 (list of species collected in Albania).

  • — Scheerpeltz, 1962b: 267 (characters).

  • — Székessy, 1963: 62 (characters).

  • — Palm, 1963b: 158 (characters; key to species of Sweden).

  • — Lohse, 1964: 220 (characters; key to central European species).

  • — Scheerpeltz, 1968a: 97 (catalog of Austrian species).

  • — Smetana, 1971: 246 (revision of and key to species of Canada and United States; characters).

  • — Shibata, 1973c: 64 (checklist of species of Taiwan).

  • — Tikhomirova, 1973a: 190 (checklist of species of USSR).

  • — Moore and Legner, 1975: 48 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 32 (list of British species).

  • — Coiffait, 1978: 296 (characters; key to species of western Palaearctic region).

  • — Moore and Legner, 1979: 41 (characters; notes).

  • — Muona, 1979: 16 (list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 154 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 126 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 204 (checklist of species of Borneo).

  • — Tóth, 1984: 93 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 139 (subgenus of Acylophorus; checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Smetana, 1988: 337 (characters; key to species of Himalayan region).

  • — Campbell and Davies, 1991: 122 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 18 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Smetana, 1993b: 50 (list of species of Czech Republic and Slovakia).

  • — Smetana, 1995: 125 (notes; revision of species of Taiwan; key to subgenera and species of Taiwan; cladistic analysis).

  • — Ciceroni and Zanetti, 1995: 33 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (additions to list of Danish species).

  • — Downie and Arnett, 1996: 394 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (18 Nearctic species; notes; characters in key).

Rhygmacera Motschulsky, 1845a: 40 (species included: nitida). Type species: Rhygmacera nitida Motschulsky, fixed by original designation and monotypy.

  • — Motschulsky, 1858: 656 (synonym of Acylophorus).

  • — Fauvel, 1874: 316 [= 1874a: 542] (synonym of Acylophorus).

  • — Ganglbauer, 1895: 380 (synonym of Acylophorus).

  • — Bernhauer and Schubert, 1916: 438 (synonym of Acylophorus).

  • — Cameron, 1932: 302 (synonym of Acylophorus).

  • — Blackwelder, 1943: 466 (synonym of Acylophorus; type species: nitida).

  • — Blackwelder, 1952: 341 (type species: nitida).

  • — Smetana, 1958: 398 (synonym of Acylophorus).

  • — Smetana, 1971: 246 (synonym of Acylophorus).

  • — Coiffait, 1978: 296 (synonym of Acylophorus).

Paracylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: schmidti). Type species: Acylophorus (Paracylophorus) schmidti Bierig, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 289 (subgenus of Acylophorus; type species: schmidti).

Neoacylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: oculifer). Type species: Acylophorus (Neoacylophorus) oculifer Bierig, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 258 (subgenus of Acylophorus; type species: oculifer).

Indoacylophorus Bierig, 1938: 123 (subgenus of Acylophorus; species included: microcephalus). Type species: Acylophorus (Indoacylophorus) microcephalus Cameron, by original designation and monotypy.

  • — Blackwelder, 1952: 199 (subgenus of Acylophorus; type species: microcephalus).

Amacylophorus Smetana, 1971: 247 (subgenus of Acylophorus; species included: pratensis). Type species: Acylophorus (Amacylophorus) pratensis LeConte, fixed by original designation and monotypy.

Palpacylophorus Smetana, 1971: 247, 268 (subgenus of Acylophorus; species included: leechi). Type species: Acylophorus (Palpacylophorus) leechi Smetana, fixed by original designation and monotypy.

acuminatus Sharp, 1876: 99 (Acylophorus; Type locality: Ega).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

aenescens Blackwelder, 1943: 468 (Acylophorus; Type locality: Dominican Republic: Sanchez).

  • Distribution: West Indies.

agilis Smetana, 1971: 262 (Acylophorus; subgenus Acylophorus; Type locality: Indiana: Parke Co., Inc. Also cited from Maryland, Massachusetts, Michigan, New York, North Carolina, Pennsylvania, Rhode Island, and Missouri).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1973: 1434 (Acylophorus; notes; North Carolina).

  • — Smetana, 1978a: 836 (Acylophorus; collecting notes; Illinois; Kentucky).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas; New Jersey).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

ahrensii Nordmann, 1837, see: glaberrimus Herbst, 1784.

allardi Levasseur, 1968: 1400 (Acylophorus; Type locality: Katanga: Luena, Haut-Lomami).

  • Distribution: Congo.

alternans Smetana, 1971: 256 (Acylophorus; subgenus Acylophorus; Type locality: New York: Staten Is. Also cited from Missouri, New Jersey, and Ontario).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

anguliceps Bierig, 1938: 133 (Acylophorus; Type locality: Costa Rica: La Caja; Vara Blanca).

  • Distribution: Costa Rica.

angusticeps Sharp, 1876: 99 (Acylophorus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

antennalis Cameron, 1932b: 135 (Acylophorus; Type locality: Lolo Damvu, Mayumbe).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Congo.

asperatus Fauvel, 1878g: 561 (Acylophorus; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Bernhauer, 1920: 18 (Acylophorus; Australia).

  • Distribution: Australia.

asperipennis Smetana, 1967i: 561 (Acylophorus; Type locality: Türkei: Gaziantep).

  • — Korge, 1971a: 32 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 299 (Acylophorus; cited as valid species; characters; Anatolia; Lebanon).

  • Distribution: Lebanon, Turkey.

bafutensis Levasseur, 1968: 1401 (Acylophorus; Type locality: Cameroun: forêt de Bafut, Nguemba).

  • Distribution: Cameroon.

balchhi Smetana, 1988: 350 (Acylophorus; Type locality: India: Meghalaya, Khasi Hills, 1000 m, Mawsynram-Balat).

  • Distribution: India.

beesoni Cameron, 1926b: 371 (Acylophorus; Type locality: Kaligad, Dehra Dun; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 304 (Acylophorus; characters; India).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1988: 341 (Acylophorus; lectotype designation: Kaligad, Dehra Dun; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal.

ventralis Coiffait, 1983d: 168 (Acylophorus; Type locality: Nepal: Bandha, vallée de Katmandou).

  • — Smetana, 1988: 342 (Acylophorus; synonym of beesoni).

bipunctatus Cameron, 1920a: 219 (Acylophorus; Type locality: Nilgiri Hills).

  • — Cameron, 1932: 308 (Anchocerus; characters; India).

  • — Scheerpeltz, 1933: 1466, 1467 (Acylophorus and Anchocerus; [Note: Name cited in two genera]; catalog).

  • — Smetana, 1988: 359 (Acylophorus; characters; India).

  • Distribution: India.

borneensis Cameron, 1933b: 348 (Acylophorus; Type locality: British North Borneo: Kamborangah, 7200 feet).

  • Distribution: Borneo.

breviceps Cameron, 1939b: 147 (Acylophorus; Type locality: New Guinea: Cyclops Mts.: Sabron, altitude 930 feet).

  • Distribution: New Guinea.

bruchi Bernhauer, 1912e: 178 (Acylophorus; Type locality: Argentinien: Prov. Buenos Aires).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Argentina.

bumbunae Bordoni, 1994a: 311 (Acylophorus; Type locality: Sierra Leone: Northern Province: presso Bumbuna).

  • Distribution: Sierra Leone.

capensis Cameron, 1945c: 716 (Acylophorus; Type locality: South Africa: Simon's Town).

  • Distribution: South Africa.

caseyi Leng, 1920: 109 (Acylophorus; replacement name for longicornis Casey).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Smetana, 1965: 49 (Acylophorus; Nova Scotia).

  • — Smetana, 1971: 253 (Acylophorus; subgenus Acylophorus; characters; Nova Scotia; Ontario; Alabama; Illinois; Massachusetts; New Jersey; New York; Pennsylvania; Wisconsin).

  • — Smetana, 1973: 1432 (Acylophorus; notes; Quebec).

  • — Smetana, 1976c: 183 (Acylophorus; Ontario; Quebec).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; Louisiana; Wisconsin; Quebec).

  • — Smetana, 1990d: 102 (Acylophorus; Maine; West Virginia; Wisconsin).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

longicornis Casey, 1915: 427 (Acylophorus; [preoccupied]; Type locality: New York: Peekskill).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of caseyi).

  • — Smetana, 1965: 49 (Quedius; synonym of caseyi).

  • — Smetana, 1971: 253 (Acylophorus; synonym of caseyi).

cephalotes Bierig, 1938: 124 (Acylophorus; Type locality: Panama: France Field, Zona del Canal).

  • Distribution: Panama.

charaa Smetana, 1988: 356 (Acylophorus; Type locality: Nepal: Lalitpur Distr., 2 km S Godavari, 1700 m).

  • Distribution: Nepal.

chillo Smetana, 1988: 343 (Acylophorus; Type locality: India: Assam, Manas, 200 m).

  • Distribution: India.

chontalenus Sharp, 1884: 326 (Acylophorus; Type locality: Nicaragua).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Nicaragua.

ciliatus Roth, 1851, see: Moeocerus.

collarti Cameron, 1935a: 375 (Acylophorus; Type locality: Nizi: Blukwa).

  • Distribution: Congo.

congener Bierig, 1938: 127 (Acylophorus; Type locality: Panama: France Field, Zona; Puerto Armuelles, Costa pacifica).

  • Distribution: Panama.

congoensis Cameron, 1932b: 134 (Acylophorus; Type locality: Blukwa, Nizi).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Bernhauer, 1942c: 364 (Acylophorus; Cameroon).

  • — Cameron, 1950g: 43 (Acylophorus; Belgian Congo).

  • Distribution: Congo.

daai Smetana, 1988: 348 (Acylophorus; Type locality: Nepal: Khandbari District, below Sheduwa, 2100–2500 m. Also cited from India: Uttar Pradesh).

  • Distribution: Nepal, India.

dampfi Bierig, 1938: 134 (Acylophorus; Type locality: Mexico: Tenancingo, Desierto del Convento, 2400 m).

  • Distribution: Mexico.

dankalensis Bordoni, 1994a: 311 (Acylophorus; Type locality: Sierra Leone: North Province: Monti Loma, Dankale, 800 m).

  • Distribution: Sierra Leone.

darlingtoni Blackwelder, 1943: 470 (Acylophorus; Type locality: Trinidad: Cedros).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • Distribution: West Indies.

delphinus Fauvel, 1905b: 181 (Acylophorus; Type locality: Pays Androy: Ambovombe).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Jarrige, 1965: 141 (Acylophorus; characters; Madagascar).

  • Distribution: Madagascar.

densipennis Bernhauer, 1929f: 125 (Acylophorus; Type locality: Haut-Uele: Moto; Ituri: La Moto: Madyu).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Cameron, 1950g: 43 (Acylophorus; Belgian Congo).

  • Distribution: Congo.

densus LeConte, 1878a: 387 (Acylophorus; Type locality: Enterprise).

  • — Horn, 1878: 152 (Acylophorus; characters; Florida; Maryland).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 254 (Acylophorus; subgenus Acylophorus; characters; Florida; Louisiana).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; Florida; Illinois; Louisiana; Texas).

  • — Smetana, 1990d: 102 (Acylophorus; Georgia).

  • Distribution: USA.

didymus Sharp, 1884: 325 (Acylophorus; Type locality: Guatemala, Dueñas).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Guatemala.

dissimilis Smetana, 1971: 259 (Acylophorus; subgenus Acylophorus; Type locality: California: Westwood).

  • — Smetana, 1973: 1432 (Acylophorus; notes; California).

  • — Smetana, 1978a: 833 (Acylophorus; collecting notes; California).

  • — Smetana, 1990d: 103 (Acylophorus; Colorado).

  • Distribution: USA.

fallax Smetana, 1971: 254 (Acylophorus; subgenus Acylophorus; Type locality: Florida: Orange Co.).

  • — Smetana, 1990d: 102 (Acylophorus; Florida; Georgia).

  • Distribution: USA.

femoralis Nordmann, 1837: 128 (Acylophorus; Type locality: Rio-Janeiro).

  • — Erichson, 1840: 520 (Acylophorus; characters; Brazil).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

filius Smetana, 1971: 266 (Acylophorus; subgenus Acylophorus; Type locality: Pennsylvania: Pittsburg. Also cited from Illinois, Iowa, Massachusetts, New York, and Pennsylvania).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1973: 1434 (Acylophorus; notes; Michigan).

  • — Puthz, 1974: 12 (Acylophorus, cited as Acylophorys; Illinois).

  • — Smetana, 1978a: 838 (Acylophorus; collecting notes; New Jersey).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

flavicollis Sachse, 1852: 143 (Acylophorus; Type locality: Georgia).

  • — Horn, 1878: 151 (Acylophorus; characters; Pennsylvania to Kansas and Texas).

  • — Blatchley, 1910: 368 (Acylophorus; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 267 (Acylophorus; subgenus Acylophorus; characters; Arkansas; Illinois; Indiana; Iowa; Kansas; KenÏtucky; Michigan; Maryland; Missouri; New Mexico; New York; Ohio; Oklahoma; Pennsylvania; South Dakota; Tennessee; Texas; Virginia).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1978a: 838 (Acylophorus; collecting notes; Illinois; Kentucky; Texas).

  • — Smetana, 1981b: 643 (Acylophorus; Arkansas; Ontario).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Smetana, 1990d: 103 (Acylophorus; Mississippi).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • — Schmidt, 1999: 108 (Acylophorus; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: USA.

flavicornis Cameron, 1918c: 214 (Acylophorus; Type locality: Borneo, Mt. Matang, W. Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Borneo.

flavipes LeConte, 1878, see: lecontei Duvivier, 1883.

flavipes Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Cameron, 1932: 304 (Acylophorus; characters; East Indies).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Cameron, 1934d: 79 (Acylophorus; type examined).

  • — Cameron, 1940: 89 (Rhygmacera; characters).

  • — Coiffait, 1982a: 234 (Acylophorus; Nepal).

  • — Smetana, 1988: 356 (Acylophorus; lectotype designation; characters; species may not occur in the Himalaya).

  • Distribution: “East Indies”.

fordi Last, 1987: 53 (Acylophorus; Type locality: Papua New Guinea: New Guinea NE: Sepalakambang, Salawaket Range, 1920 m).

  • Distribution: New Guinea.

franckei Wendeler, 1956b: 264 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

furcatus Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Bernhauer, 1922c: 233 (Acylophorus; Formosa).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog; Philippines).

  • — Cameron, 1934d: 79 (Acylophorus; type examined).

  • — Smetana, 1988: 345 (Acylophorus; lectotype designation; characters; Assam; Meghalaya; West Bengal).

  • — Smetana, 1995: 125 (Acylophorus; characters; bionomics; Taiwan).

  • Distribution: “East Indies”, India, Philippines, Taiwan.

gilensis LeConte, 1863a: 34 (Acylophorus; Type locality: Gila River below Pima villages; [Note: See lectotype designation by Smetana, 1971]).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of pronus).

  • — Smetana, 1971: 257 (Acylophorus; subgenus Acylophorus; lectotype designation; characters; Arizona; Arkansas; California; New Mexico; Texas; Utah).

  • — Smetana, 1971a: 1848 (Acylophorus; collecting notes; Arizona).

  • Distribution: USA.

glaberrimus Herbst, 1784: 151 (Staphylinus; Type locality: Berlin; [Note: See lectotype designation by Gusarov, 1993]).

  • — Gmelin, 1790: 2032 (Staphylinus; page misnumbered as 1032; characters).

  • — Turton, 1802: 516 (Staphylinus; characters; Germany).

  • — Fauvel, 1886: 61 [= 1886a: 53] (Acylophorus; Algeria).

  • — Fauvel, 1889a: 112 (Acylophorus; notes; North America and Europe).

  • — Ganglbauer, 1895: 381 (Acylophorus; characters; Europe; Mediterranean region; North America; Mexico).

  • — Fauvel, 1897d: 311 (Acylophorus; Morocco).

  • — Fauvel, 1902b: 119 (Acylophorus; Morocco; Algeria).

  • — Porta, 1907: 99 (Acylophorus; characters; Italy).

  • — Reitter, 1909: 107 (Acylophorus; characters; Germany).

  • — Johansen, 1914: 342 (Acylophorus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Munster, 1923: 203 (Acylophorus; Fennoscandia).

  • — Lea, 1925: 252, 253 (Acylophorus; Australia, introduction).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • — Bierig, 1938: 123 (Acylophorus; type species of Acylophorus).

  • — Normand, 1947a: 9 (Acylophorus; Tunisia).

  • — V. Hansen, 1952: 162 (Acylophorus; characters; Denmark).

  • — Smetana, 1958: 399 (Acylophorus; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1960d: 136 (Acylophorus; Portugal).

  • — Palm, 1963b: 158 (Acylophorus; characters; Sweden; Denmark).

  • — Lohse, 1964: 220 (Acylophorus; characters; central Europe).

  • — M. Dvořák, 1965: 89 (Acylophorus; Slovakia).

  • — Horion, 1965: 308 (Acylophorus; Spain; France; Switzerland; Holland; Britain; Italy; Albania; Bosnia Hercegovina; Serbia; Bulgaria; Greece; Romania; Hungary; Germany; Austria; Slovakia; Czech Republic; Poland; Ukraine; Denmark; Sweden).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Smetana, 1971: 247 (Acylophorus; species does not occur in North America, previous citations referred to pratensis and perhaps gilensis; [Note: One name, luctuosus Solsky, described from Mexico, is still listed as a synonym of glaberrimus as it was not studied with by Smetana]).

  • — Korge, 1971a: 32 (Acylophorus; characters; Turkey).

  • — Pope, 1977: 32 (Acylophorus; Britain).

  • — Coiffait, 1978: 298 (Acylophorus; characters; Europe; Britain; Scandinavia; North Africa).

  • — Tóth, 1984: 94 (Acylophorus; characters; Hungary).

  • — Ádám, 1987: 153 (Acylophorus; collecting notes; Hungary).

  • — Lucht, 1987: 111 (Acylophorus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Gusarov, 1993: 72 (Acylophorus; lectotype designation; synonyms; Georgia; Ukraine).

  • — Wittwer, 1993: 251 (Acylophorus; Switzerland).

  • — Smetana, 1993b: 50 (Acylophorus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Acylophorus; Italy).

  • — Schülke, 1998g: 90 (Acylophorus; Germany).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Georgia, Turkey, Australia.

glabricollis Lacordaire, 1835: 396 (Staphylinus; Type locality: Paris).

  • — Erichson, 1839a: 482 (Acylophorus; characters; Germany).

  • — Erichson, 1840: 519 (Acylophorus; characters; Europe).

  • — Heer, 1841: 584 (Acylophorus; characters; Switzerland).

  • — Redtenbacher, 1849: 826 (Acylophorus; characters; Austria).

  • — Kiesenwetter, 1851a: 418 (Acylophorus; Spain).

  • — Fairmaire and Laboulbène, 1856: 534 (Acylophorus; characters; France).

  • — Kraatz, 1857d: 480 (Acylophorus; characters; Germany).

  • — Redtenbacher, 1857: 199 (Acylophorus; characters; Austria).

  • — Peyron, 1858: 428 (Acylophorus; Turkey).

  • — Thomson, 1859: 26 (type species of Acylophorus).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; characters; France; Algeria; Cyprus; Caramania; Caucasus; USA; Mexico).

  • — Redtenbacher, 1874: 195 (Acylophorus; characters; Austria).

  • — Seidlitz, 1875: 263 (Acylophorus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 834 [= 1877: 690] (Acylophorus; characters; France).

  • — Fauvel, 1878: 131 [= 1878a: 51] (Acylophorus; Algeria).

  • — Fowler, 1888: 223 (Acylophorus; characters; Britain).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Fauvel, 1902b: 119 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Montandon, 1908: 80 (Acylophorus; Romania).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Portevin, 1929: 335 (Acylophorus; characters; France).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of glaberrimus).

  • — Pope, 1977: 32 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

ahrensii Nordmann, 1837: 127 (Acylophorus; Type locality: Grunewald prope Berolinum).

  • — Erichson, 1839a: 482 (Acylophorus; synonym of glabricollis).

  • — Erichson, 1840: 519 (Acylophorus; synonym of glabricollis).

  • — Kraatz, 1857d: 480 (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

rufilabris Zetterstedt, 1838: 65 (Staphylinus; Type locality: Lapponia Umensis).

  • — Erichson, 1840: 519 (Acylophorus; synonym of glabricollis).

  • — Kraatz, 1857d: 480 (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

nitidus Motschulsky, 1845a: 40 (Rhygmacera; Type locality: Géorgie; [Note: See lectotype designation by Gusarov, 1993]).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Coiffait, 1978: 300 (Acylophorus; characters).

  • — Gusarov, 1993: 72 (Acylophorus; lectotype designation; synonym of glaberrimus).

luctuosus Solsky, 1868a: 122 (Acylophorus; Type locality: Mexique).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Sharp, 1884: 325 (Acylophorus; notes; Mexico).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

pulcher Scriba, 1870: 81 (Acylophorus; Type locality: Santiago de Compostella Galaeciae).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Smetana, 1958: 399 (Acylophorus; synonym of glaberrimus).

  • — Coiffait, 1978: 298 (Acylophorus; synonym of glaberrimus).

glabricollis Lacordaire, 1835, see: glaberrimus Herbst, 1784.

grandis Bernhauer, 1929f: 124 (Acylophorus; Type locality: Manyema: Niemba-Kalembe).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Congo.

hatuey Bierig, 1938: 126 (Acylophorus; Type locality: Cuba: Sierra del Rosario, El Rangel; Laguna del Somorrostro; Aspiro).

  • — Blackwelder, 1943: 469 (Acylophorus; characters; Cuba).

  • Distribution: West Indies.

hayashii Smetana, 1995: 129 (Acylophorus; Type locality: Chipon, Formosa).

  • Distribution: Taiwan.

hwang Smetana, 1995: 127 (Acylophorus; Type locality: Taiwan: Koantauchi, Nantou Hsien).

  • Distribution: Taiwan.

honshuensis R. Dvořák, 1957: 10 (Acylophorus; Type locality: Nagoya-Honshu, Japon).

  • Distribution: Japan.

hubrichi Bernhauer, 1927e: 249 (Acylophorus; Type locality: Argentinien: Prov. Santa Fe, Rosario).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Argentina.

immotus† Scudder, 1900: 47 (Acylophorus; fossil; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 723 (Acylophorus; catalog; Miocene; Colorado).

  • Distribution: USA.

indignus Blackburn, 1888: 4 (Acylophorus; Type locality: near Adelaide, banks of the Torrens).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Australia.

insulanus Bierig, 1938: 125 (Acylophorus; Type locality: Cuba: El Mariel; Laguna La Canoa, Portugues; Isla de Pinos).

  • — Blackwelder, 1943: 468 (Acylophorus; characters; Cuba; Isle of Pines).

  • Distribution: USA.

iridescens Sharp, 1876: 100 (Acylophorus; Type locality: Tapajos).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Brazil.

iridiventris Bernhauer, 1927e: 248 (Acylophorus; Type locality: Brasilien: S. Catharina).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Brazil.

jalapanus Schubert, 1909: 289 (Acylophorus; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Mexico.

javanus Cameron, 1937: 30 (Acylophorus; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft).

  • Distribution: Indonesia.

khairo Smetana, 1988: 344 (Acylophorus; Type locality: India: Meghalaya, Garo Hills, Songsak, 400 m).

  • Distribution: India.

kraatzi Bernhauer, 1908d: 336 (Acylophorus; Type locality: Paraguay).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: Paraguay.

kumbuiensis Bordoni, 1994a: 312 (Acylophorus; Type locality: Sierra Leone: Eastern Province: Kambui Hills Kenema).

  • Distribution: Sierra Leone.

leechi Smetana, 1971: 268 (Acylophorus; subgenus Palpacylophorus; Type locality: Missouri: Butler Co., stream at Hwy. 60, 12 mi. SE Ellsinore. Also cited from Texas).

  • Distribution: USA.

lecontei Duvivier, 1883: 136 (Acylophorus; replacement name for flavipes LeConte).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • — Smetana, 1971: 260 (Acylophorus; subgenus Acylophorus; characters; Arkansas; Florida; Georgia; Maryland Mississippi; New Jersey; Pennsylvania; South Carolina).

  • — Smetana, 1978a: 834 (Acylophorus; collecting notes; Alabama; Kentucky; Louisiana; Mississippi; North Carolina; Texas).

  • — Smetana, 1981b: 643 (Acylophorus; collecting notes; Florida).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas; West Virginia).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

flavipes LeConte, 1878a: 387 (Acylophorus; [preoccupied]; Type locality: Capron).

  • — Horn, 1878: 152 (Acylophorus; characters; Florida).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of lecontei).

  • — Smetana, 1971: 260 (Acylophorus; synonym of lecontei).

leonensis Bordoni, 1994a: 312 (Acylophorus; Type locality: Sierra Leone: Western Area: Sugar Loaf Mt.).

  • Distribution: Sierra Leone.

lindbergi Scheerpeltz, 1958a: 26 (Acylophorus; Type locality: Van, Südost-Ufer des Van-Sees in Armenien).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Coiffait, 1978: 299 (Acylophorus; characters; Turkey).

  • Distribution: Turkey.

lomaensis Levasseur, 1971: 303 (Acylophorus; Type locality: monts Loma, Sierra Leone, 1600 m).

  • Distribution: Sierra Leone.

longiceps Cameron, 1918, see: Acylohsellus.

longicornis Casey, 1915, see: caseyi Leng, 1920.

longicornis Erichson, 1840: 520 (Acylophorus; Type locality: Cayenna).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; catalog).

  • Distribution: French Guiana.

longistylus Casey, 1915: 426 (Acylophorus; Type locality: Florida).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of pronus).

  • — Smetana, 1971: 261 (Acylophorus; subgenus Acylophorus; characters; Alabama; Florida; Louisiana; New Jersey; New York; Pennsylvania).

  • — Smetana, 1978a: 835 (Acylophorus; collecting notes; Florida; Illinois; Louisiana; Mississippi; Texas).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: USA.

lualabaensis Levasseur, 1968: 1404 (Acylophorus; Type locality: Katanga: région de Kolwezi, Zilo, Lualaba).

  • Distribution: Congo.

luctuosus Solsky, 1868, see: glaberrimus Herbst, 1784.

maculicornis Cameron, 1918c: 214 (Acylophorus; Type locality: Borneo, Mt. Matang, W. Sarawak).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; catalog).

  • Distribution: Borneo.

malaisei Scheerpeltz, 1965: 297 (Acylophorus; Type locality: N.E. Burma, Kambaiti, 2000 m).

  • Distribution: Myanmar.

mareei Bernhauer, 1943b: 290 (Acylophorus; Type locality: Kongo: Lokandu).

  • Distribution: Congo.

marginalis Cameron, 1948: 42 (Acylophorus; Type locality: Bassin du Chari, Rivière Gribingui).

  • — Tottenham, 1961: 205 (Acylophorus; Senegal).

  • Distribution: Chad, Senegal.

metallicus Blackwelder, 1943: 469 (Acylophorus; Type locality: Dominican Republic: Sanchez).

  • Distribution: West Indies.

methneri Bernhauer, 1917g: 48 (Acylophorus; Type locality: Deutsch-Ostafrika: Kaguru).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Bernhauer and Chapman, 1939: 74 (Acylophorus; Kenya).

  • — Scheerpeltz, 1974c: 25 (Acylophorus; Sudan).

  • Distribution: Sudan, Kenya, Tanzania.

mexicanus Sharp, 1884: 326 (Acylophorus; Type locality: Mexico, Oaxaca, Las Vigas).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

microcephalus Cameron, 1932: 305 (Acylophorus; Type locality: Mussoorie district: Arni Gad and Kali Gad; [Note: See lectotype designation by Smetana, 1988]. Also cited from Mussoorie district: Kali Gad).

  • — Cameron, 1932: 304 (Acylophorus; [Note: Misidentified as furcatus according to Smetana, 1988: 352]; characters; India).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Bierig, 1938: 123 (Acylophorus; type species of Indoacylophorus).

  • — Smetana, 1988: 352 (Acylophorus; lectotype designation: Arni Gad, Mussoorie; characters; Uttar Pradesh; West Bengal).

  • Distribution: India.

microcerus Fauvel, 1895: 276 (Acylophorus; Type locality: Bírmanie, Bhamo).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Cameron, 1932: 305 (Acylophorus; characters; Burma).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Scheerpeltz, 1976: 44 (Acylophorus; Nepal).

  • Distribution: Myanmar.

minutus Jarrige, 1965: 142 (Acylophorus; Type locality: Perinet, Madagascar Est).

  • Distribution: Madagascar.

muscorum Jarrige, 1965: 140 (Acylophorus; Type locality: massif du Tsaratanana, 1500 m).

  • Distribution: Madagascar.

nepa Bierig, 1938: 131 (Acylophorus; Type locality: Panama: Puerto Armuelles).

  • Distribution: Panama.

nepalicus Coiffait, 1981, see: Erichsonius.

nigeriae Tottenham, 1961: 205 (Acylophorus; [Note: Nomen nudum, cited from Senegal with Cameron as author but I find no description of the species by either Cameron or Tottenham (Article 13.1.1)]).

nitidus Motschulsky, 1845, see: glaberrimus Herbst, 1784.

obscurus Cameron, 1942a: 139 (Acylophorus; Type locality: Brunei).

  • Distribution: Brunei.

oculifer Bierig, 1938: 137 (Acylophorus; subgenus Neoacylophorus; Type locality: Panama: France Field, Zona).

  • Distribution: Panama.

orientalis Fauvel, 1907: 48 (Acylophorus; Type locality: Voi; Kisoumou [Victoria Nyanza]; baie de Kavirondo; steppe Masai pres Nairobi).

  • — Eichelbaum, 1913: 129 (Acylophorus; Tanzania).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bernhauer, 1931d: 588 (Acylophorus; Abyssinia).

  • — Scheerpeltz, 1974c: 25 (Acylophorus; Sudan).

  • Distribution: Sudan, Ethiopia, Kenya, Tanzania.

pacificus Bierig, 1938: 128 (Acylophorus; Type locality: Panama: France Field, Zona; Puerto Armuelles, Costa pacifica).

  • Distribution: Panama.

palpalis Sharp, 1884: 327 (Acylophorus; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bierig, 1938: 135 (Acylophorus; Panama).

  • Distribution: Guatemala, Panama.

parviceps Sharp, 1884: 326 (Acylophorus; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Guatemala.

persimilis Cameron, 1950d: 24 (Acylophorus; Type locality: Malacca: Perak).

  • Distribution: Indonesia.

picipennis Roth, 1851, see: Glyphesthus.

picipennis Bernhauer, 1929f: 124 (Acylophorus; [preoccupied]; Type locality: Luebo).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Congo.

picipes Erichson, 1840: 521 (Acylophorus; Type locality: Columbia).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Colombia.

planatus Sharp, 1884: 327 (Acylophorus; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Nicaragua.

plaumanni Wendeler, 1956b: 263 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

pratensis LeConte, 1863a: 34 (Acylophorus; Type locality: Kansas).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; synonym of glabricollis).

  • — Horn, 1878: 151 (Acylophorus; characters; Kansas).

  • — Ganglbauer, 1895: 381 (Acylophorus; synonym of glaberrimus).

  • — Porta, 1907: 99 (Acylophorus; synonym of glaberrimus).

  • — Bernhauer and Schubert, 1916: 438 (Acylophorus; synonym of glaberrimus).

  • — Scheerpeltz, 1933: 1466 (Acylophorus; synonym of glaberrimus; valid species?).

  • — Smetana, 1965: 50 (Acylophorus; characters; Newfoundland).

  • — Smetana, 1971: 248 (Acylophorus; subgenus Amacylophorus; characters; Newfoundland; Quebec; Massachusetts; Minnesota).

  • — Smetana, 1973: 1432 (Acylophorus; notes; Quebec).

  • — Smetana, 1981b: 643 (Acylophorus; collecting notes; Ontario).

  • — Smetana, 1990d: 102 (Acylophorus; New Hampshire).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

princeps Smetana, 1971: 255 (Acylophorus; subgenus Acylophorus; Type locality: Florida: Homestead, Dade Co.).

  • — Smetana, 1981b: 643 (Acylophorus; Florida).

  • Distribution: USA.

pronus Erichson, 1840: 521 (Acylophorus; Type locality: America septentrionalis).

  • — Horn, 1878: 151 (Acylophorus; characters; USA).

  • — Blatchley, 1910: 368 (Acylophorus; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Notman, 1919: 135 (Acylophorus; New York).

  • — Notman, 1920a: 180 (Acylophorus; New York).

  • — Notman, 1920b: 706 (Acylophorus; Florida).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Hatch, 1957: 231 (Acylophorus; characters; British Columbia; Washington).

  • — Smetana, 1965: 48 (Acylophorus; characters; Nova Scotia; Newfoundland).

  • — Smetana, 1965a: 17 (Acylophorus; Alaska).

  • — Smetana, 1971: 264 (Acylophorus; subgenus Acylophorus; characters; British Columbia; Manitoba; New Brunswick; Newfoundland; Northwest Territories; Nova Scotia; Ontario; Quebec; Alaska; Connecticut; Illinois; Massachusetts; Maine; Michigan; Minnesota; New Hampshire; New Jersey; New York; Oregon; Pennsylvania; Rhode Island; Washington; Wisconsin).

  • — Smetana, 1971b: 4 (Acylophorus; Michigan).

  • — Smetana, 1976c: 184 (Acylophorus; California; Washington).

  • — Frank, 1982: 33 (Acylophorus; list of parasites).

  • — Downie and Arnett, 1996: 395 (Acylophorus; characters; USA).

  • Distribution: Canada, USA.

proximus Sharp, 1884: 325 (Acylophorus; Type locality: Guatemala, Dueñas, Paso Antonio).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Guatemala.

pulcher Scriba, 1870, see: glaberrimus Herbst, 1784.

puncticeps Fauvel, 1895: 275 (Acylophorus; Type locality: Bírmanie, Carin Ghécu, 1300–1400 m).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Cameron, 1932: 303 (Acylophorus; characters; Burma).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Smetana, 1988: 359 (Acylophorus; characters; Burma).

  • Distribution: Myanmar.

punctiventris Sharp, 1876: 98 (Acylophorus; Type locality: Ega; Tapajos).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Brazil.

punctulatus Bierig, 1938: 130 (Acylophorus; Type locality: Costa Rica: San José, La Caja).

  • Distribution: Costa Rica.

pusillus Smetana, 1973: 1432 (Acylophorus; Type locality: U.S.A.: Florida: Lake Istokpoga, near Lake Placid).

  • Distribution: USA.

raato Smetana, 1988: 351 (Acylophorus; Type locality: Nepal: Khandbari District, Arun River at Num, 1500–1600 m. Also cited from India: Meghalaya; West Bengal).

  • — Smetana, 1992b: 10 (Acylophorus; Nepal).

  • Distribution: Nepal, India.

richteri Bernhauer, 1912e: 177 (Acylophorus; Type locality: Argentinien: Prov. Buenos).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Argentina.

rossii Bordoni, 1994a: 309 (Acylophorus; Type locality: Sierra Leone: Western Area: Sugar Loaf Mt.).

  • Distribution: Sierra Leone.

rotundicollis Cameron, 1918: 90 (Acylophorus; Type locality: Singapore: Bukit Timah).

  • — Cameron, 1921a: 379, 405 (Acylophorus; characters; Singapore).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Singapore.

ruficeps Cameron, 1941c: 383 (Acylophorus; Type locality: Philippines: Mindanao: Surigao).

  • Distribution: Philippines.

ruficollis Kraatz, 1859, see: ruficollis Motschulsky, 1858.

ruficollis Motschulsky, 1858: 657 (Acylophorus; Type locality: Indes orientales; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Bernhauer, 1920: 18 (Acylophorus; Australia).

  • — Wendeler, 1924a: 344 (Acylophorus; East Indies; Ceylon; Australia).

  • — Bernhauer, 1928: 18 (Acylophorus; Sumatra).

  • — Cameron, 1932: 303 (Acylophorus; characters; Ceylon; Penang).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Cameron, 1933d: 393 (Acylophorus; Sumatra).

  • — Last, 1964: 177 (Acylophorus; Thailand).

  • — Smetana, 1988: 349 (Acylophorus; lectotype designation; characters; Assam; Nepal).

  • — Barrion, Litsinger, and Last, 1991: 1078 (Acylophorus; associated with rice agroecosystem).

  • Distribution: “East Indies”, Nepal, Sri Lanka, Malaysia, Thailand, Indonesia, Australia.

ruficollis Kraatz, 1859: 65 (Acylophorus; [preoccupied]; Type locality: Ceylan).

  • — Lea, 1925: 252, 253 (Acylophorus; characters; Australia, introduction).

  • — Cameron, 1932: 303 (Acylophorus; synonym of ruficollis).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; synonym of ruficollis).

  • — Smetana, 1988: 349 (Acylophorus; synonym of ruficollis).

rufilabris Zetterstedt 1838, see: glaberrimus Herbst, 1784.

rufipennis Cameron, 1959: 117 (Acylophorus; Type locality: Angola: Dundo, River Luachimo).

  • Distribution: Angola.

sandersoni Smetana, 1971: 258 (Acylophorus; subgenus Acylophorus; Type locality: Arkansas: Washington Co.).

  • Distribution: USA.

schmidti Bierig, 1938: 135 (Acylophorus; subgenus Paracylophorus; Type locality: Costa Rica: San José, La Caja).

  • Distribution: Costa Rica.

scutellaris Bernhauer, 1916b: 21 (Acylophorus; Type locality: Borneo).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Cameron, 1933d: 393 (Acylophorus; Sumatra).

  • — Cameron, 1937: 30 (Acylophorus; Java).

  • Distribution: Borneo, Indonesia.

similis Wendeler, 1956b: 264 (Acylophorus; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

simplex Sharp, 1884: 325 (Acylophorus; Type locality: Mexico, Oaxaca, Mirador, Cordova).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

siyo Smetana, 1988: 344 (Acylophorus; Type locality: Nepal: Khandbari Distr., 2 km E Mansingma, 1900 m).

  • — Smetana, 1992b: 10 (Acylophorus; Nepal).

  • Distribution: Nepal.

sogai Jarrige, 1965: 142 (Acylophorus; Type locality: Mahanilazy, Ambatondrazaka, Madagascar Est).

  • Distribution: Madagascar.

tenuiceps Bernhauer, 1932b: 157 (Acylophorus; Type locality: Haut-Uele: Yebo; Moto).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Congo.

tenuipes Lea, 1929: 207 (Acylophorus; Type locality: Queensland: Brisbane).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Australia.

tibialis Cameron, 1932: 305 (Acylophorus; Type locality: Burma: Ruby Mines; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — Scheerpeltz, 1965: 112 (Acylophorus; Burma).

  • — Smetana, 1988: 357 (Acylophorus; lectotype designation; characters; Burma).

  • Distribution: Myanmar.

trigonocephalus Cameron, 1949a: 322 (Acylophorus; Type locality: Côte d'Ivoire: Mt. Tonkoui, alt. 900–1200 m).

  • Distribution: Ivory Coast.

trinitatis Blackwelder, 1943: 467 (Acylophorus; Type locality: Trinidad: foothills north of Tunapuna).

  • Distribution: West Indies.

tristis Jarrige, 1965: 140 (Acylophorus; Type locality: Tananarive).

  • Distribution: Madagascar.

usambarae Bernhauer, 1917g: 48 (Acylophorus; Type locality: West-Usambara).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • Distribution: Tanzania.

ventralis Coiffait, 1983, see: beesoni Cameron, 1926.

versicolor Sharp, 1887: 783 (Acylophorus; Type locality: Mexico, Tehuantepec).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • Distribution: Mexico.

wagenschieberi Kiesenwetter, 1850: 220 (Acylophorus; Type locality: Berolini).

  • — Kiesenwetter, 1851a: 419 (Acylophorus; characters; Germany).

  • — Redtenbacher, 1857: 199 (Acylophorus; characters; Germany).

  • — Kraatz, 1857d: 481 (Acylophorus; characters; Germany).

  • — Thomson, 1860: 182 (Acylophorus; characters; Scandinavia).

  • — Fauvel, 1874: 316 [= 1874a: 542] (Acylophorus; characters; Scandinavia; Germany; Greece).

  • — Redtenbacher, 1874: 195 (Acylophorus; characters).

  • — Seidlitz, 1875: 263 (Acylophorus; characters; Baltic region).

  • — Mulsant and Rey, 1876: 836 [= 1877: 692] (Acylophorus; characters; France).

  • — Ganglbauer, 1895: 382 (Acylophorus; characters; Germany; Greece; Sweden).

  • — Reitter, 1909: 107 (Acylophorus; characters; Germany).

  • — Johansen, 1914: 344 (Acylophorus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 439 (Acylophorus; catalog).

  • — Munster, 1923: 203 (Acylophorus; Fennoscandia).

  • — Scheerpeltz, 1933: 1467 (Acylophorus; catalog).

  • — V. Hansen, 1952: 162 (Acylophorus; characters; Denmark).

  • — Smetana, 1958: 401 (Acylophorus; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 158 (Acylophorus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 220 (Acylophorus; characters; central Europe).

  • — Horion, 1965: 311 (Acylophorus; Russia; Finland; Sweden; Norway; Denmark; Poland; Czechoslovakia; Holland; Austria; Germany).

  • — Smetana, 1967i: 563 (Acylophorus; characters).

  • — Coiffait, 1978: 300 (Acylophorus; characters; Europe; Greece).

  • — Tóth, 1984: 94 (Acylophorus; characters).

  • — Segers, 1987: 60 (Acylophorus; Belgium).

  • — Lucht, 1987: 111 (Acylophorus; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Acylophorus; Czech Republic).

  • — Sterrenburg and Schülke, 1997: 20 (Acylophorus; Netherlands).

  • — Staneic, 1998a: 5 (Acylophorus; habitat; life cycle; Poland).

  • Distribution: Russia, Finland, Sweden, Norway, Denmark, Holland, Germany, Poland, Czechoslovakia, Austria.

williamsi Bierig, 1938: 129 (Acylophorus; Type locality: Brasil: Belen, Para).

  • Distribution: Brazil.

zdenae Smetana, 1978a: 836 (Acylophorus; Type locality: Texas: Trinity Co.: 12 mi. SW Lufkin. Also cited from Illinois and Louisiana).

  • — Smetana, 1990d: 103 (Acylophorus; Arkansas).

  • Distribution: USA.

ziloensis Levasseur, 1968: 1403 (Acylophorus; Type locality: Katanga: région de Kolwezi, Zilo, Lualaba).

  • Distribution: Congo.

Anaquedius [1 species; Nearctic Region]

Anaquedius Casey, 1915: 397, 400 (species included: vernix). Type species: Quedius vernix LeConte, fixed by original designation and monotypy.

  • — Casey, 1915: 397 (type species: Quedius vernix).

  • — Scheerpeltz, 1933: 1427 (subgenus of Quedius).

  • — Blackwelder, 1952: 50 (type species: vernix).

  • — Smetana, 1971: 242 (valid genus; characters).

  • — Moore and Legner, 1975: 51 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Moore and Legner, 1979: 42 (characters; notes).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Downie and Arnett, 1996: 394 (characters).

  • — Lundgren, 1998: 50 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

vernix LeConte, 1878a: 389 (Quedius; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Massachusetts; Michigan; Canada).

  • — Horn, 1878: 167 (Quedius; characters; Massachusetts, Canada, Illinois, Michigan).

  • — Blatchley, 1910: 373 (Quedius; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1465 (Quedius; catalog).

  • — Smetana, 1965: 35 (Quedius; subgenus Anaquedius; Nova Scotia).

  • — Smetana, 1971: 242 (Anaquedius; lectotype designation: Cambr. [Cambridge, Massachusetts]; characters; New Brunswick; Nova Scotia; Ontario; Quebec; Connecticut; Illinois; Indiana; Louisiana; Maine; Maryland; Massachusetts; Michigan; Minnesota; New Hamshire; New Jersey; New York; Ohio; Pennsylvania; Rhode Island; Virginia; Wisconsin).

  • — Smetana, 1973: 1432 (Anaquedius; notes; North Carolina).

  • — Smetana, 1978a: 831 (Anaquedius; collecting notes; Florida; Louisiana; Texas).

  • — Frank, 1982: 33 (Anaquedius; list of parasites).

  • — Smetana, 1990d: 102 (Anaquedius; Arkansas).

  • — Campbell and Davies, 1991: 122 (Anaquedius; checklist of species; Canada).

  • — Downie and Arnett, 1996: 394 (Anaquedius; characters; USA).

  • Distribution: Canada, USA.

Anchocerus [10 species; Palaearctic, Oriental, and Australian Regions]

Anchocerus Fauvel, 1905a: 141 (species included: birmanus). Type species: Anchocerus birmanus Fauvel, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 439 (world catalog; 1 species).

  • — Cameron, 1925: 73 (catalog of Indian species).

  • — Cameron, 1932: 306 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1467 (world catalog supplement).

  • — Blackwelder, 1952: 51(type species: birmanus).

  • — Scheerpeltz, 1962b: 268 (characters).

  • — Last, 1975: 441 (characters).

  • — Smetana, 1988: 361 (characters; key to species of Himalayan region).

  • — Smetana, 1995: 132 (notes; species of Taiwan).

birmanus Fauvel, 1905a: 141 (Anchocerus; Type locality: Tenasserim, Kawkareet, Bírmanie; [Note: See lectotype designation by Smetana, 1977a]. Originally cited from Bírmanie. Tenasserim. Java: Buitenzorg; Malang).

  • — Fauvel, 1905e: 84 (Anchocerus; Java).

  • — Bernhauer and Schubert, 1916: 439 (Anchocerus; catalog).

  • — Bernhauer, 1922a: 151 (Anchocerus; Indochina).

  • — Cameron, 1932: 306 (Anchocerus; characters; Burma: Tenasserim).

  • — Scheerpeltz, 1933: 1467 (Anchocerus; catalog).

  • — Cameron, 1937: 30 (Anchocerus; Java).

  • — Smetana, 1977a: 249 (Anchocerus; lectotype designation: Tenasserim, Kawkareet, Bírmanie; characters).

  • — Smetana, 1988: 364 (Anchocerus; characters; Burma; Uttar Pradesh).

  • Distribution: India, Myanmar, Indonesia.

monticola Cameron, 1926b: 371 (Anchocerus; Type locality: Kaligad, Dehra Dun; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussoorie district; Kaligad).

  • — Cameron, 1932: 307 (Anchocerus; characters; India).

  • — Scheerpeltz, 1933: 1467 (Anchocerus; catalog).

  • — Smetana, 1988: 363 (Anchocerus; lectotype designation: Kaligad, Dehra Dun; characters; India).

  • Distribution: India.

nepalicus Smetana, 1988: 366 (Anchocerus; Type locality: Nepal: Khandbari District: For. above Ahale, 2300 m. Also cited from India).

  • — Smetana, 1992b: 11 (Anchocerus; Nepal).

  • Distribution: India, Nepal.

novaguinensis Last, 1975: 442 (Anchocerus; Type locality: Okapa).

  • Distribution: New Guinea.

okasaensis Last, 1975: 441 (Anchocerus; Type locality: Okasa, 4000 ft.).

  • Distribution: New Guinea.

punctatissimus Smetana, 1977a: 248 (Anchocerus; Type locality: Bhutan: Changra, 18 km S. Tongsa).

  • — Smetana, 1988: 365 (Anchocerus; characters; Bhutan; Nepal).

  • Distribution: Bhutan, Nepal.

punctus Last, 1980: 151 (Anchocerus; Type locality: New Guinea: Wau, Mount Kumbak).

  • Distribution: New Guinea.

shibatai Smetana, 1995: 132 (Anchocerus; Type locality: Taiwan: near Lienhuachih, Nantou Hsien, 700 m).

  • Distribution: Taiwan.

similis Last, 1980: 150 (Anchocerus; Type locality: New Guinea: Friedrich Wilhelmshafen).

  • Distribution: New Guinea.

wilhelmensis Last, 1980: 150 (Anchocerus; Type locality: New Guinea: Mount Wilhelm, Field Stat., 3200 m).

  • Distribution: New Guinea.

Arrowinus [1 species; Ethiopian Region]

Arrowinus Bernhauer, 1935f: 215 (species included: phaenomenalis). Type species: Arrowinus phaenomenalis Bernhauer, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 63 (type species: phaenomenalis).

  • — Newton, 1985: 208 (comments on austral distribution and taxonomic notes).

phaenomenalis Bernhauer, 1935f: 216 (Arrowinus; Type locality: Natal).

  • Distribution: South Africa.

Astrapaeus [1 species; Palaearctic Region]

Astrapaeus Gravenhorst, 1802: 199 (species included: ulmi). Type species: Staphylinus ulmi Rossi, fixed by monotypy.

  • — Latreille, 1804: 284, 288 (characters).

  • — Latreille, 1802: 130 (characters).

  • — Latreille, 1806: 284 (characters).

  • — Latreille, 1810: 179, 427 (characters; type species).

  • — Leach, 1815: 91 (characters).

  • — Billberg, 1820: 13 (characters).

  • — Dejean, 1821: 23 (species in collection).

  • — Latreille, 1825: 244 (cited as Astrapée; characters).

  • — Berthold, 1827: 331 (characters).

  • — Le Peletier and Serville, 1828: 478 (characters; notes).

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829a: 22 (catalog; Britain).

  • — Stephens, 1829: 276 (catalog; Britain).

  • — Latreille, 1829: 433 (character).

  • — Mannerheim, 1830: 7, 19 (characters).

  • — Mannerheim, 1831: 421, 433 (characters).

  • — Stephens, 1832: 200, 213 (characters).

  • — Griffith and Pidgeon, 1832: 290 (characters).

  • — Dejean, 1833: 61 (in list of species).

  • — Gistel, 1834: 8 (in list of species).

  • — Lacordaire, 1835: 355 (characters).

  • — Nordmann, 1837: 16 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Heer, 1839: 282 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Erichson, 1840: 552 (characters).

  • — Laporte, 1840: 173 (characters).

  • — Redtenbacher, 1849: 712 (characters).

  • — Schaum, 1852: 27 (in list of species; Europe).

  • — Lacordaire, 1854: 86 (characters; notes; in list of species).

  • — Fairmaire and Laboulbène, 1856: 545 (characters).

  • — Jacquelin du Val, 1856: 40 (characters).

  • — Kraatz, 1857d: 521 (characters).

  • — Redtenbacher, 1857: 205 (characters; key to Austrian species).

  • — G. Waterhouse, 1858: 23 (catalog of British species).

  • — Schaum, 1859: 26 (catalog; European species).

  • — Gredler, 1863: 101 (in list of species from Tirol).

  • — Wencker and Silbermann, 1866: 28 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Fauvel, 1874: 313 [= 1874a: 539] (characters).

  • — Redtenbacher, 1874: 202 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 798 [= 1877: 654] (characters).

  • — Heyden, Reitter, and Weise, 1891: 99 (in list of species of Europe and Caucasus).

  • — Ganglbauer, 1895: 383 (characters).

  • — Heyden, Reitter, and Weise, 1906: 176 (species of Europe and Caucasus).

  • — Porta, 1907: 89 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Petri, 1912: 80 (in list of species of Siebenbürgen).

  • — Bernhauer and Schubert, 1916: 414 (world catalog; 1 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Porta, 1926: 118 (characters).

  • — Portevin, 1929: 336 (characters; key to French species).

  • — Roubal, 1930: 386 (catalog; Slovakia).

  • — Scheerpeltz, 1933: 1425 (world catalog supplement).

  • — Blackwelder, 1952: 66 (type species: ulmi).

  • — Smetana, 1958: 322 (characters; habitus; type species: ulmi).

  • — Smetana, 1959a: 215 (species collected in Albania).

  • — Scheerpeltz, 1962b: 264 (characters; type species: ulmi).

  • — Székessy, 1963: 62 (characters).

  • — Lohse, 1964: 205 (characters; central European species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Bordoni, 1973j: 713 (in list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 294 (characters; cited as Astrapeus).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 118 (catalog; Poland).

  • — Tóth, 1984: 97 (characters; habitus; species of Hungary).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic; cited as Astrapeus).

  • — Ciceroni and Zanetti, 1995: 31 (species of Italy).

  • — Sparacio, 1995: 164 (characters; notes; Sicily).

Systolastes Gistel, 1856: 388 (species included: ulmi). Type species: Staphylinus ulmi Rossi, fixed by monotypy as “Systolastes ulmi Pan.”

  • — Blackwelder, 1952: 368 (type species; synonym of Astrapaeus).

rufipennis Stephens, 1832, see: ulmi Rossi, 1790.

rufipes Latreille, 1806, see: Tasgius.

ulmi Rossi, 1790: 248 (Staphylinus; Type locality: Etruria).

  • — Olivier, 1795:(42): 17 (Staphylinus; characters; Italy).

  • — Gravenhorst, 1802: 199 (Astrapaeus; characters).

  • — Panzer, 1805: 4 (Astrapaeus; characters; illustrations).

  • — Gravenhorst, 1806: 47 (Staphylinus).

  • — Latreille, 1806: 284 (Astrapaeus; characters).

  • — Leach, 1815: 91 (Astrapaeus; characters; Italy; France).

  • — Le Peletier and Serville, 1828: 479 (notes).

  • — Mannerheim, 1830: 19 (Astrapaeus; southern Europe).

  • — Mannerheim, 1831: 433 (Astrapaeus; southern Europe).

  • — Griffith and Pidgeon, 1832: 290 (Astrapaeus).

  • — Nordmann, 1837: 16 (Astrapaeus; southern Europe).

  • — Westwood, 1838a: 16 (Astrapaeus; characters; type species).

  • — Heer, 1839: 282 (Astrapaeus; characters; Switzerland).

  • — Erichson, 1840: 553 (Astrapaeus; characters; France; Italy; Dalmatia; Hungary).

  • — Küster, 1847: 47 (Astrapaeus; characters; Germany; France; Italy; Dalmatia; Hungary).

  • — Hochhuth, 1849: 155 (Astrapaeus; characters; Helenendorf).

  • — Redtenbacher, 1849: 712 (Astrapaeus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 545 (Astrapaeus; characters; France).

  • — Kraatz, 1857d: 523 (Astrapaeus; characters; Germany).

  • — Redtenbacher, 1857: 205 (Astrapaeus; characters; Austria).

  • — Kraatz, 1858a: 56 (Astrapaeus; Greece).

  • — Fauvel, 1874: 314 [= 1874a: 540] (Astrapaeus; characters; France; Germany; Switzerland; Austria; Italy; Sardinia; Greece; Russia; Caucasus).

  • — Redtenbacher, 1874: 202 (Astrapaeus; characters; Austria).

  • — Mulsant and Rey, 1876: 801 [= 1877: 657] (Astrapaeus; characters; France).

  • — Seidlitz, 1891a: 420 (Astrapaeus; characters; Europe).

  • — Ganglbauer, 1895: 384 (Astrapaeus; characters; southern middle Europe; Mediterranean region).

  • — Sainte-Claire Deville, 1906: 104 (Astrapaeus; Corsica).

  • — Porta, 1907: 101 (Astrapaeus; characters; Italy).

  • — Montandon, 1908: 80 (Astrapaeus; Romania).

  • — Reitter, 1909: 107 (Astrapaeus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; catalog).

  • — Portevin, 1929: 336 (Astrapaeus; characters; France).

  • — Scheerpeltz, 1933: 1425 (Astrapaeus; catalog).

  • — Smetana, 1958: 322 (Astrapaeus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 205 (Astrapaeus; characters; central Europe).

  • — Raianu, 1964: 102 (Astrapaeus; Romania).

  • — Scheerpeltz, 1965b: 406 (Astrapaeus; Sicily).

  • — Scheerpeltz, 1965c: 511 (Astrapaeus; Sicily).

  • — Horion, 1965: 251 (Astrapaeus, cited as Astrapeus; Portugal; Spain; Corsica; Italy; Croatia; Bosnia Hercegovina; Montenegro; Albania; Greece; Corfu; Albania; Turkey; Slovenia; Serbia; Hungary; Czech Republic; Slovakia; Poland ?; Ukraine; Switzerland; France; Belgium; Austria; Germany ?).

  • — Bordoni, 1974b: 14 (Astrapaeus; Italy).

  • — Coiffait, 1978: 294 (Astrapaeus; characters; Europe; Britain; Scandinavia; North Africa).

  • — Tóth, 1984: 98 (Astrapaeus; characters; Hungary).

  • — Segers, 1986: 52 (Astrapaeus; checklist of species; Belgium).

  • — Lucht, 1987: 109 (Astrapaeus; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux).

  • — Smetana, 1993b: 49 (Astrapaeus, cited as Astrapeus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Astrapaeus; Italy).

  • Distribution: Europe, England, Russia, Caucasus.

ulmineus Fabricius, 1801: 595 (Staphylinus; Type locality: Italia).

  • — Panzer, 1805: 4 (Astrapaeus; synonym of ulmi).

  • — Gravenhorst, 1806: 47 (Staphylinus; synonym of ulmi Rossi).

  • — Guérin-Méneville, 1829: pl. 9, fig. 2 (Astrapaeus; illustration).

  • — Lacordaire, 1835: 356 (Astrapaeus; characters; France).

  • — Erichson, 1840: 553 (Astrapaeus; synonym of ulmi).

  • — Laporte, 1840: 173 (Astrapaeus; characters; Europe and northern Asia).

  • — Guérin-Méneville, 1844a: 30 (Astrapaeus; France).

  • — Fairmaire and Laboulbène, 1856: 545 (Astrapaeus; synonym of ulmi).

  • — Kraatz, 1857d: 523 (Astrapaeus; synonym of ulmi).

  • — Ganglbauer, 1895: 384 (Astrapaeus; synonym of ulmi).

  • — Porta, 1907: 101 (Astrapaeus; synonym of ulmi).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; synonym of ulmi).

  • — Smetana, 1958: 323 (Astrapaeus; synonym of ulmi).

  • — Coiffait, 1978: 295 (Astrapaeus; synonym of ulmi).

rufipennis Stephens, 1832: 214 (Astrapaeus; Type locality: Coombe-wood; Devonshire).

  • — Curtis, 1829: 24 (Astrapaeus; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 389 (Ocypus; characters; England).

  • — G. Waterhouse, 1858: 23 (Astrapaeus; synonym of ulmi).

  • — Gemminger and Harold, 1868: 573 (Astrapaeus; synonym of ulmi).

  • — Ganglbauer, 1895: 384 (Astrapaeus; synonym of ulmi).

  • — Porta, 1907: 101 (Astrapaeus; synonym of ulmi).

  • — Bernhauer and Schubert, 1916: 415 (Astrapaeus; synonym of ulmi).

  • — Smetana, 1958: 323 (Astrapaeus; synonym of ulmi).

  • — Coiffait, 1978: 295 (Astrapaeus; synonym of ulmi).

ulmineus Fabricius, 1801, see: ulmi Rossi, 1790.

Beeria [1 species; Nearctic Region]

Beeria Hatch, 1957: 214, 231 (species included: punctata). Type species: Beeria punctata Hatch, fixed by original designation and monotypy.

  • — Korge, 1971a: 36 (characters).

  • — Moore and Legner, 1975: 8 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Smetana, 1978a: 816 (characters).

  • — Smetana, 1995j: 131 (related to Strouhalium and Heinzia).

  • — Moore and Legner, 1979: 44 (characters; notes).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

nematocera Casey, 1915: 436 (Philonthus; Type locality: British Columbia: Metlakatla).

  • — Scheerpeltz, 1933: 1352 (Philonthus; catalog).

  • — Hatch, 1957: 198 (Philonthus; characters; British Columbia).

  • — Smetana, 1977b: 184 (Beeria; characters; ecological notes; British Columbia; Oregon; Washington).

  • — Smetana, 1978a: 815 (Beeria; notes; collecting notes; British Columbia; Oregon; Washington).

  • — Smetana, 1981b: 631 (Beeria; Alaska).

  • — Smetana, 1990d: 95 (Beeria; British Columbia; Washington).

  • — Campbell and Davies, 1991: 120 (Beeria; checklist of species; Canada).

  • Distribution: Canada, USA.

punctata Hatch, 1957: 231 (Beeria; Type locality: Oregon: Benton Co.: Mary's Peak).

  • — Smetana, 1977b: 184 (Beeria; synonym of nematocera).

  • — Smetana, 1978a: 815 (Beeria; synonym of nematocera).

punctata Hatch, 1957, see: nematocera Casey, 1915.

Bolitogyrus [29 species; Neotropical, Palaearctic, and Oriental Regions]

Bolitogyrus Chevrolat, 1842: 641 (species included: buphthalmus). Type species: Quedius buphthalmus Erichson, fixed by monotypy (see Herman, 2001).

  • — Blackwelder, 1952: 82 (type species: cribripennis).

  • — Hammond, 1984: 205 (checklist of species of Borneo).

  • — Smetana, 1988: 314 (characters; species of Himalayan region).

  • — Smetana, 1995: 118 (revision of species of Taiwan; key to species of Taiwan).

  • — Smetana, 2000b: 329 (key to species of mainland China).

  • — Herman, 2001: 8 (discussion of availability of and type species for Bolitogyrus).

Cyrtothorax Kraatz, 1858: 366 (species included: sallei, nigricoxis, erythrurus). Type species: Cyrtothorax sallei Kraatz, fixed by subsequent designation by Scheerpeltz, 1974a: 175, 180 [Note: Blackwelder, 1952: 115 accepted C. buphthalmus Erichson as the type species designation by R. Lucas, 1920: 222, but that name was not among the originally included nominal species].

  • — Gemminger and Harold, 1868: 567 (catalog).

  • — Fauvel, 1878b: 163 [= 1878c: 83] (notes; key to species).

  • — Duvivier, 1883: 137 (catalog).

  • — Sharp, 1884: 340 (notes; Mexican and Central American species).

  • — Bernhauer and Schubert, 1916: 415 (world catalog; 9 species).

  • — Cameron, 1932: 277 (characters; key to species of British India).

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 115 (type species).

  • — Scheerpeltz, 1962b: 265 (characters).

  • — Scheerpeltz, 1974a: 175, 176 (type species designation; notes; key to species).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Smetana, 1988: 314 (synonym of Bolitogyrus).

  • — Smetana, 1995: 118 (synonym of Bolitogyrus).

bechyneorum Scheerpeltz, 1974a: 181, 189 (Cyrtothorax; Type locality: El Salvador: Gebiet des Vulcan San Vincente, Finca la Paz, Ahuachapan, Apaneca).

  • Distribution: El Salvador.

borneensis Cameron, 1942a: 138 (Cyrtothorax; Type locality: Martapura).

  • Distribution: Borneo.

bullatus Sharp, 1884: 340 (Cyrtothorax; Type locality: Panama, Boquete).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Panama).

  • Distribution: Panama.

buphthalmus Erichson, 1840: 534 (Quedius; Type locality: Mexico).

  • — Fauvel, 1878b: 164 [= 1878c: 84] (Cyrtothorax; characters; Mexico).

  • — Sharp, 1884: 340 (Cyrtothorax; characters; notes; Mexico; Guatemala).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; characters; Mexico; Guatemala; El Salvador).

  • Distribution: Mexico, Guatemala, El Salvador.

sallei Kraatz, 1858: 367 (Cyrtothorax; Type locality: Mexico).

  • — Gemminger and Harold, 1868: 567 (Cyrtothorax; synonym of buphthalmus).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; synonym of buphthalmus).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; synonym of buphthalmus).

nigricoxis Kraatz, 1858: 367 (Cyrtothorax; variety? of sallei; Type locality: Not cited).

caesareus Bernhauer, 1915f: 146 (Cyrtothorax; Type locality: Sarawak: Mts. Matang).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Borneo).

  • Distribution: Borneo.

carnifex Fauvel, 1878b: 165 [= 1878c: 85] (Cyrtothorax; Type locality: Cambodge).

  • — Bernhauer and Schubert, 1916: 415 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 180 (Cyrtothorax; characters; Cambodia).

  • Distribution: Cambodia.

costaricensis Wendeler, 1927: 8 (Cyrtothorax; Type locality: Costa Rica: Turrialba).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 181 (Cyrtothorax; characters; Costa Rica).

  • Distribution: Costa Rica.

cyanescens Sharp, 1884: 341 (Cyrtothorax; Type locality: Guatemala, Capetillo, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 181 (Cyrtothorax; characters; Guatemala).

  • Distribution: Guatemala.

cyanipennis Zheng, 1988: 306, 308 (Cyrtothorax; Type locality: China: Sichuan, Wolong, alt. 1830 m).

  • — Smetana, 2000b: 329 (Bolitogyrus; characters in key).

  • Distribution: China.

doesburgi Scheerpeltz, 1974a: 178, 187 (Cyrtothorax; Type locality: Java: Djawa; Gg. Moerie Tjolo).

  • Distribution: Indonesia.

electus Smetana and Zheng, 2001: 465 (Bolitogyrus; replacement name for elegans Smetana and Zheng).

  • Distribution: China.

elegans Smetana and Zheng, 2000: 59 (Bolitogyrus; [Preoccupied]; Type locality: China: N. Yunnan, Yulongshan mts., 2500–2800 m, Ganhaizi/Lijiang road).

  • — Smetana, 2000b: 329 (Bolitogyrus; characters in key).

  • — Smetana and Zheng, 2001: 465 (Bolitogyrus; synonym of electus).

elegans Cameron, 1937: 28 (Cyrtothorax; Type locality: Java: Batoerraden, G. Slamat).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

elegans Smetana and Zheng, 2000, see: electus Smetana and Zheng, 2001.

erythrurus Kraatz, 1858: 368 (Cyrtothorax; Type locality: Nova Granada).

  • — Fauvel, 1878b: 164 [= 1878c: 84] (Cyrtothorax; New Granada).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Colombia).

  • Distribution: Colombia.

fukienensis Scheerpeltz, 1974a: 177, 183 (Cyrtothorax; Type locality: Provinz Fukien in Südostchina, in Kuatun, Tschang-Sen).

  • — Smetana and Zheng, 2000: 55 (Bolitogyrus; characters; notes; China).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

fulgidus Sharp, 1884: 341 (Cyrtothorax; Type locality: Nicaragua, Chontales).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 182 (Cyrtothorax; characters; Nicaragua).

  • Distribution: Nicaragua.

kitawakii Smetana and Zheng, 2000: 56 (Bolitogyrus; Type locality: China: NE Sichuan, Chengkou Xian Daba Shan abv. Bashan, 1600–1900 m).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

nevermanni Scheerpeltz, 1974a: 181, 190 (Cyrtothorax; Type locality: Costarica: Carpintera).

  • Distribution: Costa Rica.

nigricoxis Kraatz, 1858, see: sallei Kraatz, 1858.

nigropolitus Smetana, 2000b: 327 (Bolitogyrus; Type locality: China: W. Sichuan: Ya'an Prefecture: Tianquan Co., Jiain Shan, Tal oberh. Labahe N. R. St., 57 km W Ya'an, 30°06N, 102°25E, 1800 m).

  • Distribution: China.

octomaculatus Cameron, 1937: 28 (Cyrtothorax; Type locality: Java: G. Tangkoeban Prahoe, alt. 4000–5000 ft.).

  • — Scheerpeltz, 1974a: 177 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

pictus Smetana and Zheng, 2000: 62 (Bolitogyrus; Type locality: China: Yun'an. okr. Puzenya, 900 m).

  • — Smetana, 2000b: 330 (Bolitogyrus; characters in key).

  • Distribution: China.

proximus Cameron, 1942a: 138 (Cyrtothorax; Type locality: Martapura).

  • Distribution: Borneo.

rufipennis Cameron, 1937: 27 (Cyrtothorax; Type locality: Java: Batoerraden, G. Slamat; Pengalengan, alt. 4000 ft.).

  • — Scheerpeltz, 1974a: 179 (Cyrtothorax; characters; Java).

  • Distribution: Indonesia.

rufomaculatus Shibata, 1979: 26 (Cyrtothorax; Type locality: near Lushan, about 1200 m alt., Nantou Hsien).

  • — Smetana, 1995: 118 (Bolitogyrus; characters; bionomics; Taiwan).

  • Distribution: Taiwan.

sallei Kraatz, 1858, see: buphthalmus Erichson, 1840.

salvini Sharp, 1884: 341 (Cyrtothorax; Type locality: Guatemala, Zapotes).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 183 (Cyrtothorax; characters; Guatemala).

  • Distribution: Guatemala.

signatus Cameron, 1932: 278 (Cyrtothorax; Type locality: Ceylon: Dikoya, alt 3800–4200 feet).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 178 (Cyrtothorax; characters; Ceylon).

  • Distribution: Sri Lanka.

strigifrons Wendeler, 1928a: 34 (Cyrtothorax; Type locality: Mexico: Jalapa).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 183 (Cyrtothorax; characters; Mexico).

  • Distribution: Mexico.

taiwanensis Hayashi, 1991a: 45 (Cyrtothorax; Type locality: Taiwan: Taitung Hsien: Taitung).

  • — Smetana, 1995: 121 (Bolitogyrus; characters; bionomics; Taiwan).

  • — Smetana, 1996g: 27 (Bolitogyrus; notes; Taiwan).

  • Distribution: Taiwan.

vietnamensis Scheerpeltz, 1974a: 178, 185 (Cyrtothorax; Type locality: Nordvietnam, Bao Lac, nördlich von Hanoi).

  • Distribution: Vietnam.

vulneratus Fauvel, 1878b: 165 [= 1878c: 85] (Cyrtothorax; Type locality: Cochinchine).

  • — Bernhauer and Schubert, 1916: 416 (Cyrtothorax; catalog).

  • — Cameron, 1932: 277 (Cyrtothorax; characters; Burma: Mt. Mooleyit. Cochin China).

  • — Scheerpeltz, 1933: 1426 (Cyrtothorax; catalog).

  • — Scheerpeltz, 1974a: 177 (Cyrtothorax; characters; Burma; Indochina).

  • — Biswas, 1986: 128 (Cyrtothorax; India).

  • — Smetana, 1988: 315 (Bolitogyrus; characters; Meghalaya; West Bengal; Nepal; Vietnam).

  • Distribution: Vietnam, Myanmar, Nepal, India.

Cheilocolpus [16 species; Neotropical Region]

Cheilocolpus Solier, 1849: 320 (species included: pyrostomus, angustatus, parvus). Type species: Staphylinus pyrostomus Solier, fixed by subsequent designation by Blackwelder, 1943: 99.

  • — Fauvel, 1874: 207 [= 1874: 433] (synonym of Philonthus).

  • — Bernhauer and Schubert, 1914: 326 (synonym of Philonthus).

  • — Cameron, 1932: 62 (synonym of Philonthus).

  • — Blackwelder, 1943: 399 (synonym of Philonthus; type species: pyrostomus).

  • — Blackwelder, 1952: 99 (type species).

  • — Smetana, 1958: 143 (synonym of Philonthus).

  • — Coiffait and Saiz, 1966: 386 (characters; key to Chilean species).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Saiz, 1971: 365, 387, 389 (characters; key to species of Chile; notes on biogeography; notes on phenetic relationships).

angustatus Solier, 1849: 320 (Staphylinus; [preoccupied]; Type locality: Chile).

  • — Kraatz, 1859a: 5 (Staphylinus; subgenus Cheilocolpus is a Philonthus; notes).

  • — Fairmaire and Germain, 1861: 433 (Staphylinus; characters; Chile).

  • — Fauvel, 1864a: 126 (Othius; characters; Chile).

  • — Fauvel, 1866a: 348 [= 1866b: 99] (Othius; characters; Chile).

  • — Germain, 1903: 417 (Heterothops; characters; Chile).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 397 (Cheilocolpus; characters; Chile).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 365, 370, 377 (Cheilocolpus; characters; Chile).

  • — Herman, 2001: 43 (Cheilocolpus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

  • Distribution: Chile.

cryptobioides Fairmaire and Germain, 1861: 439 (Lithocharis; Type locality: Aculco).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of angustatus).

  • — Coiffait and Saiz, 1966: 398 (Cheilocolpus; synonym of angustatus).

brevipennis Germain, 1903: 424 (Heterothops; Type locality: selvas subandinas de Chillan).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 392 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

centralis Saiz, 1971: 375 (Cheilocolpus; Type locality: Chile: El Arrayán, Santiago).

  • Distribution: Chile.

consors Fauvel, 1866a: 340 [= 1866b: 91] (Quedius; Type locality: Cordilliere de Santiago).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 393 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367, 370, 373 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

cryptobioides Fairmaire and Germain, 1861, see: angustatus Solier, 1849.

denticulatus Saiz, 1971: 379 (Cheilocolpus; Type locality: Chile: Bosque Relicto de Quinter, Valparaíso).

  • Distribution: Chile.

fulvicollis Fairmaire and Germain, 1861: 435 (Baptolinus; Type locality: Concepcion).

  • — Germain, 1903: 417 (Heterothops; characters; notes; Chile).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of angustatus).

  • — Coiffait and Saiz, 1966: 400 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 365, 370 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

impressifrons Solier, 1849: 316 (Staphylinus; Type locality: Chile: provincia de Santiago).

  • — Kraatz, 1859a: 5 (Philonthus; notes).

  • — Fairmaire and Germain, 1861: 430 (Philonthus; characters; Chile).

  • — Fauvel, 1864a: 125 (Philonthus; notes; Chile).

  • — Fauvel, 1866a: 343 [= 1866b: 94] (Philonthus; characters; Chile).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 396 (Cheilocolpus; characters; Chile).

  • — Coiffait and Saiz, 1968: 365 (Cheilocolpus; synonym of pyrostoma).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 367, 370, 374 (Cheilocolpus; characters; Chile).

  • — Frank, 1982: 34 (Cheilocolpus; list of parasites).

  • Distribution: Chile.

levigatus Saiz, 1971: 371 (Cheilocolpus; Type locality: Malacohuello, Sud du Chili).

  • Distribution: Chile.

magallanicus Saiz, 1971: 374 (Cheilocolpus; Type locality: Chile: Seno Otway, Rio El Ganso, Magallanes).

  • Distribution: Chile.

nanus Saiz, 1971: 377 (Cheilocolpus; Type locality: Chile: Nahuelbuta, Mallaco).

  • Distribution: Chile.

punctatus Saiz, 1971: 381 (Cheilocolpus; Type locality: Chile: Nahuelbuta, Malleco).

  • Distribution: Chile.

pygmaeus Saiz, 1971: 379 (Cheilocolpus; Type locality: Chile: Curacaví, Los Cerrillos, Santiago).

  • Distribution: Chile.

pyrostomus Solier, 1849: 320 (Staphylinus; Type locality: Chile: Illapel).

  • — Kraatz, 1859a: 5 (Staphylinus; subgenus Cheilocolpus is a Philonthus; notes).

  • — Fairmaire and Germain, 1861: 429 (Quedius; characters; Chile).

  • — Fauvel, 1864a: 125 (Quedius; notes; Chile).

  • — Fauvel, 1866a: 341 [= 1866b: 92] (Quedius; characters; Chile).

  • — Bernhauer and Schubert, 1916: 432 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 389 (Cheilocolpus; characters; Chile).

  • — Saiz, 1969: 3, 4 (Cheilocolpus; characters; collecting notes; Chile).

  • — Saiz, 1969a: 9 (Cheilocolpus; notes).

  • — Saiz, 1971: 367, 370, 371 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971a (Cheilocolpus; ecological notes).

  • — Saiz, Solervicens, and Ojeda, 1989: 52 (Cheilocolpus; collecting notes; habitus; Chile).

  • Distribution: Chile.

tenuis Saiz, 1971: 375 (Cheilocolpus; Type locality: Chile: Misituni, Tarapacá, 4000 m).

  • Distribution: Chile.

valdiviensis Coiffait and Saiz, 1966: 394 (Cheilocolpus; Type locality: Chili: Valdivia, 180 m, 40°05′S, 73°13′W).

  • — Saiz, 1971: 367, 370, 373 (Cheilocolpus; characters; Chile).

  • Distribution: Chile.

vicunianus Germain, 1903: 426 (Heterothops; Type locality: Requínoa).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Coiffait and Saiz, 1966: 392 (Cheilocolpus; characters; Chile).

  • — Saiz, 1971: 367 (Cheilocolpus; characters; Chile).

  • — Scheerpeltz, 1972: 23 (Heterothops; [Note: I assume Scheerpeltz overlooked transfer of this name to Cheilocolpus]; Argentina).

  • Distribution: Argentina, Chile.

Ctenandropus [2 species; Palaearctic, Oriental, Australian, and Oceanic Regions]

Ctenandropus Cameron, 1926b: 348 (species included: nigriceps). Type species: Ctenandropus nigriceps Cameron, fixed by original designation and monotypy.

  • — Cameron, 1932: 268 (characters).

  • — Scheerpeltz, 1933: 1425 (world catalog supplement).

  • — Blackwelder, 1952: 113 (type species: nigriceps).

  • — Scheerpeltz, 1962b: 264 (characters; type species: nigriceps).

  • — Smetana, 1988: 334 (characters; species of Himalayan region).

nigriceps Cameron, 1926b: 349 (Ctenandropus; Type locality: Asarori, Siwaliks; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Siwaliks; Asarori, Timli. Andaman Islands, Port Blair).

  • — Cameron, 1927c: 264 (Ctenandropus; Fiji; Sumatra).

  • — Cameron, 1932: 268 (Ctenandropus; characters; India; Andaman Islands; Malay Peninsula; Sumatra; Philippines).

  • — Scheerpeltz, 1933: 1425 (Ctenandropus; catalog).

  • — Cameron, 1933d: 392 (Ctenandropus; Sumatra).

  • — Cameron, 1933e: 83 (Ctenandropus; Society Islands).

  • — Cameron, 1936: 6 (Ctenandropus; Society Islands; Rapa).

  • — Coiffait, 1976: 218 (Ctenandropus; Tahiti).

  • — Legner and Moore, 1977: 175 (Ctenandropus; cited as nicriceps; found under bark).

  • — Smetana, 1988: 334 (Ctenandropus; lectotype designation: Asarori, Siwaliks; characters; Uttar Pradesh).

  • — Lecoq, 1990: 207 (Ctenandropus; Mascarene Islands).

  • Distribution: India, Andaman Islands, Malaysia, Indonesia, Philippines, Fiji, Society Islands, Rapa Islands.

magniceps Bernhauer, 1920: 17 (Heterothops; Type locality: Herberton).

  • — Lea, 1925: 231 (Heterothops; Australia).

  • — Bernhauer, 1928: 18 (Heterothops; Sumatra).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Cameron, 1933d: 392 (Heterothops; Sumatra).

  • — Smetana, 1988: 334 (Ctenandropus).

  • Distribution: Australia, Indonesia.

Euristus [1 species; Ethiopian Region]

Euristus Fauvel, 1899: 23 (species included: globus). Type species: Euristus globus Fauvel, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 409 (world catalog; 1 species).

  • — Blackwelder, 1952: 158 (type species: globus).

globus Fauvel, 1899: 24 (Euristus; Type locality: Gabon: Loango).

  • — Bernhauer and Schubert, 1916: 409 (Euristus; catalog).

  • Distribution: Gabon.

Euryporus [6 species; Palaearctic Oriental, and Australian Regions]

Euryporus Erichson, 1839a: 496 (replacement name for Pelecyphorus Nordmann). Type species: Oxyporus picipes Paykull, fixed by objective synonymy with Pelecyphorus.

  • — Heer, 1839: 282 (characters).

  • — Stephens, 1839: 384 (characters).

  • — Erichson, 1840: 553 (characters).

  • — Redtenbacher, 1849: 712 (characters).

  • — Schaum, 1852: 27 (in list of species; Europe).

  • — Lacordaire, 1854: 86 (characters; notes; in list of species).

  • — Fairmaire and Laboulbène, 1856: 545 (characters).

  • — Jacquelin du Val, 1856: 39 (characters).

  • — Kraatz, 1857d: 481 (characters).

  • — Redtenbacher, 1857: 205 (characters; species of Austria).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 26 (characters; type species: picipes).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 179 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — Wencker and Silbermann, 1866: 27 (in list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 566 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (in list of British species).

  • — Fauvel, 1874: 315 [= 1874a: 541] (characters).

  • — Redtenbacher, 1874: 196 (characters).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 803 [= 1877: 659] (characters).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1888: 223 (characters).

  • — Ganglbauer, 1895: 382 (character).

  • — Heyden, Reitter, and Weise, 1906: 176 (in list of species of Europe and Caucasus).

  • — Porta, 1907: 88 (key to Italian species).

  • — Reitter, 1908: 106 (characters).

  • — Petri, 1912: 80 (in list of species of Siebenbürgen).

  • — Johansen, 1914: 344 (characters).

  • — Bernhauer and Schubert, 1916: 414 (world catalog; 5 species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Porta, 1926: 118 (characters; key to species of Italy).

  • — Portevin, 1929: 335 (characters; key to French species).

  • — Roubal, 1930: 386 (catalog; Slovakia).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1424 (world catalog supplement).

  • — Normand, 1935: 372 (cited as Euriporus; in list of species of Tunisia).

  • — Kloet and Hincks, 1945: 167 (in list of British species).

  • — Tottenham, 1949: 376, 421 (type species: picipes Paykull; in checklist of British species).

  • — Blackwelder, 1952: 160 (type species: picipes).

  • — V. Hansen, 1952: 160 (characters; Danish species).

  • — Blackwelder, 1952: 160 (type species).

  • — Kocher, 1958: 156 (in checklist of species; Morocco).

  • — Smetana, 1958: 320 (characters; habitus of one species; type species: picipes; Czechoslovakian species).

  • — Scheerpeltz, 1962b: 264 (characters; type species: picipes).

  • — Székessy, 1963: 61 (characters).

  • — Palm, 1963b: 157 (characters; species of Sweden).

  • — Lohse, 1964: 204 (characters; central European species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Kasule, 1970: 57 (larval characters).

  • — Bordoni, 1973j: 713 (in list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (in checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 31 (in list of British species).

  • — Coiffait, 1978: 290 (characters; key to species of western Palaearctic region).

  • — Muona, 1979: 15 (in list of Scandinavian species).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 117 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 122 (in list of some species collected in Germany; collecting notes).

  • — Tóth, 1984: 94 (characters; habitus).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Silfverberg, 1992: 17 (in list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Siitonen, 1993: 230 (in list of Finnish species).

  • — Smetana, 1993b: 49 (in list of species of Czech Republic and Slovakia).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 30 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 31 (in list of species of Italy).

  • — M. Hansen, 1996: 108 (in list of Danish species).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (additions to list of Danish species).

Pelecyphorus Nordmann, 1837: 13 (species included: picipes; preoccupied by Dejean, 1834). Type species: Oxyporus picipes Paykull, fixed by monotypy.

  • — Erichson, 1839a: 496 (synonym of Euryporus).

  • — Lacordaire, 1854: 86 (synonym of Euryporus).

  • — Jacquelin du Val, 1856: 39 (synonym of Euryporus).

  • — Fauvel, 1874: 315 [= 1874a: 541] (synonym of Euryporus).

  • — Ganglbauer, 1895: 382 (synonym of Euryporus).

  • — Bernhauer and Schubert, 1916: 414 (synonym of Euryporus).

  • — Tottenham, 1949: 376 (synonym of Euryporus; type species: picipes).

  • — Blackwelder, 1952: 296 (synonym of Euryporus; type species: picipes).

  • — Smetana, 1958: 320 (synonym of Euryporus).

aeneiventris P. Lucas, 1846: 115 (Euryporus; Type locality: Algeria: environs du moulin pres Lacalle; Sicile).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Euryporus; Egypt; Algeria).

  • — Fauvel, 1886: 61 [= 1886a: 53] (Euryporus; Morocco; Algeria).

  • — Fauvel, 1897d: 310 (Euryporus; cited as Eurporus; Barbary).

  • — Fauvel, 1902b: 118 (Euryporus; Morocco; Algeria; Tunisia).

  • — Sainte-Claire Deville, 1906: 104 (Euryporus; Corsica).

  • — Porta, 1907: 100 (Euryporus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • — Normand, 1947a: 9 (Euryporus, cited as Euriporus; Tunisia).

  • — Kocher, 1958: 156 (Euryporus; Morocco).

  • — Scheerpeltz, 1961d: 132 (Euryporus; Italy).

  • — Scheerpeltz, 1965b: 406 (Euryporus; Sicily).

  • — Scheerpeltz, 1965c: 511 (Euryporus; Sicily).

  • — Coiffait, 1968c: 505 (Euryporus; Corsica).

  • — Coiffait, 1978: 293 (Euryporus; characters; Spain; Italy; Corsica; North Africa).

  • — Sparacio, 1995: 163 (Euryporus; characters; summary of distribution; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Euryporus; Italy).

  • Distribution: Egypt, Tunisia, Algeria, Morocco, Spain, Italy.

meridionalis Baudi, 1848: 133 (Euryporus; Type locality: Sardinia).

  • — Gemminger and Harold, 1868: 566 (Euryporus; synonym of aeneiventris).

  • — Fauvel, 1902b: 118 (Euryporus; synonym of aeneiventris).

  • — Porta, 1907: 100 (Euryporus; synonym of aeneiventris).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; synonym of aeneiventris).

  • — Coiffait, 1978: 293 (Euryporus; synonym of aeneiventris).

argentatus Fauvel, 1881: 164 (Euryporus; Type locality: Moeara Laboe, Sumatra).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • Distribution: Indonesia.

flavipes Fauvel, 1884, see: Pammegus.

meridionalis Baudi, 1848, see: aeneiventris P. Lucas, 1846.

multicavus Last, 1980: 154 (Euryporus; Type locality: New Guinea: Kiunga).

  • Distribution: New Guinea.

picipes Paykull, 1800: 426 (Oxyporus; Type locality: Suecia).

  • — Gyllenhal, 1810: 443 (Oxyporus; characters; Sweden).

  • — Olivier, 1811: 608 (Oxyporus; characters; Sweden).

  • — Nordmann, 1837: 14 (Pelecyphorus; Germany; Sweden).

  • — Erichson, 1839a: 496 (Euryporus; characters; Germany).

  • — Heer, 1839: 283 (Heterothops; characters; Switzerland).

  • — Stephens, 1839: 389 (Euryporus; characters; England).

  • — Erichson, 1840: 554 (Euryporus; characters; Sweden; Germany).

  • — Heer, 1841: 584 (Euryporus; Switzerland).

  • — Redtenbacher, 1849: 712 (Euryporus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 545 (Euryporus; characters; France).

  • — Kraatz, 1857d: 482 (Euryporus; characters; Germany).

  • — Redtenbacher, 1857: 205 (Euryporus; characters; Austria).

  • — Thomson, 1859: 26 (type species of Euryporus).

  • — Thomson, 1860: 180 (Euryporus; characters; Scandinavia).

  • — Fauvel, 1874: 315 [= 1874a: 541] (Euryporus; characters; France; Lapland; Scandinavia; Russia; Germany; Britain; Switzerland).

  • — Redtenbacher, 1874: 196 (Euryporus; characters; Austria).

  • — Seidlitz, 1875: 268 (Euryporus; characters; Baltic region).

  • — Fauvel, 1876a: 79 [= 1876: 264] (Euryporus; Spain).

  • — Mulsant and Rey, 1876: 806 [= 1877: 662] (Euryporus; characters; France).

  • — J. Sahlberg, 1876: 30 (Euryporus; Finland).

  • — Fowler, 1888: 223 (Euryporus; characters; Britain).

  • — Seidlitz, 1891: 395 (Euryporus; characters; Europe).

  • — Seidlitz, 1891a: 420 (Euryporus; characters; Europe).

  • — Ganglbauer, 1895: 383 (Euryporus; characters; north and middle Europe).

  • — Porta, 1907: 100 (Euryporus; characters; Italy).

  • — Reitter, 1909: 107 (Euryporus; characters; Germany).

  • — Johansen, 1914: 345 (Euryporus; characters; Denmark).

  • — Munster, 1923: 185 (Euryporus; Fennoscandia).

  • — Portevin, 1929: 336 (Euryporus; characters; France).

  • — Scheerpeltz, 1933: 1425 (Euryporus; catalog).

  • — V. Hansen, 1952: 160 (Euryporus; characters; Denmark).

  • — Smetana, 1958: 320 (Euryporus; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 204 (Euryporus; characters; central Europe).

  • — Horion, 1965: 250 (Euryporus; Russia; Finland; Sweden; Norway; Denmark; Baltics; Poland; Czechoslovakia; Hungary; Romania; Italy; Switzerland; France; Belgium; Holland; Germany; Austria).

  • — Kasule, 1970: 57 (Euryporus?; larval characters).

  • — Pope, 1977: 31 (Euryporus; Britain).

  • — Coiffait, 1978: 291 (Euryporus; characters; Europe; Scandinavia; Britain).

  • — Tóth, 1984: 95 (Euryporus; characters).

  • — Segers, 1986: 52 (Euryporus; checklist of species; Belgium).

  • — Lucht, 1987: 108 (Euryporus; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 49 (Euryporus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Euryporus; Italy).

  • — Sterrenburg and Schülke, 1997: 20 (Euryporus; Netherlands).

  • — Owen, 1999: 10 (Euryporus; collecting notes; Britain).

  • — Eyre, Luff, and Lott, 2000: 33 (Euryporus; Britain).

  • Distribution: Europe, Russia.

princeps Wollaston, 1864: 561 (Euryporus; Type locality: Grand Canary: Teror).

  • — Wollaston, 1865: 485 (Euryporus; habitat; Canary Islands).

  • — Fauvel, 1897d: 310 (Euryporus; cited as Eurporus; Canaries).

  • — Fauvel, 1902b: 118 (Euryporus; Canaries).

  • — Bernhauer and Schubert, 1916: 414 (Euryporus; catalog).

  • — Coiffait, 1978: 294 (Euryporus; characters; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 187 (Euryporus; checklist of species; Canary Islands).

  • Distribution: Canary Islands.

puncticollis Erichson, 1840, see: Tympanophorus.

warisensis Last, 1987: 51 (Euryporus; Type locality: Irian: Warris, S. of Jayapura, 450–500 m).

  • Distribution: New Guinea.

Haematodes [2 species; Neotropical Region]

Haematodes Laporte, 1835: 112 (species included: bicolor). Type species: Haematodes bicolor Laporte, fixed by monotypy.

  • — Erichson, 1839b: 340 (characters).

  • — Laporte, 1840: 175 (cited as Hematodes; characters).

  • — Lacordaire, 1854: 72 (characters; notes; list of species).

  • — Gemminger and Harold, 1868: 573 (catalog).

  • — Lynch, 1884: 115 (characters).

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 2 species).

  • — Scheerpeltz, 1933: 1420 (world catalog supplement).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 179 (type species: bicolor).

Platycnemus Nordmann, 1837: 135 (species included: lateritius). Type species: Platycnemus lateritius Nordmann, fixed by monotypy.

  • — Lacordaire, 1854: 72 (synonym of Haematodes).

  • — Lynch, 1884: 115 (synonym of Haematodes).

  • — Bernhauer and Schubert, 1916: 410 (synonym of Haematodes).

  • — Blackwelder, 1952: 312 (type species: lateritius).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place Platycnemus on the Official List of Generic Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of Generic Names in Zoology).

Platytoma Chevrolat, 1847: 263 ([Note: Nomen nudum (Article 11.6, 12.1), published without description or species and as a synonym of Haematodes. Blackwelder, 1952: 314 cited it as an isogenotypic synonym of Haematodes]).

  • — Lacordaire, 1854: 72 (synonym of Haematodes).

  • — Lynch, 1884: 115 (synonym of Haematodes).

bicolor Laporte, 1835: 113 (Haematodes; Type locality: Buenos Ayres).

  • — Blanchard, 1842: 76 (Haematodes; Uruguay; Brazil).

  • — Erichson, 1839b: 340 (Haematodes; characters; Brazil).

  • — Laporte, 1840: 175 (Haematodes; characters; Argentina).

  • — Lynch, 1884: 116 (Haematodes; characters; Argentina; Brazil; Uruguay).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; catalog).

  • — Scheerpeltz, 1933: 1420 (Haematodes; catalog).

  • Distribution: Brazil, Uruguay, Argentina.

lateritius Nordmann, 1837: 135 (Platycnemus; Type locality: Brasilia meridionalis).

  • — Blanchard, 1842: 76 (Haematodes; synonym of bicolor).

  • — Erichson, 1839b: 340 (Haematodes; synonym of bicolor).

  • — Lynch, 1884: 116 (Haematodes; synonym of bicolor).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; synonym of bicolor).

  • — Newton, 1995: 51 (requested that the International Commission of Zoological Nomenclature place lateritius on the Official List of Specific Names in Zoology).

  • — ICZN, 1996: 215 (placed on Official List of specific Names in Zoology).

lateritius Nordmann, 1837, see: bicolor Laporte, 1835.

tenuipes Kraatz, 1858: 363 (Haematodes; Type locality: Brasilia).

  • — Bernhauer and Schubert, 1916: 410 (Haematodes; catalog).

  • Distribution: Brazil.

Heinzia [1 species; Palaearctic Region]

Heinzia Korge, 1971a: 33, 36 (species included; variabilis). Type species: Heinzia variabilis Korge, fixed by original designation and monotypy.

  • — Smetana, 1977b: 181 (characters).

  • — Coiffait, 1978: 284 (characters).

variabilis Korge, 1971a: 33, 37 (Heinzia; Type locality: Anatolia bor.: Kaçkar-Daglari bei Iliça, Tal von Ardesen, 1000–1600 m).

  • — Coiffait, 1978: 284 (Heinzia; characters; Anatolia).

  • — Solodovnikov, 1998a: 16 (Heinzia; notes; Russia, Krasnodar territory).

  • Distribution: Turkey, Russia.

Hemiquedius [1 species; Nearctic Region]

Hemiquedius Casey, 1915: 397, 399 (species included: ferox). Type species: Quedius ferox LeConte, fixed by original designation and monotypy.

  • — Casey, 1915: 397 (type species: Quedius ferox).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 182 (type species: ferox).

  • — Smetana, 1971: 238 (valid genus; characters).

  • — Moore and Legner, 1975: 53 (catalog; North America north of Mexico).

  • — Smetana, 1977b: 181 (characters).

  • — Moore and Legner, 1979: 45 (characters; notes).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Downie and Arnett, 1996: 394 (characters).

  • — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 392 (1 Nearctic species; notes; characters in key).

ferox LeConte, 1878a: 388 (Quedius; Type locality: Enterprise, Florida; [Note: See lectotype designation by Smetana, 1971]. Also cited from Louisiana; Canada; Massachusetts).

  • — Horn, 1878: 166 (Quedius; characters; Massachusetts to Canada, Florida, Louisiana).

  • — Blatchley, 1910: 372 (Quedius; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1965: 35 (Quedius; subgenus Hemiquedius; Nova Scotia).

  • — Smetana, 1971: 239 (Hemiquedius; lectotype designation: Enterprise, Florida; characters; Ontario; Nova Scotia; Quebec; Alabama; Florida; Georgia; Illinois; Indiana; Iowa; Louisiana; Maine; Maryland; massachusetts; Michigan; Missouri; New Jersey; New York; North Carolina; Pennsylvania; Wisconsin).

  • — Smetana, 1971a: 1848 (Hemiquedius; notes).

  • — Smetana, 1978a: 831 (Hemiquedius; collecting notes; Florida; Illinois; Texas).

  • — Frank, 1982: 36 (Hemiquedius; list of parasites).

  • — Smetana, 1990d: 101 (Hemiquedius; Arkansas; Georgia; Louisiana; Mississippi; New Hampshire; New York; Tennessee).

  • — Campbell and Davies, 1991: 122 (Hemiquedius; checklist of species; Canada).

  • — Downie and Arnett, 1996: 394 (Hemiquedius; characters; USA).

  • Distribution: Canada, USA.

Heterothops [149 species; Neotropical, Nearctic, Palaearctic, Oriental, and Australian Regions]

Heterothops Stephens, 1829a: 23 (species included: binotatus and two unavailable names). Type species: Staphylinus binotatus Gravenhorst, fixed by monotypy.

  • — Stephens, 1829: 284 (catalog; Britain).

  • — Stephens, 1832: 200 (characters).

  • — Stephens, 1833: 256 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 480 (characters).

  • — Heer, 1839: 280 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Erichson, 1840: 515 (characters).

  • — Redtenbacher, 1849: 711 (characters).

  • — Schaum, 1852: 26 (list of species; Europe).

  • — Lacordaire, 1854: 83 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 532 (characters).

  • — Jacquelin du Val, 1856: 38 (characters).

  • — Kraatz, 1857d: 483 (characters).

  • — Redtenbacher, 1857: 198 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 26 (characters; type species: dissimilis).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 180 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (characters).

  • — Wencker and Silbermann, 1866: 27 (cited as Heterotops; list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 567 (catalog).

  • — Jacquelin du Val, 1868: 71 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 309 [= 1874a: 535] (characters; key to species).

  • — Redtenbacher, 1874: 196 (characters; key to Austrian species).

  • — Fauvel, 1875a: XXXIV [= 1875b: 236] (catalog).

  • — Mulsant and Rey, 1876: 808 [= 1877: 664] (characters; key to species).

  • — Horn, 1878: 152 (characters; key for North American species).

  • — Heyden, 1880: 70 (list of species of Siberian region).

  • — Duvivier, 1883: 136 (catalog).

  • — Sharp, 1884: 323 (notes; Mexican and Central American species).

  • — Lynch, 1884: 122 (characters).

  • — Sharp, 1888: 224 (characters; key to British species).

  • — Seidlitz, 1891: 400 (key to some European species).

  • — Seidlitz, 1891a: 426 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 40 (list of species of Siberian region).

  • — Ganglbauer, 1895: 384 (adult and larval characters; key to middle European species).

  • — Heyden, 1896: 31 (list of species of Siberian region).

  • — Fauvel, 1897d: 310 (catalog of species of Barbary; Madeira; Canaries).

  • — Germain, 1903: 414 (characters; notes).

  • — Heyden, Reitter, and Weise, 1906: 176 (list of species of Europe and Caucasus).

  • — Porta, 1907: 89 (key to Italian species).

  • — Reitter, 1908: 106, 107 (characters; key to species).

  • — Blatchley, 1910: 368 (characters).

  • — Petri, 1912: 80 (list of species of Siebenbürgen).

  • — Johansen, 1914: 345 (characters; key to species).

  • — Bernhauer and Schubert, 1916: 411 (world catalog; 48 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Munster, 1923: 184 (characters; key to Scandinavian species).

  • — Winkler, 1925: 387 (catalog for Palaearctic region).

  • — Cameron, 1925: 71 (catalog of Indian species).

  • — Porta, 1926: 118 (characters; key to species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 336 (key to French species).

  • — Roubal, 1930: 385 (catalog; Slovakia).

  • — Cameron, 1932: 265 (characters; key to species of British India).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1421 (world catalog supplement).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Paulian, 1941: 254 (larval characters).

  • — Blackwelder, 1943: 464 (characters; type species: binotatus; key to species of West Indies).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 167 (list of British species).

  • — Tottenham, 1949: 376, 421 (type species: binotatus Gravenhorst; notes; checklist of British species).

  • — Blackwelder, 1952: 186 (type species: binotatus).

  • — V. Hansen, 1952: 158 (characters; key to Danish species).

  • — V. Hansen, 1952: 158 (characters; key to Danish species).

  • — Hatch, 1957: 214, 230 (characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 314 (characters; habitus of one species; type species: binotatus; key to Czechoslovakian species).

  • — Kocher, 1958: 156 (checklist of species; Morocco).

  • — Smetana, 1959a: 215 (list of species collected in Albania).

  • — Scheerpeltz, 1962b: 263 (characters; type species: binotatus).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Székessy, 1963: 59 (characters).

  • — Palm, 1963b: 154 (characters; key to species of Sweden).

  • — Lohse, 1964: 203 (characters; key to central European species).

  • — Pototskaia, 1967: 79 (larval characters).

  • — Smetana, 1967g: 466, 470 (natural history notes; key to central European species).

  • — Scheerpeltz, 1968a: 86 (catalog of Austrian species).

  • — Kasule, 1970: 57 (larval characters).

  • — Smetana, 1971: 14 (characters; key to species of America north of Mexico; species group classification).

  • — Smetana, 1971a: 1838 (additions to key to species).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Last, 1975: 425 (characters; species of New Guinea).

  • — Moore and Legner, 1975: 54 (catalog; North America north of Mexico).

  • — Moore and Legner, 1976: 539 (includes seashore species; notes).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 30 (list of British species).

  • — Coiffait, 1978: 300 (characters; key to species of western Palaearctic region).

  • — Smetana, 1978a: 821 (key to North American species).

  • — Topp, 1978: 312 (larval characters).

  • — Israelson, 1979a: 261 (discussion of characters of punctation, chaetotaxy, and aedeagus).

  • — Moore and Legner, 1979: 46 (characters; notes).

  • — Uhlig, 1979: 255 (some species collected in Germany).

  • — Uhlig, Vogel, and Sieber, 1980: 248 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 113 (catalog; Poland).

  • — Uhlig and Vogler, 1981: 121 (list of some species collected in Germany; habitat and collecting notes).

  • — Tóth, 1984: 98 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 115 (checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 52 (checklist of species; Belgium).

  • — Drugmand 1987b (comments on distribution of Belgian species).

  • — Ádám, 1987: 152 (list of two species with collecting notes; Hungary).

  • — Smetana, 1988: 318 (characters; key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 169 (supplement to key to species of Lohse, 1964).

  • — Angelini, 1991: 191 (list of species from high elevations of Calabria, Italy).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Campbell and Davies, 1991: 121 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Ito, 1994a: 179 (key to Japanese species).

  • — Smetana, 1995: 121 (revision of species of Taiwan; key to species of Taiwan).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 30 (additional records to Danish list of species).

  • — Ciceroni and Zanetti, 1995: 31 (list of species of Italy).

  • — M. Hansen, 1996: 108 (list of Danish species).

  • — Cho, 1996b: 108 (key to species of Korea).

  • — Downie and Arnett, 1996: 388 (characters; key to species of Northeastern North America).

  • — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 74 (list of species collected in 1997; Denmark).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (18 Nearctic species; notes; characters in key).

Trichopygus Nordmann, 1837: 137 (species included: subuliformis, dissimilis, nitens, pumilio). Type species: Tachyporus dissimilis Gravenhorst, fixed by subsequent designation by Tottenham, 1939d: 229.

  • — Erichson, 1839a: 480 (synonym of Heterothops).

  • — Lacordaire, 1854: 83 (synonym of Heterothops).

  • — Jacquelin du Val, 1856: 38 (synonym of Heterothops).

  • — Fauvel, 1874: 309 [= 1874a: 535] (synonym of Heterothops).

  • — Lynch, 1884: 122 (synonym of Heterothops).

  • — Ganglbauer, 1895: 386 (synonym of Heterothops).

  • — Bernhauer and Schubert, 1916: 411 (synonym of Heterothops).

  • — Cameron, 1932: 265 (synonym of Heterothops).

  • — Tottenham, 1939d: 229 (type species designation: dissimilis).

  • — Blackwelder, 1943: 464 (synonym of Heterothops; type species: dissimilis).

  • — Tottenham, 1949: 376 (synonym of Heterothops; type species: dissimilis).

  • — Blackwelder, 1952: 396 (type species).

  • — Smetana, 1958: 314 (synonym of Heterothops).

  • — Smetana, 1971: 14 (synonym of Heterothops).

  • — Coiffait, 1978: 301 (synonym of Heterothops).

acuminatus Fairmaire, 1860, see: Quedius.

aeneiventris Smetana, 1971a: 1836 (Heterothops; Type locality: Arizona: Cochise Co.: Huachuca Mts., Ramsey Canyon, 5200 ft.).

  • — Smetana, 1978a: 818 (Heterothops; collecting notes; Arizona).

  • Distribution: USA.

amabilis Last, 1975: 428 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

angusticeps Reitter, 1891: 18 (Heterothops; Type locality: Kasalinsk).

  • — Eppelsheim, 1892: 329 (Heterothops; Turkestan).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog; Turkestan; Transcaspian region).

  • — Coiffait, 1978: 317 (Heterothops; characters; Transcaspian region).

  • Distribution: Kazakhstan, Uzbekistan.

apicicornis Bernhauer, 1917h: 86 (Heterothops; Type locality: West-Kolumbien: Umgebung von Cali am Rio Cauca, St. Antonio, 2000 m).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Colombia.

apterus Lea, 1925: 232 (Heterothops; Type locality: Tasmania: Cradle Mountain).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Australia.

aquitanicus Coiffait, 1977d: 295 (Heterothops; Type locality: Arcachon, Plage du Pilat).

  • — Coiffait, 1978: 315 (Heterothops; characters; France).

  • Distribution: France.

argus Wendeler, 1956b: 263 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

arizonicus Smetana, 1971a: 1834 (Heterothops; Type locality: Arizona: Cochise Co.: Chiricahua Mts., Portal, 5000 ft.).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; Arizona).

  • Distribution: USA.

armeniacus Coiffait, 1977c: 142 (Heterothops; Type locality: Erevan, Kamaker, Arménie soviétique).

  • — Coiffait, 1978: 312 (Heterothops; characters; Armenia).

  • — Tronquet, 1981: 71 (Heterothops; Afghanistan).

  • Distribution: Armenia, Afghanistan.

asperatus Smetana, 1971: 34 (Heterothops; Type locality: California: Samoa, Humboldt Co. Also cited from British Columbia).

  • — Smetana, 1971b: 1 (Heterothops; Washington).

  • — Orth, Moore, and Fisher, 1978: 188 (Heterothops; characters; collecting notes; California).

  • Distribution: Canada, USA.

baicalensis Fauvel, 1900d: 233 (Heterothops; Type locality: Riviere Sludianka, Baical; Daourie).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; catalog).

  • — Smetana, 1967d: 213 (Heterothops; Mongolia).

  • — Smetana 1975c: 176 (Heterothops; Mongolia).

  • — Coiffait, 1978: 318 (Heterothops; characters; central Asia).

  • Distribution: Russia, Mongolia.

balthasari Smetana, 1967g: 468 (Heterothops; Type locality: Böhmen, Böhmisches Mittelgebirge, Berg Raná).

  • — Coiffait, 1978: 312 (Heterothops; cited as balthazari; characters; central Europe).

  • — Tóth, 1984: 100 (Heterothops; cited as balthazari; characters).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • Distribution: Czech Republic.

basicornis Bernhauer, 1917h: 85 (Heterothops; Type locality: Bolivien: Yuracares).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Bolivia.

bicolor Smetana, 1975c: 173 (Heterothops; Type locality: Mongolie: Bajanchongor aimak: Oase Echin gol, cca 90 km NO vom Grenzposten Caganbulag, 950 m).

  • Distribution: Mongolia.

bimaculatus Fauvel, 1878g: 557 (Heterothops; Type locality: Nouvelle-Galles du Sud, Sydney).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • Distribution: Australia.

binotatus Gravenhorst, 1802: 28 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 316 (Staphylinus; characters; Germany).

  • — Gyllenhal, 1810: 312 (Staphylinus; synonym of subuliformis Gravenhorst).

  • — C. Sahlberg, 1830: 319 (Staphylinus; synonym of subuliformis Gravenhorst).

  • — Stephens, 1833: 257 (Heterothops; characters; England).

  • — Westwood, 1838a: 16 (Heterothops; type species).

  • — Stephens, 1839: 401 (Heterothops; characters; England).

  • — Kraatz, 1857d: 485 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 985 (Heterothops; characters; Austria).

  • — Fauvel, 1874: 310 [= 1874a: 536] (Heterothops; characters; France; Russia; Germany; Britain; Italy; Corsica; Sardinia; Cyprus; Caramania).

  • — Redtenbacher, 1874: 197 (Heterothops; characters).

  • — Fauvel, 1876a: 78 [= 1876: 263] (Heterothops; Algeria; Tunisia).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Heterothops; Algeria; Tunisia).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Heterothops; Algeria).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; coast of North Sea and Baltic Sea; Mediterranean coast).

  • — Fauvel, 1902b: 118 (Heterothops; Algeria; Tunisia).

  • — Porta, 1907: 101 (Heterothops; characters; Italy).

  • — Sainte-Claire Deville, 1906: 103 (Heterothops; Corsica).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 346 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Munster, 1923: 184 (Heterothops; Fennoscandia).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1421 (Heterothops; catalog).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Scheerpeltz, 1960d: 136 (Heterothops; Portugal).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Norway).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Horion, 1965: 245 (Heterothops; Syria; Cyprus; Turkey; Greece; Dalmatia; Italy; France; Spain; Tunisia; Algeria; Spain; Portugal; Ireland; Britain; Belgium; Holland; Norway; Denmark; Sweden; Germany).

  • — Smetana, 1967g: 470 (Heterothops; characters).

  • — Szujecki, 1969a: 258 (Heterothops; notes; Poland).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 308 (Heterothops; characters; Europe; Britain; Scandinavia; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 100 (Heterothops; characters).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 169 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • — Owen, 1997a: 149 (Heterothops; collecting sites; Ireland).

  • — Herman, 2001: 25 (Heterothops; synonymy).

  • Distribution: Tunisia, Algeria, Europe, Turkey, Russia.

sericans Stephens, 1832: 221 (Quedius; Type locality: Suffolk).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

holmensis Stephens, 1833: 257 (Heterothops; Type locality: on the sea-shore near Holm, in Norfolk).

  • — Stephens, 1839: 401 (Heterothops; variety of binotatus; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

  • — Gemminger and Harold, 1868: 566 (Heterothops; synonym of binotatus).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of binotatus).

  • — Porta, 1907: 101 (Heterothops; synonym of binotatus).

  • — Bernhauer and Schubert, 1916: 411 (Heterothops; synonym of binotatus).

  • — Coiffait, 1978: 309 (Heterothops; synonym of binotatus).

kirbiella Stephens, 1833: 257 (Heterothops; Type locality: Sussex).

  • — Stephens, 1839: 401 (Heterothops; variety of binotatus; characters; England).

  • — G. Waterhouse, 1858: 22 (Heterothops; synonym of binotatus).

  • — Gemminger and Harold, 1868: 567 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Ganglbauer, 1895: 387 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Porta, 1907: 101 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; cited as kirbyella; synonym of binotatus).

  • — Coiffait, 1978: 309 (Heterothops; cited as kirbyellus; synonym of binotatus).

binotatus Erichson, 1840: 516 (Heterothops; [preoccupied]; [Note: Bernhauer and Schubert, 1916: 412 listed Erichson's reference for binotatus as a subsequent citation to binotatus Gravenhorst. Erichson attributed binotatus to no one so he evidently described it as new but it may be the same as binotatus Gravenhorst]; Type locality: Anglia).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Peyron, 1858: 428 (Heterothops; Turkey).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 811 [= 1877: 667] (Heterothops; characters; France).

  • — Fowler, 1888: 224 (Heterothops; characters; Britain).

  • — Herman, 2001: 25 (Heterothops; synonym of binotatus).

binotatus Erichson, 1840, see: binotatus Gravenhorst, 1802.

bonariensis Lynch, 1884: 123 (Heterothops; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Argentina.

boops Bernhauer, 1910b: 383 (Heterothops; Type locality: Mexiko: Cuernavaca).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Mexico.

brevipennis Germain, 1903, see: Cheilocolpus.

brunnipennis Kiesenwetter, 1858: 56 (Heterothops; Type locality: bei Nauplia [bei Tirynth]; [Note: See lectotype designation by Israelson, 1979a]).

  • — Baudi, 1870: 383 (Heterothops; cited as brunneipennis; characters; Sardinia).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Fowler, 1888: 225 (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; cited as brunneipennis; variety of dissimilis).

  • — Scheerpeltz, 1933: 1422 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Gridelli, 1934: 82 (Heterothops; cited as brunneipennis; variety of dissimilis; Karakoram region).

  • — C. Koch, 1936a: 76 (Heterothops; cited as brunneipennis; Libya).

  • — Normand, 1947a: 9 (Heterothops; cited as brunneipennis; subspecies of dissimilis; Tunisia).

  • — Cameron, 1951: 283 (Heterothops; cited as brunneipennis; Egypt).

  • — Smetana, 1958: 318 (Heterothops; cited as brunneipennis; aberration of dissimilis).

  • — Scheerpeltz, 1963: 14 (Heterothops; cited as brunneipennis; Afghanistan).

  • — Focarile, 1964: 57 (Heterothops; cited as valid species; Italy).

  • — Coiffait, 1973a: 275 (Heterothops; cited as brunneipennis; variety of dissimilis; Morocco).

  • — Coiffait, 1978: 314 (Heterothops; cited as brunneipennis; variety of dissimilis; characters).

  • — Muona, 1979: 15 (Heterothops; cited as brunneipennis; synonym of dissimilis).

  • — Israelson, 1979a: 263, 265 (Heterothops; lectotype designation; valid species distinct from dissimilis; Swedish record of brunnipennis are minutus; notes; discussion; characters).

  • Distribution: Egypt, Libya, Europe, Afghanistan, “Karakorum region”.

brunneus Smetana, 1971: 32 (Heterothops; Type locality: California: Antelope Spgs., Inyo Co.).

  • — Smetana, 1978a: 821 (Heterothops; collecting notes; California).

  • Distribution: USA.

buettikeri Coiffait, 1981b: 238 (Heterothops; Type locality: Arabie Saoudite: Hofuf).

  • — Gusarov, 1997: 288 (Heterothops; Saudi Arabia).

  • Distribution: Saudi Arabia.

buluensis Last, 1975: 438 (Heterothops; Type locality: New Guinea: Garaina, Bulu Valley).

  • Distribution: New Guinea.

burdurensis Štourač, 2000: 67 (Heterothops; Type locality: Turquie: Burdur, Çerçin lac Burdur, 870 m).

  • Distribution: Turkey.

californicus LeConte, 1863, see: fusculus LeConte, 1863.

campbelli Smetana, 1971: 22 (Heterothops; Type locality: Illinois: Pine Hills Field Sta., Union Co.).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; North Carolina; Oklahoma; Texas).

  • — Smetana, 1981b: 632 (Heterothops; collecting notes; Florida; Illinois; Kentucky; Michigan).

  • — Smetana, 1990d: 96 (Heterothops; Arkansas; Louisiana; Mississippi).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • Distribution: USA.

canariensis Israelson, 1979a: 267 (Heterothops; Type locality: Los Tiles, Gran Canaria, Canary Islands).

  • — Wollaston, 1864: 562 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; notes; habitat; Canary Islands).

  • — Wollaston, 1865: 485 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; habitat; notes; Canary Islands).

  • — Coiffait, 1954: 167 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 267]; Canary Islands).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (Heterothops; checklist of species; Canary Islands).

  • — Assing, 1999a: 346 (Heterothops; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Heterothops; Canary Islands).

  • Distribution: Canary Islands.

carbonatus Fall, 1901: 220 (Heterothops; Type locality: maritime southern California).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Hatch, 1957: 231 (Heterothops; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 23 (Heterothops; characters; British Columbia; Arizona; California; Idaho; Oregon; Washington).

  • — Smetana, 1971a: 1833 (Heterothops; collecting notes; Arizona).

  • — Smetana, 1971b: 1 (Heterothops; British Columbia).

  • Distribution: Canada, USA.

castaneus Lea, 1925: 238 (Heterothops; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

cephalota Coiffait, 1977d: 297 (Heterothops; Type locality: Darvaz, Noulvantch, Tadjikistan).

  • — Coiffait, 1978: 313 (Heterothops; characters; Transcaucasia).

  • Distribution: Tajikistan, “Transcaucasus”.

clarki Lea, 1925: 237 (Heterothops; Type locality: Western Australia: Swan River).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Australia.

cognatus Sharp, 1874: 20 (Heterothops; Type locality: Japan).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Watanabe, 1961: 354 (Heterothops; Izu Islands).

  • — Nakane, 1963a: 94 (Heterothops; characters; habitus photograph; Japan).

  • — Watanabe and Shibata, 1972: 65 (Heterothops; Yaku-shima Island).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Heterothops; Korea).

  • — Watanabe, 1993d (Heterothops; Sarushima Island).

  • — J. Li, 1993: 38 (Heterothops; China).

  • — Cho, 1996b: 108 (Heterothops; characters; Korea; also listed from China).

  • — Watanabe, 1998: 314 (Heterothops; Japan).

  • Distribution: Japan, Korea, China.

confertus Smetana, 1971: 25 (Heterothops; Type locality: Arizona: Madera Cn. S. Rita Mts., 6200 ft., S. Cruz Co.).

  • — Smetana, 1971a: 1834 (Heterothops; collecting notes; Arizona).

  • — Smetana, 1978a: 818 (Heterothops; collecting notes; Texas).

  • Distribution: USA.

conformis Smetana, 1971: 30 (Heterothops; Type locality: British Columbia: 8 mi. W. Creston. Also cited from Arizona, California, Colorado, Idaho, Nevada, New Mexico, Oregon, Washington).

  • — Smetana, 1971a: 1834 (Heterothops; collecting notes; California).

  • — Smetana, 1971b: 1 (Heterothops; British Columbia).

  • — Smetana, 1973: 1422 (Heterothops; California; Idaho; Oregon).

  • — Smetana, 1978a: 819 (Heterothops; collecting notes; Arizona; California).

  • Distribution: Canada, USA.

conticens† Scudder, 1900: 48 (Heterothops; Type locality: Florissant, Colorado).

  • — Handlirsch, 1907: 723 (Heterothops; cited as conticeus; catalog; Miocene; Colorado).

  • Distribution: USA.

conviva Smetana, 1967d: 212 (Heterothops; Type locality: Mongolei: Chentej aimak, 20 km SW von Somon Batnorov, 1000 m).

  • — Smetana 1975c: 176 (Heterothops; Mongolia).

  • Distribution: Mongolia.

crisbus Last, 1975: 436 (Heterothops; Type locality: Okasa, North Fore census division, Okapa district, 3000 ft.).

  • Distribution: New Guinea.

discoideus Fairmaire and Germain, 1861, see: Loncovilius.

dissimilis Gravenhorst, 1802: 125 (Tachyporus; Type locality: Brunsvigae).

  • — Gravenhorst, 1806: 23 (Tachinus; characters; Germany).

  • — Erichson, 1839a: 480 (Heterothops; characters; Germany).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 517 (Heterothops; characters; Germany; Sweden).

  • — Kolenati, 1846: 21 (Heterothops; Caucasus; Iberia).

  • — P. Lucas, 1846: 113 (Heterothops; notes; Algeria).

  • — Jacobsen, 1849: 707 (Heterothops; characters; Denmark).

  • — Hochhuth, 1849: 146 (Heterothops; characters; Lenkoran).

  • — Redtenbacher, 1849: 712 (Heterothops; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Jacquelin du Val, 1857: pl. 15, fig. 74 (Heterothops; [Note: Misidentification: Bernhauer and Schubert, 1916: 412 cited binotatus Jacquelin du Val as a synonym of dissimilis; Jacquelin du Val did not describe the species as new, he attributed it to Erichson]; illustration).

  • — Kraatz, 1857d: 485 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Thomson, 1859: 26 (type species of Heterothops).

  • — Thomson, 1860: 180 (Heterothops; [Note: Misidentification: Bernhauer and Schubert, 1916: 412 cited praevius Thomson as a synonym of dissimilis; Thomson did not describe the species as new, he attributed it to Erichson]; characters; Scandinavia).

  • — Thomson, 1860: 181 (Heterothops; characters; Scandinavia).

  • — Hochhuth, 1862: 43 (Heterothops; characters; Caucasus; Altai).

  • — Gredler, 1863: 99 (Heterothops; Austria).

  • — Wollaston, 1871: 298 (Heterothops; Madeira and Canary Islands).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; characters; France; Madeira and Canary Islands; Barbary; Caucasus; Transcaucasus; Daghestan; Altai region; Siberia).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 823 [= 1877: 679] (Heterothops; characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fauvel, 1878: 130 [= 1878a: 50] (Heterothops; Morocco; Algeria; Tunisia).

  • — Fauvel, 1886: 60 [= 1886a: 52] (Heterothops; North Africa).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Palaearctic region).

  • — Fauvel, 1902b: 118 (Heterothops; Barbary; Egypt; Madeira; Canaries).

  • — Sainte-Claire Deville, 1906: 104 (Heterothops; Corsica).

  • — Porta, 1907: 103 (Heterothops; characters; Italy).

  • — Peyerimhoff, 1907: 14 (Heterothops; Sinai).

  • — Montandon, 1908: 80 (Heterothops; Romania).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 347 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Munster, 1923: 185 (Heterothops; Fennoscandia).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Gridelli, 1930: 78 (Heterothops; Libya).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Uyttenboogaart, 1937: 82 (Heterothops; characters; notes).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Smetana, 1958: 318 (Heterothops; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1961: 7 (Heterothops; Iran).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Scheerpeltz, 1965c: 510 (Heterothops; Sicily).

  • — Horion, 1965: 247 (Heterothops; widespread in southern and central Europe; Denmark; Britain; Sweden; Finland).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Smetana, 1967i: 557 (Heterothops; Turkey).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Coiffait, 1973a: 275 (Heterothops; Morocco).

  • — Osella and Zanetti, 1975: 113 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 314 (Heterothops; characters; Europe; Britain; Scandinavia; Madeira; Canaries; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 102 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Nowosad, 1990: 124 (Heterothops; collected from mole nest; notes; Poland).

  • — Maachi, 1991: 251 (Heterothops; Morocco).

  • — Janák and Vysoky, 1992: 137 (Heterothops; collected with Formica rufa and truncorum; Bohemia).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Outerelo, Palmer and Pons, 1995: 80 (Heterothops; Mallorca).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; probably a species complex; Italy).

  • — Ryabukhin, 1999: 69 (Heterothops; catalog; habitat; distribution; northeastern Russia).

  • — Kashcheev, 1999a: 58 (Heterothops; Kazakhstan).

  • Distribution: Libya, Tunisia, Algeria, Morocco, Canary Islands, Maderia, Europe, Russia, Armenia, Azerbaijan, Turkey, Iran, Kazakhstan.

flavolimbatus Motschulsky, 1860: 571 (Heterothops; Type locality: Armenie; Daghestan).

  • — Hochhuth, 1862: 42 (Heterothops; characters; Armenia; Daghestan).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; synonym of dissimilis).

  • — Mulsant and Rey, 1876: 826 [= 1877: 682] (Heterothops; characters; France).

  • — Fowler, 1888: 225 (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

minutus Motschulsky, 1860: 572 (Heterothops; [preoccupied]; Type locality: fl. Irtysch dans la Sibérie occidentale).

  • — Hochhuth, 1862: 43 (Heterothops; notes; Siberia).

  • — Fauvel, 1874: 313 [= 1874a: 539] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

pygmaeus Gemminger and Harold, 1868: 567 (Heterothops; replacement name for minutus Motschulsky).

  • — Fauvel, 1874: 313 [= 1874a: 539] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

distinguendus Hochhuth, 1872: 130 (Heterothops; Type locality: Not cited, but presumably from Gouvernements Kiew and/or Volhynien).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

  • — Coiffait, 1978: 310 (Heterothops; synonym of praevius).

  • — Gusarov, 1992a: 786 [= 1993a: 76] (Heterothops; synonym of dissimilis).

parvicornis Mulsant and Rey, 1876: 826 [= 1877: 682 (Heterothops; Type locality: Lille).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Coiffait, 1978: 314 (Heterothops; variety of dissimilis; characters).

sericans Mulsant and Rey, 1876: 820 [= 1877: 676] (Heterothops; [preoccupied]; Type locality: Provence).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Europe).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog; western Mediterranean region).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

  • — Tóth, 1984: 102 (Heterothops; variety of dissimilis; characters).

paradoxus Mulsant and Rey, 1876: 823 [= 1877: 679] (Heterothops; Type locality: Not cited).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of sericans).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of sericans).

fraudulentus Coiffait, 1977c: 143 (Heterothops; variety of dissimilis; [Note: Nomen nudum (Article 15.2]; cited from Portugal, environs de Faro).

  • — Coiffait, 1978: 314 (Heterothops; variety of dissimilis; characters).

distinguendus Hochhuth, 1872, see: dissimilis Gravenhorst, 1802.

dolichocephalus Lea, 1925: 231 (Heterothops; Type locality: Tasmania; Waratah).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Australia.

eidmanni Scheerpeltz, 1936a: 502 (Heterothops; Type locality: Brasilien: Mendes, E. d. Rio).

  • Distribution: Brazil.

exilis Casey, 1886, see: occidentis Casey, 1886.

exilis Erichson, 1840: 518 (Heterothops; Type locality: Columbia).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Colombia.

fallax Lynch, 1884: 124 (Heterothops; Type locality: Chacabuco).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Argentina.

fauveli Bernhauer and Schubert, 1916: 412 (Heterothops; replacement name for flavicollis Fauvel).

  • — Bernhauer, 1920: 17 (Heterothops; Australia).

  • — Lea, 1925: 230 (Heterothops; synonym of xantholinoides).

  • — Scheerpeltz, 1933: 1424 (Heterothops; synonym of xantholinoides).

  • — Herman, 2001: 26 (Heterothops; synonymy).

  • Distribution: Australia.

xantholinoides W. J. MacLeay, 1873: 141 (Philonthus; [preoccupied by xantholinoides Wollaston]; Type locality: Gayndah).

  • — Fauvel, 1877: 275 (Heterothops; characters; Australia).

  • — Bernhauer and Schubert, 1916: 437 (Quedius; catalog).

  • — Bernhauer, 1920: 17 (Heterothops; Australia).

  • — Lea, 1925: 230 (Heterothops; characters; Australia).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Herman, 2001: 26 (Heterothops; synonym of fauveli).

flavicollis Fauvel, 1878g: 559 (Heterothops; [preoccupied]; Type locality: Australie méridionale: Nouvelle-Galles du Sud, Sydney).

  • — Lea, 1925: 230 (Heterothops; synonym of xantholinoides).

  • — Scheerpeltz, 1933: 1424 (Heterothops; synonym of xantholinoides).

flavicollis Fauvel, 1878, see: fauveli Bernhauer and Schubert, 1916.

flavicollis Motschulsky, 1858, see: Gabronthus.

flavicornis Bernhauer, 1917h: 85 (Heterothops; Type locality: Bolivien: Yuracares).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Bolivia.

flavolimbatus Motschulsky, 1860, see: dissimilis Gravenhorst, 1802.

flavus Scheerpeltz, 1972: 122 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, NW Tal, 460 m).

  • Distribution: Argentina.

formicetorum Bernhauer, 1912d: 46 (Heterothops; Type locality: Argentinien: Buenos Aires).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • Distribution: Argentina.

forsteri Scheerpeltz, 1960c: 128 (Heterothops; Type locality: Bolivien: Yungas de Arepucho, Sihuencas, 2200–2500 m).

  • Distribution: Bolivia.

franzi Coiffait, 1982: 82 (Heterothops; Type locality: Nepal: environs de Pina, region de Rara-See).

  • — Smetana, 1988: 328 (Heterothops; characters; Uttar Pradesh; Nepal).

  • Distribution: Nepal, India.

frater Last, 1975: 433 (Heterothops; Type locality: Okasa, 3000 ft., North Fore census division, Okapa subdistrict).

  • Distribution: New Guinea.

fraternus Smetana, 1971: 35 (Heterothops; Type locality: Washington: Paradise Park, alt. 6000 ft., Mt. Rainier. Also cited from Oregon and Washington).

  • — Smetana, 1971b: 1 (Heterothops; Washington).

  • Distribution: USA.

fraudulentus Coiffait, 1977 [Note: Nomen nudum], see: dissimilis Gravenhorst, 1802.

fulvipennis Bernhauer, 1921, see: laxus Blackwelder, 1944.

fulvipennis Motschulsky, 1860, see: praevius Erichson, 1839.

fulvus Scheerpeltz, 1972: 119 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1170 m).

  • Distribution: Argentina.

fumigatus LeConte, 1863, see: fusculus LeConte, 1863.

fundatus Last, 1980: 151 (Heterothops; Type locality: New Guinea: Mt. Wilhelm, 3900 m).

  • Distribution: New Guinea.

funeralis Last, 1975: 432 (Heterothops; Type locality: Karimui, 3500 ft, S.W. Goroka, 6°32′[S],144°47′[E]).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea; Irian).

  • Distribution: New Guinea.

fusculus LeConte, 1863a: 35 (Heterothops; Type locality: Cambridge, Massachusetts; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; variety of fumigatus; characters).

  • — Blatchley, 1910: 369 (Heterothops; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; synonym of fumigatus).

  • — Smetana, 1971: 26 (Heterothops; cited as valid species; lectotype designation; characters; Alberta; British Columbia; Manitoba; Nova Scotia; Ontario; Quebec; Saskatchewan; Arizona; California; Colorado; Idaho; Illinois; Iowa; Maine; Massachusetts; Michigan; Minnesota; Montana; Nevada; New Hampshire; New Jersey; New Mexico; New York; North Carolina; North Dakota; Oregon; Pennsylvania; Washington; Wisconsin).

  • — Smetana, 1971b: 1 (Heterothops; Michigan).

  • — Levesque and Levesque, 1996: 292 (Heterothops; seasonal abundance; Quebec).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • Distribution: Canada, USA.

californicus LeConte, 1863a: 35 (Heterothops; Type locality: San Francisco, California; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; variety of fumigatus; characters).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; synonym of fumigatus).

  • — Hatch, 1957: 231 (Heterothops; synonym of fumigatus).

  • — Smetana, 1971: 26 (Heterothops; lectotype designation; synonym of fusculus).

fumigatus LeConte, 1863a: 35 (Heterothops; Type locality: Detroit; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 153 (Heterothops; characters; Michigan; Texas; California; middle States; Canada).

  • — Fall and Cockerell, 1907: 166 (Heterothops; New Mexico).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Hatch, 1957: 231 (Heterothops; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1965: 34 (Heterothops; Newfoundland).

  • — Smetana, 1971: 26 (Heterothops; lectotype designation; synonym of fusculus).

gemellus Smetana, 1971: 19 (Heterothops; Type locality: California: Pasadena).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; California).

  • Distribution: USA.

gestroi Gridelli, 1922: 25 (Heterothops; Type locality: Persia settentrionalis).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Coiffait, 1978: 309 (Heterothops; characters; Iran; Lebanon; Uzbekistan).

  • Distribution: Lebanon, Iran, Uzbekistan.

gijiensis Last, 1987: 54 (Heterothops; Type locality: Irian: Vogelkop, Lake Anggi Giji, 2000–2100 m).

  • Distribution: New Guinea.

gnarus Last, 1975: 435 (Heterothops; Type locality: Daulo Pass, 8000 ft., Osaro Climbu Divide).

  • Distribution: New Guinea.

grandiceps Bernhauer, 1917h: 84 (Heterothops; Type locality: Brasilien: Blumenau).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Brazil.

gressitti Last, 1987: 53 (Heterothops; Type locality: Papua New Guinea: New Guinea NE: Asaro-Chimbu Div., Daulo Pass, 2800 m).

  • Distribution: New Guinea.

giulianii Smetana, 1978a: 819 (Heterothops; Type locality: California: Inyo Co.: Rock Creek, 10,000 ft.).

  • Distribution: USA.

hindustanus Cameron, 1932: 267 (Heterothops; Type locality: Chakrata district: Korawa Khud, alt. 9000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Coiffait, 1982: 33 (Heterothops; Nepal).

  • — Smetana, 1988: 325 (Heterothops; lectotype designation; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal, Myanmar.

holmensis Stephens, 1833, see: binotatus Gravenhorst, 1802.

hornabrooki Last, 1975: 430 (Heterothops; Type locality: N.E. New Guinea: Daulo Pass, North of Goroka, Eastern Highlands, 8200 ft.).

  • Distribution: New Guinea.

impressifrons Germain, 1903: 422 (Heterothops; Type locality: cordilleras de Chillan, cerca de las Termas).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • Distribution: Chile.

indicus Cameron, 1926b: 365 (Heterothops; Type locality: Mossy Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussoorie district; Arni Gad. Chakrata and Simla Hills).

  • — Scheerpeltz, 1933: 1423 (Heterothops; synonym of oculatus).

  • — Smetana, 1988: 322 (Heterothops; valid species; lectotype designation: Mossy Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

insolitus Sharp, 1884: 323 (Heterothops; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Guatemala.

iranicus Štourač, 2000: 68 (Heterothops; Type locality: SE Iran: 40 km SW Zábolí).

  • Distribution: Iran.

ishiharai Ito, 1994a: 177 (Heterothops; Type locality: Japan: Yamanashi Pref.: Gozaishi Spa, alt. 1400 m).

  • Distribution: Japan.

jureceki Štourač, 2000: 70 (Heterothops; Type locality: Tockoe, Ros. or., Gub Samara).

  • Distribution: Russia.

kentiae Lea, 1925: 238 (Heterothops; Type locality: Lord Howe Island, summit of Mount Gower).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

khairo Smetana, 1988: 326 (Heterothops; Type locality: India: Dehra Dun).

  • Distribution: India.

kirbiella Stephens, 1833, see: binotatus Gravenhorst, 1802.

laeticolor Reitter, 1891: 17 (Heterothops; Type locality: Kasalinsk).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Coiffait, 1978: 318 (Heterothops; characters; Transcaspian region; Transcaucasus).

  • Distribution: Kazakhstan, “Transcaucasus”.

laticeps Fauvel, 1878g: 558 (Heterothops; Type locality: Australie orientale: Queensland, Wide Bay).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; catalog).

  • — Lea, 1925: 230 (Heterothops; valid species; characters; Australia).

  • Distribution: Australia.

laxus Blackwelder, 1944: 144 (Heterothops; replacement name for fulvipennis Bernhauer).

  • Distribution: Argentina.

fulvipennis Bernhauer, 1921c: 172 (Heterothops; [preoccupied]; Type locality: Argentinien: Prov. Jujuy, bei Hornadita, 3400 und bei Cueva Iturbe, 3700 m).

  • — Scheerpeltz, 1933: 1422 (Heterothops; catalog).

  • — Blackwelder, 1944: 144 (Heterothops; synonym of laxus).

limbatus Heer, 1839, see: Quedius.

limosus Motschulsky, 1858, see: praevius Erichson, 1839.

longiceps Wendeler, 1956b: 262 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

luctuosus Fauvel, 1877: 275 (Heterothops; Type locality: Australie).

  • — Fauvel, 1878g: 558 (Heterothops; Australia).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

macrops Coiffait, 1977c: 142 (Heterothops; Type locality: Arménie, Dvine, 15 km au Sud de Erevan, Arménie soviétique).

  • — Coiffait, 1978: 313 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

magniceps Bernhauer, 1920, see: Ctenandropus.

malaisei Scheerpeltz, 1965: 280 (Heterothops; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

marmotae Smetana, 1971a: 1837 (Heterothops; Type locality: Ontario; Ottawa, Kanata).

  • — Smetana, 1973: 1421 (Heterothops; Ontario).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; Canada).

  • Distribution: Canada.

marocanus Coiffait, 1977d: 296 (Heterothops; Type locality: Maroc: Aguelmane Sidi Ali).

  • — Coiffait, 1978: 316 (Heterothops; characters; Morocco).

  • Distribution: Morocco.

mediocris Fall, 1907, see: occidentis Casey, 1886.

mediofuscus Lea, 1925: 233 (Heterothops; Type locality: Tasmania: Waratah).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

megalops Cameron, 1959: 117 (Heterothops; Type locality: Angola: Vila Luso).

  • Distribution: Angola.

melanocerus Solsky, 1871: 156 (Heterothops; Type locality: Samarkand).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Solsky, 1874: 171 (Heterothops; characters; Turkestan).

  • — Eppelsheim, 1892: 329 (Heterothops; valid species; Turkestan).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Coiffait, 1978: 314 (Heterothops; characters; Transcaucasus).

  • — Tronquet, 1981: 71 (Heterothops; Afghanistan).

  • Distribution: Uzbekistan, Afghanistan.

meurguesae Tronquet, 1981: 80 (Heterothops; Type locality: Afghanistan: Kenjan, 2000 m).

  • Distribution: Afghanistan.

mexicanus Schubert, 1909: 290 (Heterothops; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d'Orizaba).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Mexico.

microtophilus Coiffait, 1977c: 141 (Heterothops; Type locality: Goukassian Zouigakhpiour).

  • — Coiffait, 1978: 310 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

minor Smetana, 1971: 36 (Heterothops; Type locality: Ontario: Ottawa. Also cited from Alberta, British Columbia, Manitoba, Nova Scotia, Maine, New Hampshire, and Washington).

  • — Smetana, 1973: 1422 (Heterothops; Quebec).

  • — Smetana, 1976c: 169 (Heterothops; Alberta; Newfoundland; Northwest Territories).

  • — Smetana, 1981b: 633 (Heterothops; Newfoundland).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; Canada; USA).

  • Distribution: Canada, USA.

minutus Motschulsky, 1860, see: dissimilis Gravenhorst, 1802.

minutus Wollaston, 1860: 53 (Heterothops; Type locality: Palmeira above Funchal; [Note: See lectotype designation by Israelson, 1979a]).

  • — Fauvel, 1869: 486 (Heterothops; synonym of praevius).

  • — Fauvel, 1869a: 151 (Heterothops; synonym of praevius).

  • — Wollaston, 1871: 298l, 299 (Heterothops; notes; synonym of dissimilis).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; synonym of dissimilis).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of dissimilis).

  • — Porta, 1907: 103 (Heterothops; synonym of dissimilis).

  • — Bernhauer and Schubert, 1916: 412 (Heterothops; synonym of dissimilis).

  • — Uyttenboogaart, 1937: 81 (Heterothops; characters; notes; Canary Islands).

  • — Smetana, 1958: 318 (Heterothops; synonym of dissimilis).

  • — Smetana, 1963b: 39 (Heterothops; Madeira).

  • — Coiffait, 1978: 314 (Heterothops; synonym of dissimilis).

  • — Israelson, 1979a: 264, 266 (Heterothops; lectotype designation; valid species; Swedish specimens identified as brunnipennis identical to minutus; characters; species may be introduced into Madeira; specimens from Canary and Cape Verde Islands identified as minutus are different species; Madeira; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters; all central Europe).

  • — Borges, 1990: Table II (Heterothops; cited as valid species; Azores).

  • — Hodge and Jones, 1995: 41 (Heterothops; characters; Britain).

  • — Štourač, 1998b: 135 (Heterothops; notes; Czech Republic).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 53 (Heterothops; catalog; Poland).

  • Distribution: Azores, Madeira, Britain, Sweden, central Europe.

mirabilis Last, 1975: 426 (Heterothops; Type locality: New Guinea: Okapa).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea).

  • Distribution: New Guinea.

mirus Lea, 1925: 234 (Heterothops; Type locality: Lord Howe Island).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Lord Howe Island.

montanus Iablokov-Khnzorian, 1966: 174 (Heterothops; Type locality: ArmSSR: Argats, u ozera Karilich).

  • — Coiffait, 1978: 311 (Heterothops; characters; Armenia).

  • Distribution: Armenia.

montanus Last, 1975, see: tumulus Herman, 2001.

myrmeciae Lea, 1929: 205 (Heterothops; Type locality: Victoria: Northern Gippsland; Fern Tree Gully).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

nidicola Thomson, 1870, see: niger Kraatz, 1868.

nidorum Smetana, 1967d: 211 (Heterothops; Type locality: Mongolei: Cojbalsan aimak, 80 km NV von Cojbalsan, 700 m).

  • — Smetana 1975c: 175 (Heterothops; Mongolia).

  • Distribution: Mongolia.

niger Kraatz, 1868: 352 (Heterothops; Type locality: Thüringen).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; cited as nigra; synonym of praevius).

  • — Redtenbacher, 1874: 197 (Heterothops; characters).

  • — Mulsant and Rey, 1876: 818 [= 1877: 674] (Heterothops; characters; France).

  • — Fowler, 1888: 225 (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Joy, 1906a: 199 (Heterothops; notes; mole's nest; Britain).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Montandon, 1908: 80 (Heterothops; Romania).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Scheerpeltz, 1933: 1423 (Heterothops; variety of praevius).

  • — Paulian, 1941: 256 (Heterothops; larval characters).

  • — Smetana, 1958: 317 (Heterothops; characters; ecology; Czechoslovakia).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Smetana, 1964d: 78 (Heterothops; Czechoslovakia).

  • — Focarile, 1964: 57 (Heterothops; Italy).

  • — Horion, 1965: 247 (Heterothops; Denmark; Sweden; Britain; Holland; Belgium; France; Switzerland; Italy; Czechoslovakia; Poland; Germany; Austria).

  • — Pototskaia, 1967: 79 (Heterothops; larval characters).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Bolov, 1969a: 329 (Heterothops; collecting notes; Russia).

  • — Osella and Zanetti, 1975: 112 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 312 (Heterothops; characters; Europe; Britain; Scandinavia).

  • — Israelson, 1979a: 263, 265 (Heterothops; subspecies of praevius; discussion of status; characters; collecting notes).

  • — Tóth, 1984: 100 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988a: 80 (Heterothops; characters).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Nowosad, 1990: 120 (Heterothops; collected from mole nest; notes; Poland).

  • — Janák and Vysoky, 1992: 137 (Heterothops; collected with Formica pratensis; Bohemia).

  • — Welch, 1993: 229 (Heterothops; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • Distribution: Britain, Sweden, Denmark, Holland; Belgium, France, Germany, Poland, Czechoslovakia, Switzerland, Italy, Austria, Hungary, Romania, Russia.

nidicola Thomson, 1870: 127 (Heterothops; Type locality: Bastad in nordvestra Skåne).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Fowler, 1888: 225 (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 317 (Heterothops; synonym of niger).

  • — Coiffait, 1978: 312 (Heterothops; synonym of niger).

nigerrimus Bernhauer, 1914b: 67 (Heterothops; Type locality: Ghissar-Gebirge in Turkestan).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1963: 14 (Heterothops; Afghanistan).

  • — Coiffait, 1978: 316 (Heterothops; characters; Transcaspian region).

  • Distribution: Tajikistan, Afghanistan.

nigritulus R. Sahlberg, 1847: 790 (Quedius; Type locality: Rio Janeiro).

  • — Fauvel, 1891: 120 (Heterothops).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Brazil.

nigrofrater Lea, 1925: 236 (Heterothops; Type locality: Victoria: Warburton, Belgrave, Killara; Melbourne. Tasmania: Mount Wellington).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

nitens Nordmann, 1837: 138 (Trichopygus; Type locality: Taurus-Chersonesus).

  • — Erichson, 1840: 518 (Heterothops; characters; Chersonesus Tauricus).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Germany).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Germany).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Russia.

obscuripennis Lea, 1925: 237 (Heterothops; Type locality: South Australia: Kangaroo Island).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

occidentis Casey, 1886: 263 (Heterothops; replacement name for exilis Casey).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Smetana, 1971: 17 (Heterothops; characters; California; New Mexico).

  • — Legner and Moore, 1977: 175 (Heterothops; cited as occidentalis; found under bark).

  • Distribution: USA.

exilis Casey, 1886: 208 (Heterothops; [preoccupied]; Type locality: California: Monterey Co.).

  • — Scheerpeltz, 1933: 1423 (Heterothops; synonym of occidentis).

  • — Smetana, 1971: 17 (Heterothops; synonym of occidentis).

mediocris Fall, 1907: 220 (Heterothops; Type locality: Las Vegas Hot Springs, New Mexico; Riverside, California).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Smetana, 1971: 17 (Heterothops; synonym of occidentis).

ocularis Blackwelder, 1943: 465 (Heterothops; Type locality: Trinidad: mouth of cave in Aripo Valley, elevation 2600 feet).

  • Distribution: West Indies.

oculatus Fauvel, 1895: 275 (Heterothops; Type locality: Tenasserim, mont Mooleyit, 1800–1900 m; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Cameron, 1932: 267 (Heterothops; characters; Burma; India).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Coiffait, 1982a: 234 (Heterothops; Pakistan).

  • — Smetana, 1988: 320 (Heterothops; lectotype designation; characters; Burma; Nepal).

  • — Smetana, 1992b: 10 (Heterothops; Nepal).

  • — Smetana, 1995: 122 (Heterothops; characters; bionomics; Taiwan).

  • Distribution: Pakistan, India, Nepal, Myanmar, Taiwan.

okapaensis Last, 1975: 431 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

orbicularis Sharp, 1884: 324 (Heterothops; Type locality: Guatemala, San Juan in Vera Paz; Panama, Volcan de Chiriqui 3000 to 4000 feet, Boquete 3500 feet).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • Distribution: Guatemala, Panama.

papuanus Cameron, 1937a: 104 (Quedius; subgenus Sauridus; Type locality: New Guinea: Mt. Tafa, alt. 8500 ft.).

  • — Last, 1975: 425, 438 (Heterothops; New Guinea).

  • Distribution: New Guinea.

paradoxus Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

paralleliceps Bernhauer, 1917h: 87 (Heterothops; Type locality: West-Kolumbien: Quindici Pass bei Linia, 3600 m).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1960c: 128 (Heterothops; Bolivia).

  • Distribution: Colombia, Bolivia.

parvicornis Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

parvipennis Scheerpeltz, 1972: 116 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1000 m).

  • Distribution: Argentina.

persimilis Cameron, 1932: 267 (Heterothops; Type locality: Simla Hills: Kotgarh, alt. 7000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1965: 111 (Heterothops; Burma).

  • — Smetana, 1988: 326 (Heterothops; lectotype designation; characters; Himachal Pradesh; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal, Myanmar.

picipennis Fauvel, 1877: 276 (Heterothops; Type locality: Australie, Victoria).

  • — Fauvel, 1878g: 558 (Heterothops; notes; Australia).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Lea, 1925: 231 (Heterothops; from nest of Myrmecia sp.; Australia).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Scheerpeltz, 1935a: 655 (Heterothops; Australia).

  • Distribution: Australia.

picticollis Lea, 1925: 232 (Heterothops; Type locality: Victoria: Dividing Range; Ferntree Gully; Killara).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

pictus Lea, 1925: 239 (Heterothops; Type locality: Victoria: Victorian Alps, Dividing Range, Belgrave, Ferntree Gully, Warburton, Bright, Wandin, Emerald. Tasmania, Zeehan, Western Australia: Donnybrook).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • Distribution: Australia.

praevius Erichson, 1839a: 480 (Heterothops; Type locality: Mark Brandenburg).

  • — Lacordaire, 1835: 385 (Emus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited subuliformis Boisduval and Lacordaire as a synonym of praevius Erichson; Lacordaire did not describe the species as new, he attributed it to Gravenhorst]; characters; France).

  • — Nordmann, 1837: 137 (Trichopygus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited dissimilis Nordmann as a synonym of praevius Erichson; Nordmann did not describe the species as new, he attributed it to Gravenhorst]; characters; Germany; Russia).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 516 (Heterothops; characters; Germany).

  • — Märkel, 1841: 217 (Heterothops; in ant nest).

  • — Redtenbacher, 1849: 711 (Heterothops; characters; Austria).

  • — Rosenhauer, 1856: 71 (Philonthus; Granada).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Kraatz, 1857d: 484 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; characters; France; Europe; Caspian Sea; Iran; Buchara).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 814 [= 1877: 670] (Heterothops; characters; larval characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ragusa, 1892: 167 (Heterothops; Sicily).

  • — Ganglbauer, 1895: 387 (Heterothops; characters; Europe; eastern Mediterranean region).

  • — Fauvel, 1902b: 118 (Heterothops; Algeria).

  • — Porta, 1907: 102 (Heterothops; characters; Italy).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 346 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Šulc, 1922 (Heterothops; osmeterium; anatomy; histology; physiology).

  • — Portevin, 1929: 336 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1423 (Heterothops; catalog).

  • — Bernhauer, 1936: 5 (Heterothops; Mongolia).

  • — Paulian, 1941: 256 (Heterothops; larval characters).

  • — Hinton, 1945: 69 (Heterothops; characters).

  • — Jeannel and Jarrige, 1949: 377 (Heterothops; cave species; Spain; Algeria).

  • — V. Hansen, 1952: 159 (Heterothops; characters; Denmark).

  • — Scheerpeltz, 1958a: 23 (Heterothops; Turkey).

  • — Smetana, 1958: 316 (Heterothops; characters; ecology; Czechoslovakia).

  • — Scheerpeltz, 1963: 14 (Heterothops; Afghanistan).

  • — Palm, 1963b: 156 (Heterothops; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Scheerpeltz, 1965c: 510 (Heterothops; Sicily).

  • — Horion, 1965: 246 (Heterothops; Caucasus; Transbaikal; Afghanistan; Russia; Finland; Sweden; Norway; Britain; Denmark; France; Belgium; Holland; Germany; Austria; Czechoslovakia; Poland; Romania; Portugal; Balaearics; Italy).

  • — Pototskaia, 1967: 79 (Heterothops; larval characters).

  • — Smetana, 1967g: 470 (Heterothops; characters).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Osella and Zanetti, 1975: 110 (Heterothops; collected from nest of Talpa europaea; notes; Italy).

  • — Pope, 1977: 30 (Heterothops; Britain).

  • — Coiffait, 1978: 310 (Heterothops; characters; Europe; Britain; Scandinavia; North Africa).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • — Tóth, 1984: 101 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1988a: 80 (Heterothops; characters).

  • — Lohse and Lucht, 1989: 169 (Heterothops; characters).

  • — Nowosad, 1990: 117 (Heterothops; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy).

  • — Ryabukhin, 1999: 70 (Heterothops; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Algeria, Europe, Russia, Iran, Afghanistan, Uzbekistan, Mongolia.

limosus Motschulsky, 1858: 660 (Heterothops; cited as Heteropos; Type locality: Styrie).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

fulvipennis Motschulsky, 1860: 572 (Heterothops; Type locality: environs de St. Pétersbourg sur les bords de la Néva).

  • — Fauvel, 1874: 311 [= 1874a: 537] (Heterothops; synonym of praevius).

  • — Ganglbauer, 1895: 387 (Heterothops; synonym of praevius).

  • — Porta, 1907: 102 (Heterothops; synonym of praevius).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of praevius).

  • — Smetana, 1958: 316 (Heterothops; synonym of praevius).

  • — Coiffait, 1978: 310 (Heterothops; synonym of praevius).

primarius Last, 1975: 426 (Heterothops; Type locality: N.E. New Guinea: Mount Michael, 8–1000 ft.).

  • Distribution: New Guinea.

pumilio Nordmann, 1837, see: quadripunctulus Gravenhorst, 1806.

pusillus Coiffait, 1982: 83 (Heterothops; Type locality: Nepal: environs de Pina, region de Rara-See).

  • — Smetana, 1988: 323 (Heterothops; characters; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1992b: 10 (Heterothops; Nepal).

  • Distribution: Nepal, India.

pusio LeConte, 1863a: 35 (Heterothops; Type locality: New York).

  • — Horn, 1878: 153 (Heterothops; characters; New York; Pennsylvania; Ohio; Arizona).

  • — Blatchley, 1910: 369 (Heterothops; characters; Indiana).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Smetana, 1971: 20 (Heterothops; characters; British Columbia; Ontario; Quebec; Arizona; Arkansas; California; Illinois; Indiana; Iowa; Kentucky; Maine; Massachusetts; Michigan; Missouri; New Jersey; Ohio; Oregon; Pennsylvania; Wisconsin).

  • — Smetana, 1973: 1421 (Heterothops; Quebec).

  • — Legner and Moore, 1977: 176 (Heterothops; found under bark).

  • — Smetana, 1978a: 817 (Heterothops; collecting notes; Arizona; Arkansas; California; Oklahoma; Texas).

  • — Smetana, 1981b: 631 (Heterothops; Ontario).

  • — Smetana, 1990d: 96 (Heterothops; Saskatchewan).

  • — Downie and Arnett, 1996: 388 (Heterothops; characters; USA).

  • — Schmidt, 1999: 108 (Heterothops; food for adults and larvae used for rearing species; rearing methods).

  • Distribution: Canada, USA.

pygmaeus Gemminger and Harold, 1868, see: dissimilis Gravenhorst, 1802.

quadriceps Bernhauer, 1917h: 87 (Heterothops; Type locality: Argentinien: Prov. Tucuman).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Argentina.

quadripunctulus Gravenhorst, 1806: 24 (Tachinus; Type locality: Brunsvigae).

  • — Zetterstedt, 1828: 72 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 413 cited subuliformis Zetterstedt as a synonym of quadripunctulus; Zetterstedt did not describe the species as new, he attributed it to Gravenhorst]; characters; Lapland).

  • — Erichson, 1839a: 481 (Heterothops; characters; Germany).

  • — Heer, 1839: 281 (Heterothops; characters; Switzerland).

  • — Erichson, 1840: 517 (Heterothops; characters; Germany).

  • — Jacobsen, 1849: 707 (Heterothops; characters; Denmark).

  • — Redtenbacher, 1849: 712 (Heterothops; cited as quadripunctatus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 533 (Heterothops; characters; France).

  • — Kraatz, 1857d: 486 (Heterothops; characters; Germany).

  • — Redtenbacher, 1857: 199 (Heterothops; characters; Austria).

  • — Thomson, 1860: 181 (Heterothops; characters; Scandinavia).

  • — Gredler, 1863: 99 (Heterothops; Austria).

  • — Fauvel, 1874: 312 [= 1874a: 538] (Heterothops; characters; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Tirol; Russia).

  • — Redtenbacher, 1874: 196 (Heterothops; characters; Austria).

  • — Seidlitz, 1875: 263 (Heterothops; characters; Baltic region).

  • — Mulsant and Rey, 1876: 829 [= 1877: 685] (Heterothops; characters; France).

  • — J. Sahlberg, 1876: 30 (Heterothops; Finland).

  • — Fowler, 1888: 225 (Heterothops; characters; Britain).

  • — Ganglbauer, 1895: 388 (Heterothops; characters; north and middle Europe).

  • — Porta, 1907: 104 (Heterothops; characters; Italy).

  • — Reitter, 1909: 108 (Heterothops; characters; Germany).

  • — Johansen, 1914: 347 (Heterothops; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Portevin, 1929: 337 (Heterothops; characters; France).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — V. Hansen, 1952: 160 (Heterothops; characters; Denmark).

  • — Smetana, 1958: 319 (Heterothops; characters; ecology; Czechoslovakia).

  • — Palm, 1963b: 157 (Heterothops; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 204 (Heterothops; characters; central Europe).

  • — Smetana, 1964d: 78 (Heterothops; Czechoslovakia).

  • — Horion, 1965: 249 (Heterothops; Russia; Finland; Sweden; Norway; Denmark; Britain; Poland; Czechoslovakia; Hungary; Hercegovina; Italy; Switzerland; Austria; Germany; France; Holland).

  • — Smetana, 1967g: 471 (Heterothops; characters).

  • — Szujecki, 1968a: 733 (Heterothops; Poland).

  • — Coiffait, 1978: 316 (Heterothops; characters; Britain; France).

  • — Israelson, 1979a: 265, 266 (Heterothops; characters).

  • — Tóth, 1984: 101 (Heterothops; characters; Hungary).

  • — Lucht, 1987: 108 (Heterothops; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 171 (Heterothops; characters).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Heterothops; Italy ?).

  • Distribution: Europe, Russia.

pumilio Nordmann, 1837: 138 (Trichopygus; Type locality: Berolini).

  • — Erichson, 1839a: 481 (Heterothops; synonym of quadripunctulus).

  • — Erichson, 1840: 517 (Heterothops; synonym of quadripunctulus).

  • — Redtenbacher, 1849: 712 (Heterothops; synonym of quadripunctulus).

  • — Kraatz, 1857d: 486 (Heterothops; synonym of quadripunctulus).

  • — Ganglbauer, 1895: 388 (Heterothops; synonym of quadripunctula).

  • — Porta, 1907: 104 (Heterothops; synonym of quadripunctula).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; synonym of quadripunctula).

  • — Smetana, 1958: 319 (Heterothops; synonym of quadripunctulus).

  • — Coiffait, 1978: 316 (Heterothops; synonym of quadripunctula).

rambouseki Blackwelder, 1943: 465 (Heterothops; Type locality: Cuba: district of Jamaica, province of Habana. Also cited from Jamaica).

  • Distribution: West Indies.

remotus Sharp, 1884: 323 (Heterothops; Type locality: Mexico, Mirador; Guatemala, Cerro Zunil 4000 to 5000 feet, San Geronimo; Panama, Volcan de Chiriqui 2500 to 4000 feet, Pena Blanca 3000 to 4000 feet).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Panama, Guatemala, Mexico.

riparius Peyerimhoff, 1929: 56 (Heterothops; Type locality: Hoggar de 1,700 a 2,400 m: Adrar Amezzeroui, Aguelmane Imerrera, Tihentekert, Arrem de Tinozel, Tezeit. Tifedest, vers 1150 m: Oued Aguelil, oued Ahetes. Haut Igharghar a Timenain. Tassili occidental, 600–800 m: Tim Tahart et Tin Eselmaken, Amguid).

  • — Peyerimhoff, 1931: 32 (Heterothops; characters; Algeria).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Jarrige, 1958: 92 (Heterothops; Algeria).

  • — Coiffait, 1966b: 338 (Heterothops; Mauritania).

  • — Coiffait, 1978: 311 (Heterothops; characters; Sahara).

  • Distribution: Algeria, Mauritania.

rotundiceps Sharp, 1889: 35 (Heterothops; Type locality: Nikko; Kiga).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Watanabe and Shibata, 1972: 65 (Heterothops; Yaku-shima Island).

  • — Cho, 1996b: 109 (Heterothops; characters; Korea).

  • Distribution: Japan, Korea.

rubicundulus Last, 1980: 152 (Heterothops; Type locality: New Guinea: Mt. Wilhelm, Keglsugl).

  • Distribution: New Guinea.

rufosuturalis Lea, 1925: 234 (Heterothops; Type locality: Queensland: Goodna; Brisbane; New South Wales: Tweed River).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

rufulus Last, 1980: 152 (Heterothops; Type locality: New Guinea: Mount Wilhelm, Field Station).

  • Distribution: New Guinea.

rufus Last, 1987: 53 (Heterothops; Type locality: Irian: Vogelkop, Fak Fak, S coast of Bomberai, 100–700 m).

  • Distribution: New Guinea.

saano Smetana, 1988: 329 (Heterothops; Type locality: India: Kashmir, Aru).

  • Distribution: India.

saltus Last, 1975: 427 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

saphaa Smetana, 1988: 330 (Heterothops; Type locality: Nepal: 148 Manang Dist., Marayandi, 3800–4100 m, oberhalb Manang Richtung Thorung).

  • Distribution: Nepal.

semicupreus Fauvel, 1878g: 557 (Heterothops; Type locality: Australie orientale: Queensland, Wide Bay).

  • — Bernhauer and Schubert, 1916: 413 (Heterothops; catalog).

  • — Lea, 1929: 205 (Heterothops; characters).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

sericans Stephens, 1832, see: binotatus Gravenhorst, 1802.

sericans Mulsant and Rey, 1876, see: dissimilis Gravenhorst, 1802.

simulator Last, 1980: 153 (Heterothops; Type locality: New Guinea: Mount Wilhelm).

  • Distribution: New Guinea.

simulatrix Smetana, 1971: 37 (Heterothops; Type locality: California: Mokell. Hill, Calaveras Co.).

  • Distribution: USA.

sordidus Smetana, 1971: 32 (Heterothops; Type locality: British Columbia: 8 mi. W. Creston).

  • — Smetana, 1973: 1422 (Heterothops; Northwest Territories).

  • — Smetana, 1981b: 632 (Heterothops; collecting notes; Alaska).

  • Distribution: Canada, USA.

stiglundbergi Israelson, 1979: 259 (Heterothops; Type locality: Sweden: Lomma).

  • — Lohse and Lucht, 1989: 170 (Heterothops; characters).

  • — Silfverberg, 1992: 17 (Heterothops; Finland; Russia; Sweden; Denmark; Lithuania).

  • — Smetana, 1993b: 49 (Heterothops; Czech Republic).

  • — Assing, Frisch, Kahlen, et al., 1998: 139 (Heterothops; all central Europe).

  • — Ødegaard, 1999: 72 (Heterothops; invasive species associated with compost heaps in Sweden, Demark, Finland, and Norway).

  • — Burakowski, Mroczkowski, and Stefańska, 2000: 53 (Heterothops; catalog; Poland).

  • Distribution: Europe.

subterraneus Bernhauer, 1921c: 173 (Heterothops; Type locality: Argentinien: Prov. Buenos Aires, Monte Veloz).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Argentina.

tantillus Lea, 1925: 235 (Heterothops; Type locality: Queensland: Cairns district).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

tantulus Last, 1975: 429 (Heterothops; Type locality: Okapa, 5800 ft.).

  • Distribution: New Guinea.

tanygnathoides Reitter, 1891: 17 (Heterothops; Type locality: Kasalinsk, am Aralsee).

  • — Eppelsheim, 1892: 329 (Heterothops; Turkestan).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Coiffait, 1978: 317 (Heterothops; cited as tanygnatoides; characters; Transcaspian region).

  • Distribution: Kazakhstan, Uzbekistan.

taurus Blackburn, 1888: 4 (Heterothops; Type locality: about 30 miles north of Port Lincoln).

  • — Blackburn, 1891: 69 (Quedius; characters; notes).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Lea, 1925: 240 (Quedius; synonym of hybridus).

  • Distribution: Australia.

tenuicornis Sharp, 1884: 324 (Heterothops; Type locality: Guatemala, near the city).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • Distribution: Guatemala.

thaxteri Bernhauer, 1917h: 86 (Heterothops; Type locality: Argentinien).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • — Frank, 1982: 37 (Heterothops; list of parasites).

  • Distribution: Argentina.

tibialis Fauvel, 1878g: 559 (Heterothops; Type locality: Australie méridionale: Victoria).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Lea, 1925: 231 (Heterothops; characters; Australia).

  • Distribution: Australia.

titschacki Bernhauer, 1941c: 289 [= 1951: 279] (Heterothops; Type locality: Süd Peru: Palco, 2600 m).

  • Distribution: Peru.

topali Scheerpeltz, 1972: 112 (Heterothops; Type locality: Argentinien: Rio Negro, El Bolson, Mt. Piltriquitron, 1500 m).

  • Distribution: Argentina.

tovarensis Fauvel, 1891: 120 (Heterothops; Type locality: Colonie Tovar).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • Distribution: Venezuela.

tumulus Herman, 2001: 45 (Heterothops; replacement name for montanus Last).

  • Distribution: New Guinea.

montanus Last, 1975: 434 (Heterothops; [preoccupied] Type locality: N.E. New Guinea: Mount Hagen, 9000 ft., Western Highlands District).

  • — Last, 1987: 55 (Heterothops; Papua New Guinea).

  • — Herman, 2001: 45 (Heterothops; synonym of tumulus).

tzaw Smetana, 1995: 124 (Heterothops; Type locality: Taiwan: near Tungpu, Nantou-Hsien, 2600 m).

  • Distribution: Taiwan.

ubiquitosus Lea, 1925: 236 (Heterothops; Type locality: Tasmania: Mount Wellington, Hobart, Huon River, Latrobe, Stanley, Ulverstone, Launceston, Frankford, Burnie. Victoria: Melbourne, Beaconfield, Emerald, Belgrave, Ferntree Gully. South Australia: Myponga, Lucindale).

  • — Scheerpeltz, 1933: 1424 (Heterothops; catalog).

  • Distribution: Australia.

velatus Wendeler, 1956b: 262 (Heterothops; Type locality: Brasilien: Nova Teutonia).

  • Distribution: Brazil.

vicunianus Germain, 1903, see: Cheilocolpus.

wouei Drugmand, 1988a: 74 (Heterothops; Type locality: Vierves-sur-Viroin, Province de Namur, Belgique; also included larval characters).

  • Distribution: Belgium.

willbergi Reitter, 1891: 18 (Heterothops; Type locality: Kasalinsk).

  • — Bernhauer and Schubert, 1916: 414 (Heterothops; catalog).

  • — Coiffait, 1978: 318 (Heterothops; cited as wilbergi; characters; Transcaucasus).

  • Distribution: Kazakhstan.

wittei Bernhauer, 1927, see: Rolla.

wollastoni Israelson, 1979a: 266 (Heterothops; Type locality: S. Antao, Cape Verde Islands).

  • — Wollaston, 1867: 235 (Heterothops; [Note: Misidentified as minutus according to Israelson, 1979a: 266] characters; notes; Cape Verde Islands).

  • Distribution: Cape Verde Islands.

xantholinoides W. J. MacLeay, 1873, see: fauveli Bernhauer and Schubert, 1916.

Indoquedius [14 species; Palaearctic and Oriental Regions]

Indoquedius Blackwelder, 1952: 199 (subgenus of Quedius; name made available by Blackwelder's designation of type species [Articles 13.3, 50.1]; species included: see Cameron, 1932: 281, 300). Type species: Quedius oculatus Fauvel, fixed by original designation by Blackwelder, 1952: 199, and selected from among species included by Cameron, 1932 (see Herman, 2001).

  • — Cameron, 1932: 281, 300 (subgenus of Quedius; species included: oculatus, filicornis, bipunctatus; characters; [Note: Cameron (1932) did not designate a type species for Indoquedius so the name was unavailable (Article 13.3). Blackwelder (1952) made the name available when he designated the type species]).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Korge, 1963a: 85 (subgenus of Quedius; characters; notes).

  • — Shibata, 1973c: 63 (subgenus of Quedius; checklist of species of Taiwan).

  • — Shibata, 1984: 137 (subgenus of Quedius; checklist of species of Japan).

  • — Smetana, 1988: 300 (valid genus; characters; key to species of Himalayan region).

  • — Smetana, 1995: 112 (characters; key to and revision of species of Taiwan).

  • — Herman, 2001: 9 (type species; author of name is Blackwelder, not Cameron).

aberrans Cameron, 1932: 299 (Quedius; subgenus Raphirus; Type locality: Sikkim: Rungbong Valley).

  • — Smetana, 1988: 311 (Indoquedius; characters; India).

  • Distribution: India.

akalita Smetana, 1977, see: sikkimensis Cameron, 1932.

baliyo Smetana, 1988: 304 (Indoquedius; Type locality: Nepal: Prov. Bagmati: Malemchi, 2800 m. Also cited from India).

  • — Smetana, 1992b: 10 (Indoquedius; Nepal).

  • Distribution: Nepal, India.

bing Smetana, 1995: 113 (Indoquedius; Type locality: Taiwan: Nantou Hsien: Hwy. 14, Fengnan, 700 m).

  • Distribution: Taiwan.

bipunctatus Eppelsheim, 1895b: 390 (Quedius; Type locality: Parasnath-Gebirge; [Note: See lectotype designation by Smetana, 1988]).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 301 (Quedius; subgenus Indoquedius; characters; Bihar).

  • — Scheerpeltz, 1933: 1442 (Quedius; synonym of grandiceps).

  • — Smetana, 1988: 305 (Indoquedius; cited as valid species; lectotype designation; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

conicus Champion, 1922, see: filicornis Eppelsheim, 1895.

cyanipennis Coiffait, 1982, see: sikkimensis Cameron, 1932.

daai Smetana, 1988: 307 (Indoquedius; Type locality: Inde: Kumaon, UP, Bhim Tal env. 1500 m. Also from Himachal Pradesh).

  • Distribution: India.

filicornis Eppelsheim, 1895c: 55 (Quedius; Type locality: Sikkim im Himalaya).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Cameron, 1932: 301 (Quedius; subgenus Indoquedius; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1988: 309 (Indoquedius; characters; Sikkim; Uttar Pradesh; Nepal).

  • — Smetana, 1992b: 10 (Indoquedius; Nepal).

  • Distribution: India, Nepal.

conicus Champion, 1922: 33 (Quedius; subgenus Raphirus; Type locality: Ranikhet Division of Kumaon).

  • — Cameron, 1932: 301 (Quedius; synonym of filicornis).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of filicornis).

  • — Smetana, 1988: 309 (Indoquedius; synonym of filicornis).

formosae Cameron, 1949: 176 (Quedius; subgenus Indoquedius; Type locality: Formosa: Tamari, Taito).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1995: 114 (Indoquedius; characters; Taiwan).

  • Distribution: Taiwan.

grandiceps Kraatz, 1879: 122 (Quedius; Type locality: Amur).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1442 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Herman, 2001: 83 (Indoquedius).

  • Distribution: Russia.

juno Sharp, 1874: 24 (Quedius; Type locality: Yamato).

  • — Sharp, 1889: 29 (Quedius; Japan).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Smetana, 1988: 300 (Indoquedius).

  • Distribution: Japan.

oculatus Fauvel, 1895: 274 (Quedius; Type locality: Bírmanie, Carin Asciuii Ghécu, 1400–1500 m; [Note: See lectotype designation by Smetana, 1988]. Also cited from Bírmanie: Carin Cheba, 900–1100 m).

  • — Bernhauer and Schubert, 1916: 430 (Quedius; catalog).

  • — Cameron, 1932: 300 (Quedius; subgenus Indoquedius; characters; Burma; Tonkin).

  • — Scheerpeltz, 1933: 1454 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Smetana, 1988: 306 (Indoquedius; lectotype designation: Carin Asciuii Ghécu, 1400–1500 m; characters; Burma).

  • Distribution: Myanmar, Vietnam.

praeditus Sharp, 1889: 29 (Quedius; Type locality: Miyanoshita; Junsai).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • — Yuh, Paik, Kwon, and Lee, 1985: 249 (Quedius; subgenus Indoquedius; Korea).

  • — Smetana, 1988: 300 (Indoquedius).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Cho, 1996b: 113 (Quedius; subgenus Indoquedius; characters; Korea).

  • — Watanabe, 1996a: 15 (Indoquedius; cave collection; Japan).

  • Distribution: Japan, Korea, China.

saathi Smetana, 1988: 310 (Indoquedius; Type locality: India: Meghalaya, Khasi Hills, Shillong, 1850–1950 m).

  • Distribution: India.

shibatai Smetana, 1995: 115 (Indoquedius; Type locality: Taiwan: near Fenchifu, Chiai).

  • Distribution: Taiwan.

sikkimensis Cameron, 1932: 294 (Quedius; subgenus Raphirus; Type locality: Sikkim: Lachung, alt. 9000–10,000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1462 (Quedius; catalog).

  • — Smetana, 1988: 302 (Indoquedius; lectotype designation; characters; Bhutan; Arunachal Pradesh).

  • Distribution: India, Bhutan.

akalita Smetana, 1977a: 246 (Quedius; subgenus Indoquedius; Type locality: Bhutan: Thimphu).

  • — Smetana, 1988: 302 (Indoquedius; synonym of sikkimensis).

cyanipennis Coiffait, 1982a: 273 (Algon; Type locality: Bhoutan: Batbalithan, Bhumtang, 2600 m).

  • — Smetana, 1988: 302 (Indoquedius; synonym of sikkimensis).

Ioma [1 species; Ethiopian Region]

Ioma Blackwelder, 1952: 199 (replacement name for Tachinopsis Fauvel). Type species: Tachinopsis setigera Fauvel, fixed by objective synonymy with Tachinopsis.

Tachinopsis Fauvel, 1899: 22 (species included: setigera; preoccupied by Coquillett, 1897). Type species: Tachinopsis setigera Fauvel, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 440 (world catalog; 1 species).

  • — Blackwelder, 1952: 369 (synonym of Ioma; type species: setigera).

setigera Fauvel, 1899: 22 (Tachinopsis; Type locality: Abyssinie: Bogos, Sciotel; Haut-Sénégal: Kayes).

  • — Bernhauer and Schubert, 1916: 440 (Tachinopsis; synonym of setigera).

  • — Coiffait, 1968a: 137 (Tachinopsis; Chad).

  • — Scheerpeltz, 1974c: 25 (Tachinopsis; Sudan).

  • Distribution: Ethiopia, Sudan, Chad, Mali.

Loncovilius [7 species; Neotropical Region]

Loncovilius Germain, 1903: 439 (species included; semiflavus, lividipennis). Type species: Quedius semiflavus Fairmaire and Germain, fixed by subsequent designation by Blackwelder, 1952: 226.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 226 (type species).

  • — Coiffait and Saiz, 1966: 401 (characters; key to Chilean species).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Saiz, 1971: 387, 389 (notes on biogeography; notes on phenetic relationships).

  • — Herman, 2001: 24 (synonymy).

Lienturius Coiffait and Saiz, 1966: 402 (subgenus of Loncovilius; species included: germaini, discoideus, leiocephalus, heeri). Type species: Staphylinus leiocephalus Solier, fixed by original designation.

  • — Saiz, 1971: 382 (subgenus of Loncovilius; key to species of Chile).

Apoquedius Scheerpeltz, 1972: 24, 25 (subgenus of Quedius; species included: aeneipennis, piciformis). Type species: Quedius aeneipennis Fairmaire and Germain, fixed by original designation.

  • — Herman, 2001: 24 (synonym of Loncovilius).

aeneipennis Fairmaire and Germain, 1861: 428 (Quedius; [Note: Preoccupied secondary homonym in Quedius]; Type locality: Aculco).

  • — Fauvel, 1866a: 339 [= 1866b: 90] (Quedius; characters; Chile).

  • — Germain, 1903: 435 (Quedius; characters; Chile).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 404 (Loncovilius; synonym of heeri).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of heeri).

  • — Scheerpeltz, 1972: 24 (Quedius; subgenus Apoquedius; type species of subgenus; cited as valid species; Argentina).

  • — Herman, 2001: 26 (Loncovilius; valid name).

  • Distribution: Chile, Argentina.

heeri Blackwelder, 1944: 144 (Quedius; replacement name for aeneipennis Fairmaire and Germain; [Note: Unnecessary replacement name; the senior name in Quedius, aeneipennis Heer, was originally published without a description, so is an unavailable name]).

  • — Coiffait and Saiz, 1966: 404 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Herman, 2001: 26 (Loncovilius; synonym of aeneipennis).

brevis Saiz, 1971: 384 (Loncovilius; subgenus Lienturius; Type locality: Chile: Chiloé).

  • Distribution: Chile.

chilensis Bernhauer and Schubert, 1914: 332 (Philonthus; replacement name for cribripennis Germain; [Note: This name is older than germaini. It is preoccupied as secondary homonym by chilensis Solier, a name described in Staphylinus and formerly in Philonthus but now in Bisnius]).

  • — Scheerpeltz, 1933: 1344 (Philonthus; synonym of germaini).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; synonym of germaini).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of germaini).

  • — Herman, 2001: 26 (Loncovilius; valid name).

  • Distribution: Chile.

cribripennis Germain, 1903: 412 (Philonthus; [preoccupied]; Type locality: los arenales maritimos de las provincias centrales, Chile).

  • — Bernhauer and Schubert, 1914: 332 (Philonthus; synonym of chilensis).

  • — Scheerpeltz, 1933: 1344 (Philonthus; synonym of germaini).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; synonym of germaini).

  • — Coiffait and Saiz, 1968: 365 (Loncovilius; synonym of germaini).

germaini Scheerpeltz, 1933: 1344 (Philonthus; replacement name for cribripennis Germain).

  • — Coiffait and Saiz, 1966: 406 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971: 382 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Herman, 2001: 26 (Loncovilius; synonym of chilensis).

cribripennis Germain, 1903, see: chilensis Bernhauer and Schubert, 1914.

discoideus Fairmaire and Germain, 1861: 427 (Heterothops; Type locality: Santiago; Quillota).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (Heterothops; characters; Chile).

  • — Germain, 1903: 409, 444 (Bolitobius; cited as a new species as “discoideus mihi” with discoideus Fairmaire and Germain as a synonym; characters).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 407 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1969: 3, 4 (Loncovilius; characters; collecting notes; Chile).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1971a (Loncovilius; ecological notes).

  • — Saiz, Solervicens, and Ojeda, 1989: 50 (Loncovilius; collecting notes; habitus; Chile).

  • Distribution: Chile.

germaini Scheerpeltz, 1933, see: chilensis Bernhauer and Schubert, 1914.

heeri Blackwelder, 1944, see: aeneipennis Fairmaire and Germain, 1861.

leiocephalus Solier, 1849: 318 (Staphylinus; Type locality: Chile).

  • — Kraatz, 1859a: 5 (Quedius; notes).

  • — Fairmaire and Germain, 1861: 429 (Quedius; characters; Chile).

  • — Fauvel, 1864a: 124 (Quedius; notes; Chile).

  • — Fauvel, 1866a: 338 [= 1866b: 89] (Quedius; characters; Chile).

  • — Germain, 1903: 428 (Quedius; characters; notes; Chile).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 403 (Loncovilius; subgenus Lienturius; characters; Chile).

  • — Saiz, 1969a: 9 (Loncovilius; subgenus Lienturius; notes).

  • — Saiz, 1971: 382, 383 (Loncovilius; subgenus Lienturius; characters; Chile).

  • Distribution: Chile.

lividipennis Fairmaire and Germain, 1861: 430 (Quedius; Type locality: Chillan).

  • — Fauvel, 1866a: 338 [= 1866b: 89] (Quedius; characters; Chile).

  • — Germain, 1903: 443 (Loncovilius; characters).

  • — Bernhauer and Schubert, 1916: 427 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 409 (Loncovilius; subgenus Loncovilius; characters; Chile).

  • — Saiz, 1971: 385 (Loncovilius; subgenus Loncovilius; Chile).

  • — Scheerpeltz, 1972: 23 (Quedius; subgenus Loncovilius; Argentina).

  • Distribution: Chile, Argentina.

semiflavus Fairmaire and Germain, 1861: 428 (Quedius; Type locality: Concepcion).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of lividipennis).

  • — Germain, 1903: 443 (Loncovilius; characters).

  • — Bernhauer and Schubert, 1916: 434 (Quedius; catalog).

  • — Coiffait and Saiz, 1966: 411 (Loncovilius; subgenus Loncovilius; characters; Chile).

  • — Saiz, 1971: 385 (Loncovilius; subgenus Loncovilius; Chile).

  • Distribution: Chile.

Lonia [1 species; Australian Region]

Lonia E. Strand, 1943: 96 (replacement name for Colonia Olliff). Type species: Colonia regalis Olliff, fixed by objective synonymy with Colonia.

  • — Blackwelder, 1952: 227 (type species).

Colonia Olliff, 1887: 493 (species included: regalis; preoccupied by Gray, 1829, and Schaufuss, 1850). Type species: Colonia regalis Olliff, fixed by monotypy.

  • — Bernhauer and Schubert, 1914: 397 (world catalog; 1 species).

  • — Blackwelder, 1952: 102 (type species: regalis).

regalis Olliff, 1887: 494 (Colonia; Type locality: Lismore, Richmond River, New South Wales).

  • — Bernhauer and Schubert, 1914: 397 (Colonia; catalog).

  • — Lea, 1925: 229 (Colonia; Australia).

  • Distribution: Australia.

Mimosticus [1 species; Neotropical Region]

Mimosticus Sharp, 1884: 327 (species included: viridipennis). Type species: Mimosticus viridipennis Sharp, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 1 species).

  • — Blackwelder, 1944: 145 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 249 (type species: viridipennis).

viridipennis Sharp, 1884: 328 (Mimosticus; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

  • — Bernhauer and Schubert, 1916: 437 (Mimosticus; catalog).

  • Distribution: Panama.

Paratolmerus [2 species; Palaearctic and Oriental Regions]

Paratolmerus Cameron, 1932: 169 (species included: pilosiventris). Type species: Paratolmerus pilosiventris Cameron, fixed by monotypy.

  • — Scheerpeltz, 1933: 1376 (world catalog supplement).

  • — Blackwelder, 1952: 294 (type species: pilosiventris).

  • — Smetana, 1988: 335 (characters).

pilosiventris Cameron, 1932: 170 (Paratolmerus; Type locality: Sikkim: Gopaldhara, Rungbong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1376 (Paratolmerus; catalog).

  • — Smetana, 1988: 336 (Paratolmerus; lectotype designation; characters; Sikkim; Nepal).

  • Distribution: India, Nepal.

siamensis Rougemont, 1991: 221 (Paratolmerus; Type locality: Thailand: Kanchanaburi Prov.: river Kwai Noi at Ban Sai Yok).

  • Distribution: Thailand.

Parisanopus [1 species; Neotropical Region]

Parisanopus Brèthes, 1900: 216 (species included: castaneipennis). Type species: Parisanopus castaneipennis Brèthes, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 415 (world catalog; 1 species).

  • — Scheerpeltz, 1933: 1426 (world catalog supplement).

  • — Blackwelder, 1944: 144 (in checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 294 (type species: castaneipennis).

castaneipennis Brèthes, 1900: 216 (Parisanopus; Type locality: Palermo, Buenos Aires).

  • — Bernhauer and Schubert, 1916: 415 (Parisanopus; catalog).

  • — Scheerpeltz, 1933: 1426 (Parisanopus; catalog; Argentina; Paraguay).

  • Distribution: Argentina, Paraguay.

Philonthellus [1 species; Neotropical Region]

Philonthellus Bernhauer, 1939: 13 (subgenus of Philonthus; species included: chilenus). Type species: Philonthus (Philonthellus) chilenus Bernhauer, by monotypy.

  • — Blackwelder, 1952: 301 (type species: chilenus).

  • — Coiffait and Saiz, 1968: 347, 349 (valid genus; characters).

chilenus Bernhauer, 1939: 13 (Philonthus; subgenus Philonthellus; Type locality: Isla Calbuco, Süd-Chile).

  • — Coiffait and Saiz, 1968: 349 (Philonthellus; characters; Chile).

  • — Saiz, 1971b: 341 (Philonthellus; Chile).

  • — Scheerpeltz, 1972: 18 (Philonthus; Argentina).

  • Distribution: Chile, Argentina.

evitendus Tottenham, 1939d: 234 (Philonthus; replacement name for chilensis Bernhauer; [Note: Unnecessary replacement name, Bernhauer used chilenus]).

  • — Coiffait and Saiz, 1968: 349 (Philonthellus; synonym of chilenus).

evitendus Tottenham, 1939, see: chilenus Bernhauer, 1939.

Pseudorientis [3 species; Palaearctic Region]

Pseudorientis Watanbe, 1970: 70 (species included: shinobuae). Type species: Pseudorientis shinobuae Watanabe, fixed by original designation and monotypy.

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Smetana, 1977b: 181 (characters).

  • — Shibata, 1984: 115 (checklist of species of Japan).

  • — Smetana, 1995c: 345 (key to species).

gongga Smetana, 1995c: 344 (Pseudorientis; Type locality: China: Sichuan, Gongga Shan above Camp 3, 3050 m).

  • — Smetana, 1998d: 98 (Pseudorientis; characters of female; China).

  • Distribution: China.

shinobuae Watanabe, 1970: 72 (Pseudorientis; Type locality: Japan: Honshu: Gumma Pref.: near Lake Marunuma, Oku-Nikko).

  • Distribution: Japan.

uenoi Smetana, 1995c: 341 (Pseudorientis; Type locality: China: Yunnan, Heishui, 35 km N Lijiang, 27°13′N, 100°19′E).

  • Distribution: China.

Quediocafus [3 species; Australian Region]

Quediocafus Cameron, 1945d: 791 (species included: hudsoni). Type species: Quediocafus hudsoni Cameron, fixed by original designation and monotypy.

  • — Blackwelder, 1952: 334 (type species: hudsoni).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (endemic to New Zealand).

hudsoni Cameron, 1945d: 791 (Quediocafus; Type locality: New Zealand: Hokitika).

  • Distribution: New Zealand.

insolitus Sharp, 1886a: 379 (Quedius; Type locality: New Zealand; Dunedin).

  • — Broun, 1893: 1029 (Quedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Newton, 1996: 160 (Quediocafus).

  • — Marris, 2000: 175 (Quediocafus; habitat; Bollons Island [Antipodes]; also known from New Zealand).

  • Distribution: New Zealand, Antipodes Islands.

taieriensis Broun, 1894: 424 (Quedius; Type locality: Taieri, Otago).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Newton, 1996: 160 (Quediocafus).

  • Distribution: New Zealand.

Quediomacrus [2 species; Neotropical Region]

Quediomacrus Sharp, 1884: 339 (species included: puniceipennis, pollens). Type species: Quedius puniceipennis Solsky, fixed by subsequent designation by R. Lucas, 1920: 563.

  • — Bernhauer and Schubert, 1916: 410 (world catalog; 2 species).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Blackwelder, 1952: 334 (type species).

pollens Sharp, 1884: 339 (Quediomacrus; Type locality: Guatemala, near the city and at Purula).

  • — Bernhauer and Schubert, 1916: 410 (Quediomacrus; catalog).

  • Distribution: Guatemala.

puniceipennis Solsky, 1868a: 123 (Quedius; Type locality: Mexique).

  • — Sharp, 1884: 339 (Quediomacrus; Mexico).

  • — Bernhauer and Schubert, 1916: 410 (Quediomacrus; catalog).

  • — Smetana, 1971b: 4 (Quediomacrus; Mexico).

  • — Frank, 1982: 45 (Quediomacrus; list of parasites).

  • Distribution: Mexico.

Quediomimus [1 species; Australian Region]

Quediomimus Cameron, 1948c: 724 (species included: brookesi). Type species: Quediomimus brookesi Cameron, fixed by original designation and monotypy.

brookesi Cameron, 1948c: 725 (Quediomimus; Type locality: New Zealand: Katikati: Bay of Plenty).

  • — Klimaszewski, Newton, and Thayer, 1996: 151 (Quediomimus; endemic to New Zealand).

  • Distribution: New Zealand.

Quediopsis [2 species; Australian Region]

Quediopsis Fauvel, 1878g: 560 (species included: lugubris, abdominalis). Type species: Quediopsis lugubris Fauvel, fixed by subsequent designation by Blackwelder, 1952: 335.

  • — Duvivier, 1883: 136 (catalog).

  • — Bernhauer and Schubert, 1916: 437 (world catalog; 2 species).

abdominalis Fauvel, 1878g: 561 (Quediopsis; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 437 (Quediopsis; catalog).

  • Distribution: Australia.

lugubris Fauvel, 1878g: 560 (Quediopsis; Type locality: Australie méridionale, Victoria).

  • — Bernhauer and Schubert, 1916: 437 (Quediopsis; catalog).

  • Distribution: Australia.

Quedius [789 species (3 doubtful); Neotropical, Nearctic, Ethiopian, Palaearctic, Oriental, and Australian Regions]

Quedius Stephens, 1829a: 22 (species included: tristis, variabilis, fulvipes, impressus, rufitarsis, nitidus and cyanipennis along with 32 unavailable names; [Note: The same questions and controversy concerning the authorship and date of publication for Philonthus apply to Quedius. The issue is discussed in my “note” attached to the citation of the original description for Philonthus]). Type species: Staphylinus levicollis Brullé, fixed by subsequent designation in Opinion 1851 (ICZN, 1996: 215) [Note: Curtis, (1837: pl. 638) fixed Staphylinus tristis Gravenhorst as the type species Quedius. Gravenhorst, who misidentified the species, attributed the name to Fabricius so his use of the name is unavailable. Blackwelder (1952: 335) fixed Quedius fulgidus Fabricius, a species that was in Staphylinus, Ocypus, or Pseudocypus since its description. The problem is discussed by Newton, 1995: 49 and action by the Commission resolved the problem].

  • — Curtis, 1829: 24 (catalog; Britain).

  • — Stephens, 1829: 277 (catalog; Britain).

  • — Stephens, 1832: 200, 214 (characters).

  • — Curtis, 1837: pl. 638 (characters; type species; habitat; larval characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 483 (characters).

  • — Stephens, 1839: 385 (characters).

  • — Erichson, 1840: 523 (characters).

  • — Schaum, 1852: 26 (list of species; Europe).

  • — Lacordaire, 1854: 84 (characters; notes; list of species).

  • — Fairmaire and Laboulbène, 1856: 535 (characters).

  • — Jacquelin du Val, 1856: 37 (characters).

  • — Giebel, 1856: 72 (fossil species).

  • — Kraatz, 1857d: 486 (characters).

  • — Redtenbacher, 1857: 199 (characters; key to Austrian species).

  • — Thomson, 1858: 30 (characters).

  • — G. Waterhouse, 1858: 22 (catalog of British species).

  • — Thomson, 1859: 25 (characters; type species: molochinus).

  • — Schaum, 1859: 25 (catalog; European species).

  • — Thomson, 1860: 173 (characters).

  • — LeConte, 1861a: 64 (characters).

  • — LeConte, 1863: 23 (list of North American species).

  • — Gredler, 1863: 99 (list of species from Tirol).

  • — Fauvel, 1866a: 337 [= 1866b: 88] (characters).

  • — Wencker and Silbermann, 1866: 27 (list of species; collecting notes; France).

  • — Gemminger and Harold, 1868: 568 (catalog).

  • — Jacquelin du Val, 1868: 69 (catalog; European species).

  • — Sharp, 1871d: 11 (list of British species).

  • — Fauvel, 1874: 266 [= 1874a: 492] (characters; larval characters; key to species).

  • — Redtenbacher, 1874: 197 (characters; key to Austrian species).

  • — Oustalet, 1874: 171 (characters; description of fossil species).

  • — Fauvel, 1875a: XXXII [= 1875b: 234] (catalog).

  • — Mulsant and Rey, 1876: 613 [= 1877: 469] (characters; key to French species).

  • — Sharp, 1876: 100 (notes).

  • — Fauvel, 1877: 267 (notes; key to species).

  • — Provancher, 1877: 244, 728 (characters; key to species).

  • — Fauvel, 1878f: 272 (characters; notes; key to New Guinea species).

  • — Horn, 1878: 154 (characters; notes; key to North American species).

  • — Heyden, 1880: 71 (list of species of Siberian region).

  • — Duvivier, 1883: 137 (catalog).

  • — Sharp, 1884: 328 (notes; Mexican and Central American species).

  • — Lynch, 1884: 119 (characters).

  • — Sharp, 1888: 226 (adult and larval characters; key to British species).

  • — Seidlitz, 1891: 395 (key to some European species).

  • — Seidlitz, 1891a: 421 (key to some European species).

  • — Heyden, Reitter, and Weise, 1891: 99 (list of species of Europe and Caucasus).

  • — Heyden, 1893: 41 (list of species of Siberian region).

  • — Ganglbauer, 1895: 391 (characters; key to middle European species).

  • — Heyden, 1896: 32 (list of species of Siberian region).

  • — Fauvel, 1897d: 305 (catalog of species of Barbary; Canaries).

  • — Germain, 1903: 414 (characters; notes).

  • — Heyden, Reitter, and Weise, 1906: 172 (list of species of Europe and Caucasus).

  • — Porta, 1907: 90 (key to Italian species).

  • — Reitter, 1908: 106, 108 (characters; key to subgenera).

  • — Blatchley, 1910: 369 (characters; key to Indiana species).

  • — Petri, 1912: 76 (list of species of Siebenbürgen).

  • — Johansen, 1914: 349 (characters; key to species).

  • — Casey, 1915: 398 (type species: Quedius molochinus).

  • — Bernhauer and Schubert, 1916: 417 (world catalog; 278 species).

  • — Leng, 1920: 109 (catalog of North American species).

  • — Munster, 1923: 186 (characters; key to subgenera and Scandinavian species).

  • — Gridelli, 1924d (anatomy; characters; key to Palaearctic subgenera and species; revision of species).

  • — Winkler, 1925: 388 (catalog for Palaearctic region).

  • — Cameron, 1925: 72 (catalog of Indian species).

  • — Porta, 1926: 109 (characters; key to species of Italy).

  • — Bruch, 1928a: 429 (catalog; Argentina).

  • — Portevin, 1929: 337 (characters; key to French species).

  • — Roubal, 1930: 387 (catalog; Slovakia).

  • — Cameron, 1932: 280 (characters; key to subgenera and species of British India).

  • — Joy, 1932: 108 (key to British species).

  • — Scheerpeltz, 1933: 1427 (world catalog supplement).

  • — Normand, 1935: 372 (list of species of Tunisia).

  • — Voris, 1939 (egg, larval, and pupal characters).

  • — Paulian, 1941: 259 (larval characters).

  • — Blackwelder, 1944: 144 (checklist of species from Mexico, Central and South America, and the West Indies).

  • — Kloet and Hincks, 1945: 167 (list of British species).

  • — Jeannel and Jarrige, 1949: 377 (cave species; notes).

  • — Tottenham, 1949: 376, 421 (type species: levicollis Brullé = tristis Gravenhorst; checklist of British species).

  • — V. Hansen, 1952: 115 (characters; key to Danish species).

  • — Hatch, 1957: 215 (characters; key to subgenera and species of Pacific Northwest).

  • — Kocher, 1958: 157 (checklist of species; Morocco).

  • — Smetana, 1958: 327 (characters; type species: tristis Gravenhorst; key to Czechoslovakian species).

  • — Smetana, 1959a: 215 (list of species collected in Albania).

  • — Smetana, 1960e: 150 (key to European species and subgenus).

  • — Coiffait, 1961a: 58 (key to and list of species of molochinus complex).

  • — Scheerpeltz, 1962b: 267 (characters).

  • — Ferreira, 1962a: 18 (list of Portuguese species).

  • — Coiffait, 1963a: 372 (key to subgenera).

  • — Székessy, 1963: 61 (characters).

  • — Palm, 1963b: 124 (characters; key to subgenera and species of Sweden).

  • — Lohse, 1964: 205 (characters; key to central European subgenera and species).

  • — Smetana, 1965: 38 (key to Nearctic species of subgenus Quedius).

  • — Coiffait and Saiz, 1966: 411 (Chilean species).

  • — Pototskaia, 1967: 79 (larval characters).

  • — Coiffait and Saiz, 1968: 365 (list of Chilean species).

  • — Scheerpeltz, 1968a: 87 (catalog of Austrian species).

  • — Bolov, 1969a: 328 (collecting notes for some species; Russia).

  • — Kasule, 1970: 57, 59 (larval characters; notes).

  • — Smetana, 1971 (revision of and key to subgenus and species of America north of Mexico; characters).

  • — Korge, 1971a: 37 (characters).

  • — Shibata, 1973c: 63 (checklist of species of Taiwan).

  • — Bordoni, 1973b: 41 (key to subgenera).

  • — Bordoni, 1973j: 710 (list of species from islands near Sicily).

  • — Tikhomirova, 1973a: 188 (checklist of species of USSR).

  • — Moore and Legner, 1975: 56 (catalog; North America north of Mexico).

  • — Bordoni, 1976b: 88 (revision of and key to Italian species).

  • — Pototskaia, 1976: 171 (larval characters).

  • — Smetana, 1977b: 181 (characters).

  • — Pope, 1977: 31 (list of British species).

  • — Coiffait, 1978: 9 (adult and larval characters; key to subgenera and species of western Palaearctic region).

  • — Smetana, 1978b: 87 (key to species of Pacific coastal area of Siberia, from Kamchatka Peninsula northwards).

  • — Topp, 1978: 312 (larval characters).

  • — Moore and Legner, 1979: 47 (characters; notes).

  • — Muona, 1979: 15 (list of Scandinavian species).

  • — Uhlig, 1979: 255 (some species collected in Germany).

  • — M. Dvořák, 1979: 116 (some species collected in Slovakia).

  • — Uhlig, Vogel, and Sieber, 1980: 248 (some species collected in Germany).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 119 (catalog; Poland).

  • — Spahr, 1981: 96 (references to specimens from amber and copal).

  • — Uhlig and Vogler, 1981: 122 (list of some species collected in Germany; habitat and collecting notes).

  • — Hammond, 1984: 207 (checklist of species of Borneo).

  • — Tóth, 1984: 102 (characters; habitus; key to species of Hungary).

  • — Shibata, 1984: 119 (checklist of species of Japan).

  • — Frank, 1986a: 371 (checklist of species of Florida).

  • — Shibata, 1986a: 125 (supplemental list of species from Taiwan).

  • — Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

  • — Segers, 1986: 49 (checklist of species; Belgium).

  • — Dettner, 1987: 30 (no defensive glands).

  • — Ádám, 1987: 152 (list of six species with collecting notes; Hungary).

  • — Smetana, 1988: 181 (characters; notes; key to genera of Himalayan region; discussion of difficulty of defining subgenera).

  • — Lohse and Lucht, 1989: 171 (notes; supplement to key to species of Lohse, 1964).

  • — Angelini, 1991: 193 (list of species from high elevations of Calabria, Italy).

  • — M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

  • — Campbell and Davies, 1991: 121 (checklist of species; Alaska and Canada).

  • — Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

  • — Siitonen, 1993: 230 (list of Finnish species).

  • — Welch, 1993: 229 (ovariole structure and development).

  • — Smetana, 1993b: 49 (list of species of Czech Republic and Slovakia).

  • — Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).

  • — Smetana, 1995: 14, 24 (revision of species of Taiwan; key to subgenera and species of Taiwan; cladistic analysis).

  • — Newton, 1995: 51 (requested that the International Commission on Zoological Nomenclature set aside all previous designations of type species for Quedius and designate Staphylinus levicollis Brullé as type species).

  • — M. Hansen, 1996: 107 (list of Danish species).

  • — Ciceroni and Zanetti, 1995: 31 (list of species of Italy).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 244 (additions to list of Danish species).

  • — Cho, 1996b: 110 (key to species of Korea).

  • — ICZN, 1996: 215 (type species designated as Staphylinus levicollis Brullé; placed on Official List of Generic Names in Zoology).

  • — Downie and Arnett, 1996: 388 (characters; key to subgenera and species of Northeastern North America).

  • — Lundgren, 1998: 52 (list of species from Florida; records for other States listed for each species).

  • — M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

  • — Staniec, 1999a: 55 (key to known pupae of Poland).

  • — Newton, Thayer, Ashe, and Chandler, 2000: 393 (91 Nearctic species; notes; characters in key).

  • — Naomi, Kuranishi, Saito, and Maruyama, 2000: 110 (collected from Kuril Islands and Kamchatka).

  • — Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 60 (necrophilous species in Mexico).

Raphirus Stephens, 1829a: 23 (species included: semiobscurus, attenuatus, praecox along with six unavailable names). Type species: Staphylinus attenuatus Gravenhorst, fixed by subsequent designation by Westwood, 1838a: 16.

  • — Stephens, 1829: 282 (catalog; Britain).

  • — Stephens, 1832: 200 (characters).

  • — Stephens, 1833: 241 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Erichson, 1839a: 483 (synonym of Quedius).

  • — Stephens, 1839: 385 (characters).

  • — Lacordaire, 1854: 84 (synonym of Quedius).

  • — Kraatz, 1857d: 486 (synonym of Quedius).

  • — G. Waterhouse, 1858: 22 (synonym of Quedius).

  • — Thomson, 1859: 26 (characters; type species: attenuatus).

  • — Thomson, 1860: 177 (characters).

  • — Mulsant and Rey, 1876: 759 [= 1877: 615] (subgenus of Quedius; characters; key to species).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 109 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 398, 416 (characters; type species: fulvicollis; notes; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 87 (subgenus of Quedius; characters; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Tottenham, 1945: 70 (Genus dubium because attenuatus, the species designated as type species, was misidentified by Stephens).

  • — Blackwelder, 1952: 336 (type species).

  • — Hatch, 1957: 215, 228 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 327, 366 (synonym of Quedius).

  • — Lohse, 1964: 212 (subgenus of Quedius; characters; key to central European species).

  • — Smetana, 1971: 39, 183 (subgenus of Quedius; characters; notes; key to North American species).

  • — Coiffait, 1978: 9, 13, 125 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1988: 231 (notes; key to species of Himalayan region).

  • — Shibata, 1984: 134 (subgenus of Quedius; checklist of species of Japan).

  • — Ádám, 1987: 152 (list of two species with collecting notes; Hungary).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 24, 25, 65 (subgenus of Quedius; characters; key to species of Taiwan).

Microsaurus Dejean, 1833: 61 (species included: lateralis, fuliginosus, molochinus, nitidus, scitus, laevigatus, ochripennis, praecox, attenuatus, scintillans, boops). Type species: Staphylinus lateralis Gravenhorst, fixed by subsequent designation by Westwood, 1838a: 16.

  • — Stephens, 1835: 435 (characters).

  • — Westwood, 1838a: 16 (characters; type species).

  • — Stephens, 1839: 384 (characters).

  • — Lacordaire, 1854: 84 (synonym of Quedius).

  • — Kraatz, 1857d: 486 (synonym of Quedius).

  • — Thomson, 1859: 25 (characters; type species: lateralis).

  • — Thomson, 1860: 174 (characters).

  • — Mulsant and Rey, 1876: 685 [= 1877: 541] (subgenus of Quedius; characters; key to species).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 109 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 398, 408 (characters; type species: Quedius fulgidus; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (subgenus of Quedius; revision of concept; characters; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Tottenham, 1949: 376, 421 (type species: lateralis Gravenhorst; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 247 (type species).

  • — Hatch, 1957: 215, 219 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1958: 327 (subgenus of Quedius).

  • — Lohse, 1964: 206 (subgenus of Quedius; characters; key to central European species).

  • — Smetana, 1971: 38, 49 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973g: 124 (subgenus of Quedius; characters; key to species groups).

  • — Bordoni, 1974: 1 (characters; notes; anatomy; key to species and species groups of Italy).

  • — Smetana, 1976: 113 (subgenus of Quedius; key to species of Mexico).

  • — Coiffait, 1978: 9, 13, 15 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Shibata, 1984: 119 (subgenus of Quedius; checklist of species of Japan).

  • — Ádám, 1987: 152 (cited as valid genus; list of four species with collecting notes; Hungary).

  • — Smetana, 1988: 185 (subgenus of Quedius; characters; key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 24, 25 (subgenus of Quedius; characters; revision of and key to species of Taiwan).

  • — Smetana, 1996: 19 (subgenus of Quedius; key to species of apicicornis and beesoni species groups from China).

Aemulus Gistel, 1834: 8 (species included: lateralis, fuliginosus, molochinus, nitidus, scitus, boops). Type species: Staphylinus fuliginosus Gravenhorst, fixed by subsequent designation by Blackwelder, 1952: 40.

  • — Blackwelder, 1952: 40 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

Thanatomanes Gistel, 1856: 388 (species included: lateralis, fulgidus, xanthopus, impressus, molochinus, fuliginosus, boops, carbonarius, attenuatus). Type species: Staphylinus impressus Panzer, fixed by subsequent designation by Blackwelder, 1952: 384 [Note: Blackwelder cited Gravenhorst as the author of the type species of Thanatomanes; Gravenhorst did not describe impressus, he attributed it to Panzer, furthermore, Gistel cited Panzer as the author].

  • — Blackwelder, 1952: 384 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

Ediquus Mulsant and Rey, 1876: 615, 616 [= 1877: 471, 472] (subgenus of Quedius; species included: longicornis, microps). Type species: Quedius microps Gravenhorst, fixed by subsequent designation by Tottenham, 1939d: 229.

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 109 (subgenus of Quedius; characters; key to species).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (synonym of Microsaurus).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (synonym of Quedius).

  • — Tottenham, 1939d: 229 (type species designation: microps).

  • — Tottenham, 1949: 376, 422 (type species: microps Gravenhorst; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 144 (type species).

  • — Smetana, 1958: 327 (synonym of Quedius).

  • — Smetana, 1971: 38 (synonym of Quedius).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Microsaurus).

  • — Smetana, 1988: 181 (synonym of Quedius subgenus Microsaurus).

  • — Smetana, 1995: 24 (synonym of Quedius).

Sauridus Mulsant and Rey, 1876: 616, 700 [= 1877: 472, 556] (subgenus of Quedius; species included: ochropterus, picipes, nigriceps, robustus, peltatus, dubius, montanus, anceps, cyanescens, umbrinus, suturalis, limbatus, bicolor, oblitteratus, maurorufus, riparius, kraatzi, lucidulus, scintillans). Type species: Staphylinus picipes Mannerheim, fixed by subsequent designation by Tottenham, 1940: 50.

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 109 (characters).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Tottenham, 1940: 50 (type species: picipes).

  • — Tottenham, 1949: 377, 423 (type species: picipes Mannerheim; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 344 (synonym of Raphirus; type species).

  • — Hatch, 1957: 215, 226 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Coiffait, 1963a: 373 (subgenus of Quedius; key to species; key to species groups and species using aedeagus).

  • — Coiffait, 1978: 9, 13, 67 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Ádám, 1987: 153 (cited as valid genus; list of two species with collecting notes; Hungary).

  • — Smetana, 1988: 181 (synonym of Quedius subgenus Raphirus).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Quedionuchus Sharp, 1884: 336 (species included: laevigatus, impunctus, spinipes, femoralis, angustus, nigerrimus). Type species: Quedius impunctus Solsky, fixed by subsequent designation by Blackwelder, 1952: 334.

  • — Sharp, 1888: 245 (characters; key to British species).

  • — Fauvel, 1895: 274 (synonym of Quedius).

  • — Ganglbauer, 1895: 391 (characters; subgenus of Quedius).

  • — Reitter, 1909: 108, 111 (subgenus of Quedius; characters; key to species).

  • — Casey, 1915: 397, 399 (notes; type species: Quedius laevigatus).

  • — Gridelli, 1924d: 74 (subgenus of Quedius; key to species).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Joy, 1932: 107 (key to British species).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Tottenham, 1939d: 229 (type species designation: laevigatus).

  • — Tottenham, 1949: 377, 422 (type species: plagiatus Mannerheim = laevigatus Gyllenhal; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 334 (subgenus of Quedius).

  • — Hatch, 1957: 215, 229 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1958: 328, 357 (subgenus of Quedius).

  • — Korge, 1964: 118 (key to species of western Palaearctic).

  • — Kasule, 1970: 57, 59 (cited as valid genus; larval characters).

  • — Smetana, 1971: 39, 147 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973b: 45 (subgenus of Quedius; characters).

  • — Smetana, 1976a (subgenus of Quedius; review of and key to species of Central America).

  • — Coiffait, 1978: 9, 13, 53 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Shibata, 1984: 129 (subgenus of Quedius; checklist of species of Japan).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

Ediquus Reitter, 1887: 211 (subgenus of Quedius; species included: przewalskii; preoccupied by Mulsant and Rey, 1876). Type species: Quedius (Ediquus) przewalskii Reitter, fixed by monotypy.

  • — Blackwelder, 1952: 144 (type species: przewalskii).

Prionidus Bernhauer, 1907a: 287 (subgenus of Quedius; species included: iheringi; preoccupied by Uhler, 1886). Type species: Quedius (Prionidus) iheringi Bernhauer, fixed by monotypy.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Blackwelder, 1952: 321 (type species: iheringi).

Euryquedius Reitter, 1909: 108 (subgenus of Quedius; species included: curtus). Type species: Quedius curtus Erichson, fixed by original designation and monotypy.

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1427 (Velleiopsis; catalog).

  • — Blackwelder, 1952: 160 (type species: curtus).

  • — Bordoni, 1973b: 42 (subgenus of Quedius; characters).

  • — Coiffait, 1978: 9, 13, 15 (subgenus of Quedius; characters; key to species of western Palaearctic region).

Tenebrobius Rambousek, 1915a: 27, 29 (subgenus of Quedius; species included: bernhaueri). Type species: Quedius (Tenebrobius) bernhaueri Rambousek, fixed by monotypy.

  • — Rambousek, 1916: 130 (description; subgenus of Quedius; cited as a new genus).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Gridelli, 1924d: 18 (synonym of Microsaurus).

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 375 (type species: bernhaueri).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Microsaurus).

Quediellus Casey, 1915: 398, 402 (species included: nanulus, debilis, helenae, humilis, densiventris). Type species: Quedius debilis Horn, fixed by original designation.

  • — Casey, 1915: 398 (type species: Quedius debilis).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 334 (type species: debilis).

  • — Smetana, 1971: 39 (synonym of Quedius).

Quediochrus Casey, 1915: 398, 420 (species included: spelaeus, quadriceps). Type species: Quedius spelaeus Horn, fixed by original designation and monotypy.

  • — Casey, 1915: 398 (type species: Quedius spelaeus).

  • — Bernhauer and Schubert, 1916: 417 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 334 (type species: spelaeus).

  • — Hatch, 1957: 230 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 38 (synonym of Quedius).

Paraquedius Casey, 1915: 397, 400 (species included: puncticeps). Type species: Quedius puncticeps Horn, fixed by original designation and monotypy.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 293 (type species: puncticeps).

  • — Hatch, 1957: 214, 215 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 40 (subgenus of Quedius).

  • — Korge, 1971a: 36 (subgenus of Quedius; characters).

Distichalius Casey, 1915: 398, 404 (species included: capucinus, nevadensis, marginalis, brunnipennis, agnatus, virginicus, oculeus, sparsus). Type species: Staphylinus capucinus Gravenhorst, fixed by original designation.

  • — Casey, 1915: 398 (type species: Quedius capucinus).

  • — Scheerpeltz, 1933: 1428 (cited as Distichalis; subgenus of Quedius).

  • — Blackwelder, 1952: 130 (type species: capucinus).

  • — Hatch, 1957: 215, 217 (subgenus of Quedius; characters; key to species of Pacific Northwest).

  • — Smetana, 1971: 39, 153 (subgenus of Quedius; characters; key to species of Canada and USA).

  • — Bordoni, 1973b: 48 (subgenus of Quedius; characters).

  • — Coiffait, 1978: 9, 13, 52 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1988: 227 (key to species of Himalayan region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1995: 25, 57 (subgenus of Quedius; characters; key to species of Taiwan).

Megaquedius Casey, 1915: 421 (species included: laxatus, explanatus, manitobensis). Type species: Quedius explanatus LeConte, fixed by original designation.

  • — Casey, 1915: 399, 421 (type species: Quedius explanatus).

  • — Leng, 1920: 110 (catalog of North American species).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 236 (type species: explanatus).

  • — Hatch, 1957: 230 (subgenus of Quedius; characters; species of Pacific Northwest).

  • — Smetana, 1971: 38, 42 (subgenus of Quedius; key to species of USA and Canada).

Anastictodera Casey, 1915: 421 (species included: compransor). Type species: Quedius compransor Fall, fixed by monotypy.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 50 (type species: compransor).

  • — Smetana, 1971: 38 (synonym of Quedius).

Cyrtoquedius Bernhauer, 1917h: 92 (subgenus of Quedius; species included: labiatus, frenatus, clypealis, basiventris, jocosus, graciliventris, concolor, arrogans, falvicaudus, rufinasus, mexicanus, verecundus, protensus, flavinasus, ochropygus, laeviventris). Type species: Quedius basiventris Sharp, fixed by subsequent designation by Blackwelder, 1952: 115.

  • — Cameron, 1932: 280 (subgenus of Quedius).

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

  • — Blackwelder, 1952: 115 (subgenus of Quedius).

  • — Scheerpeltz, 1955a: 172 (key to neotropical species).

Quediopsis Portevin, 1929: 337 (subgenus of Quedius; species included: riparius, kraatzi; preoccupied by Fauvel, 1878). Type species: Quedius riparius F. Kellner, fixed by subsequent designation by Blackwelder, 1952: 335.

  • — Scheerpeltz, 1933: 1429 (synonym of Quedius).

  • — Smetana, 1958: 328 (synonym of Quedius).

  • — Smetana, 1971: 40 (synonym of Quedius).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Quediops).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

Arphirus Tottenham, 1945: 70 (proposed as replacement for Raphirus Stephens because he considered its type species to be of uncertain identity and the genus to be a “genus dubium”). Type species: Staphylinus semiobscurus Marsham, by original designation.

  • — Tottenham, 1948a: 243 (may be same as Sauridus; characters and key to species).

  • — Tottenham, 1949: 377, 422 (type species: semiobscurus Marsham = rufipes Erichson; subgenus of Quedius; checklist of British species).

  • — Blackwelder, 1952: 63 (type species: semiobscurus).

  • — Hatch, 1957: 228 (synonym of Raphirus).

  • — Smetana, 1958: 328 (synonym of Quedius).

  • — Smetana, 1971: 40 (synonym of Quedius).

Farus Blackwelder, 1952: 165 (replacement name for Ediquus Reitter; subgenus of Quedius). Type species: Quedius przewalskii Reitter, fixed by objective synonymy with Ediquus.

Pridonius Blackwelder, 1952: 321 (subgenus of Quedius; replacement name for Prionidus Bernhauer). Type species: Quedius iheringi Bernhauer, fixed by objective synonymy with Prionidus.

  • — Scheerpeltz, 1933: 1428 (subgenus of Quedius).

Quediops Coiffait, 1963: 373 (subgenus of Quedius; species included: plancus). Type species: Quedius plancus Erichson, fixed by original designation and monotypy.

  • — Bordoni, 1973g: 117 (subgenus of Quedius; characters; key to species).

  • — Coiffait, 1978: 9, 13, 63 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Microquedius Coiffait, 1967e: 419 (subgenus of Quedius; species included: alpestris, spurius, subalpestris, haberfelneri cited as haberfellneri, marcuzzi, xanthippae, auricomus). Type species: Philonthus alpestris Heer, fixed by original designation.

  • — Bordoni, 1974a: 204 (subgenus of Quedius; notes; key to species of Italy).

  • — Coiffait, 1978: 9, 13, 60 (subgenus of Quedius; characters; key to species of western Palaearctic region).

  • — Lohse and Lucht, 1989: 171 (subgenus of Quedius; notes).

  • — Smetana, 1993b: 50 (synonym of Quedius subgenus Raphirus).

Quedionthus Bordoni, 1973b: 53 (subgenus of Quedius; species included: riparius [Note: Replacement name for Quediopsis Portevin but not so stated]). Type species: Quedius riparius F. Kellner, fixed by original designation and objective synonymy with Quediopsis.

  • — Coiffait, 1977c: 136 (synonym of Quediops).

  • — Coiffait, 1978: 9 (synonym of Quedius subgenus Quediops).

abdominalis Eppelsheim, 1878a: 419 (Quedius; Type locality: Caucasus).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; catalog).

  • — Gridelli, 1924d: 25 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Coiffait, 1978: 158 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • Distribution: Caucasus.

aberrans Cameron, 1932, see: Indoquedius.

abietoides Coiffait, 1967 [Note: Nomen nudum], see: abietum Kiesenwetter, 1858.

abietum Kiesenwetter, 1858: 57 (Quedius; Type locality: Parnes bei Athen; Aetolische Alpen).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; characters; Illyria; Sicily; Greece; Sardinia; Morocco).

  • — Mulsant and Rey, 1876: 664 [= 1877: 520] (Quedius; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Morocco).

  • — Fairmaire, 1879: 161 (Quedius; characters; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Morocco; Tunisia; Algeria).

  • — Ganglbauer, 1895: 400 (Quedius; subgenus Microsaurus; characters; Illyria; Mediterranean region).

  • — Xambeu, 1900: 49 (Quedius; larval characters).

  • — Fauvel, 1902b: 114 (Quedius; Morocco; Algeria; Tunisia).

  • — Porta, 1907: 126 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; not yet in Germany).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; catalog).

  • — Gridelli, 1924d: 57 (Quedius; subgenus Microsaurus; characters; Sardinia; Sicily; Macedonia; Spain; Tunisia).

  • — Gridelli, 1929: 34 (Quedius; characters).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Roubal, 1941b: 125, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Normand, 1947a: 9 (Quedius; Tunisia).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Microsaurus; Italy).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Microsaurus; Crete).

  • — Scheerpeltz, 1965b: 407 (Quedius; Sicily).

  • — Korge, 1971a: 47 (Quedius; subgenus Microsaurus; Turkey).

  • — Bordoni, 1974: 48 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 171 (Quedius; subgenus Microsaurus; characters; Mediterranean region; North Africa).

  • — Tóth, 1984: 113 (Quedius; characters).

  • — Sparacio, 1995: 164 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Tunisia, Algeria, Morocco, Spain, France, Italy, “Illyria”, Macedonia, Greece, Turkey.

reboudi Fairmaire, 1877 ([Note: Nomen nudum (Articles 11.6, 12.1); although the original description for reboudi is cited as Petites Nouvelles Entomologiques 2: 111, the species is neither on page 111 nor on any other page of the volume and there is evidently no description of the species. The first publication of the name appears to have been in 1879, when it was cited as a junior synonym; all subsequent uses have been as a junior synonym]).

  • — Fairmaire, 1879: 161 (Quedius; synonym of abietum).

  • — Fauvel, 1897d: 306 (Quedius; species was not described).

  • — Porta, 1907: 126 (Quedius; synonym of abietum).

  • — Bernhauer and Schubert, 1916: 417 (Quedius; synonym of abietum).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog; the species not described).

  • — Bordoni, 1974: 48 (Quedius; synonym of abietum).

  • — Coiffait, 1978: 171 (Quedius; cited as “nomen nudum ?”; synonym of abietum).

abietoides Coiffait, 1967e: 393 (Quedius; subgenus Microsaurus; variety of abietum; [Note: Nomen nudum (Article 15.2)]; cited from Itxassou, Basses-Pyrénées).

abkasicus Coiffait, 1963a: 390, 410 (Quedius; subgenus Sauridus; [Note: Cited on page 390 in a key to the species of Sauridus but listed in Microsaurus on page 410]; Type locality: U.R.S.S.: Abkasie, Ritsa).

  • — Coiffait, 1978: 220 (Quedius; subgenus Sauridus; characters; Abkhasia).

  • Distribution: Georgia.

abnormalis Sharp, 1889: 30 (Quedius; Type locality: Nikko).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • Distribution: Japan.

abruzzensis Drugmand, 1988c: 206 (Quedius; subgenus Raphirus; Type locality: Abruzzes: l'Aquila, versant nord du mont Luco, 900 m).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; cited as “specie dubbia”; Italy).

  • Distribution: Italy.

acatl Smetana, 1975: 322 (Quedius; subgenus Raphirus; Type locality: Mexico: Durango: 10 mi. W. El Salto).

  • Distribution: Mexico.

acco Smetana, 1996: 4 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, 2750 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

acuminatus Fairmaire, 1860, see: maurorufus Gravenhorst, 1806.

acuminatus Hochhuth, 1849: 151 (Quedius; Type locality: Kaukasus).

  • — Hochhuth, 1862: 47 (Quedius; Russia).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; characters; France; Caucasus; Iran).

  • — Mulsant and Rey, 1876: 784 [= 1877: 784] (Quedius; subgenus Raphirus; characters; France).

  • — Porta, 1907: 147 (Quedius; subgenus Raphirus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 153 (Quedius; subgenus Raphirus; characters; Pyrenees; Caucasus; Persia; Germany).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1429 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 55 (Quedius; aedeagal illustration).

  • — Fagel, 1960b: 120, 123 (Quedius; characters).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe Caucasus; Iran).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Korge, 1964: 123 (Quedius; subgenus Raphirus; Turkey).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; Caucasus; Iran; Turkey; Austria; Istria; Bosnia).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; characters; Pyrenees; Greece; Transcaucasus; Lebanon).

  • — Coiffait, 1978d: 165 (Quedius; Turkey).

  • — Tóth, 1984: 125 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Austria).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy ?).

  • Distribution: Europe, Russia, Armenia, Turkey, Lebanon, Iran.

phenicius Coiffait, 1967e: 423 (Quedius; subgenus Raphirus; subspecies of acuminatus; Type locality: Aïn Batara, Liban).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; subspecies of acuminatus; characters).

khnzoriani Coiffait, 1967e: 423 (Quedius; subgenus Raphirus; subspecies of acuminatus; Type locality: Aragatch, Karagel, 3200 m, Arménie russe).

  • — Coiffait, 1978: 265 (Quedius; subgenus Raphirus; subspecies of acuminatus; characters).

adanensis Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; replacement name for gridellii Jarrige).

  • — Coiffait, 1978: 172 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Turkey.

gridellii Jarrige, 1952a: 134 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Adana, Cilicie).

  • — Coiffait, 1967e: 394 (Quedius; synonym of adanensis).

  • — Coiffait, 1978: 172 (Quedius; synonym of adanensis).

adjacens Cameron, 1926b: 368 (Quedius; subgenus Microsaurus; Type locality: Chakrata district, Deoban, 9300 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 287 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1988: 205 (Quedius; subgenus Microsaurus; lectotype designation; characters; Kashmir; Uttar Pradesh; Himachal Pradesh).

  • — Smetana, 1996: 1 (Quedius; subgenus Microsaurus; characters; China).

  • — Smetana, 1999b: 214 (Quedius; subgenus Microsaurus; China).

  • Distribution: India, China.

adnexus Smetana, 1965: 44, 45 (Quedius; subgenus Distichalius; Type locality: Newfoundland: Griguet).

  • — Smetana, 1971: 196 (Quedius; subgenus Raphirus; characters; Newfoundland).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; Canada).

  • Distribution: Canada.

adustus Sharp, 1889: 32 (Quedius; Type locality: Oyama; Nishi; Nara; Chiuzenji; Nishimura; Numata).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Watanabe, 1996a: 13 (Quedius; cave collection; Japan).

  • — Watanabe, 1998: 314 (Quedius; subgenus Microsaurus; Japan).

  • Distribution: Japan, China.

advena Sharp, 1884: 329 (Quedius; Type locality: Mexico, Las Vigas).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Campbell 1975a: 319 (Quedius; subgenus Raphirus; characters; Mexico).

  • Distribution: Mexico.

aeneipennis Heer, 1834 [Note: Nomen nudum], see: alpestris Heer, 1839.

aeneipennis Fairmaire and Germain, 1861, see: Loncovilius.

aeneiventris Broun, 1910: 14 (Quedius; Type locality: Waimarino).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

aenescens Mäklin, 1852: 315 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1965]).

  • — Mäklin, 1853: 190 (Quedius; characters; Edgecombe Island; Kadjak Island; Sitka Island).

  • — Horn, 1878: 164 (Quedius; synonym of brunnipennis).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 406 (Distichalius; cited as brunneipennis; [Note: Misidentified as brunnipennis according to Smetana, 1971: 193]; characters; Alaska; British Columbia).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; cited as aenesceus; catalog).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1965: 45, 58 (Quedius; subgenus Distichalius; lectotype designation; characters).

  • — Smetana, 1971: 193 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Alaska; Oregon; Washington).

  • Distribution: Canada, USA.

agnatus Casey, 1915: 406 (Distichalius; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1964: 38 (Quedius; subgenus Distichalius; Alaska).

  • — Smetana, 1965: 43, 45 (Quedius; subgenus Distichalius; characters; Labrador).

  • — Smetana, 1965a: 13 (Quedius; subgenus Distichalius; Alaska).

  • — Smetana, 1971: 193 (Quedius; synonym of aenescens).

aeneus Fauvel, 1877: 274 (Quedius; Type locality: Australie, Victoria).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: Australia.

aereipennis Bernhauer, 1929d: 111 (Quedius; Type locality: Da-laen-saen bei Nong-po; [Note: See lectotype designation by Smetana, 1996a]).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Smetana, 1996a: 49 (Quedius; subgenus Raphirus; lectotype designation; characters; China).

  • Distribution: China.

aetolicus Kraatz, 1858a: 58 (Quedius; Type locality: ätolische Alpen).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; synonym of cruentus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of cruentus).

  • — Porta, 1907: 115 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Gridelli, 1924d: 54 (Quedius; subgenus Microsaurus; characters; England; Corsica; Algeria; Sardinia; Italy).

  • — Donisthorpe, 1930: 31 (Quedius; characters; Britain).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Roubal, 1941b: 123, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Bordoni, 1974: 45 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 154 (Quedius; subgenus Microsaurus; characters; Europe; Corsica; Sardinia; North Africa; England; France; Portugal).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Europe, Algeria.

subapicalis Joy, 1913a: 76 (Quedius; Type locality: Borckenhurst, New Forest).

  • — Bernhauer and Schubert, 1916: 435 (Quedius; catalog).

  • — Gridelli, 1924d: 55 (Quedius; synonym of aetolicus).

  • — Scheerpeltz, 1933: 1430 (Quedius; synonym of aetolicus).

  • — Bordoni, 1974: 45 (Quedius; synonym of aetolicus).

  • — Pope, 1977: 31 (Quedius; synonym of aetolicus).

  • — Coiffait, 1978: 154 (Quedius; synonym of aetolicus).

afghanicus Coiffait, 1977c: 139 (Quedius; subgenus Sauridus; Type locality: Afghanistan: Paktui, Khat Gaï, 2600 m).

  • — Coiffait, 1978: 232 (Quedius; subgenus Sauridus; characters; Afghanistan).

  • Distribution: Afghanistan.

africanus Peyerimhoff, 1917, see: cinctus Paykull, 1790.

afrofuliginosus Gusarov, 1991a: 9 (Quedius; Type locality: Algeria).

  • Distribution: Algeria.

agathis Broun, 1893: 1407 (Quedius; Type locality: Hunua Range, near Clevedon and at Maketu).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

agnatus Casey, 1915, see: aenescens Mäklin, 1852.

akalita Smetana, 1977, see: Indoquedius.

alatus Coiffait, 1967, see: asturicus Bernhauer, 1918.

albanicus Bernhauer, 1914d: 3 (Quedius; Type locality: Albanien).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Bernhauer, 1923d: 177 (Quedius; subgenus Raphirus; characters; Albania).

  • — Gridelli, 1925: 166 (Quedius; subgenus Raphirus; characters; Albania).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Coiffait, 1978: 283 (Quedius; subgenus Raphirus; characters; Albania).

  • Distribution: Albania.

albanicus Bernhauer, 1926, see: illyricus Wendeler, 1928.

alentejensis Coiffait, 1963a: 391, 412 (Quedius; subgenus Sauridus; [Note: Cited in the subgenus Microsaurus on page 412 but that assignment is probably erroneous]; Type locality: Portugal: Castro Verde).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; characters; Portugal).

  • Distribution: Portugal.

aliiceps Cameron, 1948c: 726 (Quedius; subgenus Raphirus; Type locality: New Zealand: Auckland Island: Carnley Harbour, Musgrave Pen).

  • — Gourlay, 1950: 187 (Quedius; notes).

  • Distribution: New Zealand.

almeriensis Coiffait, 1964 [Note: Nomen nudum], see: cobosi Coiffait, 1964.

alpestris Heer, 1839: 280 (Philonthus; Type locality: Mühlebachalp; [Note: See lectotype designation by Assing, 1999b]. Also cited from Flössalp; Val Emet; Beverserthal; Alpen um Genf).

  • — Kiesenwetter, 1848: 63 (Quedius; synonym of satyrus).

  • — Kraatz, 1857d: 517 (Quedius; characters; Alps).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Fauvel, 1874: 303 [= 1874a: 529] (Quedius; characters; France; Germany; Switzerland; Austria; Italy).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 763 [= 1877: 619] (Quedius; subgenus Raphirus; characters; France).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Alpes; Sudeten; Carpathians).

  • — Porta, 1907: 145 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 142 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 46 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 381 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960b: 120 (Quedius; notes).

  • — Smetana, 1962a: 137 (Quedius; larval characters).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 289 (Quedius; subgenus Raphirus; mountains of central Europe; France; Italy; Slovenia; Switzerland; Germany; Austria; Czechoslovakia; Poland).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius; type species of Microquedius).

  • — Scheerpeltz, 1968a: 92 (Quedius; catalog; Austria).

  • — Tikhomirova, 1973a: 190 (Quedius; checklist; Soviet Union).

  • — Bordoni, 1974a: 205, 206 (Quedius; subgenus Microquedius; characters; Italy).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Alpes; Carpathians; Sudeten).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 137 (Quedius; catalog; Poland).

  • — Tóth, 1984: 121 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Assing, 1999b: 43 (Quedius; lectotype designation: Mühlebachalp; characters; notes).

  • — Štourač, 1998b: 135 (Quedius; subgenus Raphirus; notes; Czech Republic).

  • Distribution: Europe.

aeneipennis Heer, 1834: 75 (Staphylinus; subgenus Philonthus; [Note: Nomen nudum, published without characters]).

  • — Heer, 1839: 280 (Philonthus; synonym of alpestris).

  • — Kraatz, 1857d: 517 (Quedius; synonym of alpestris).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

satyrus Kiesenwetter, 1847: 75 (Quedius; Type locality: Riesengebirge).

  • — Kiesenwetter, 1848: 63 (Quedius; characters; Riesengebirge).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 825 (Philonthus; characters; Austria).

  • — Kraatz, 1857d: 517 (Quedius; synonym of alpestris).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of alpestris).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Scheerpeltz, 1933: 1430 (Quedius; doubtful synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

biseriatus Stierlin, 1875: 473 (Philonthus; Type locality: St. Bernhard).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of alpestris).

  • — Porta, 1907: 145 (Quedius; synonym of alpestris).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of alpestris).

  • — Gridelli, 1925: 142 (Quedius; synonym of alpestris).

  • — Scheerpeltz, 1933: 1430 (Quedius; doubtful synonym of alpestris).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Coiffait, 1978: 198 (Quedius; synonym of alpestris).

spurius Lokay, 1921: 71 (Quedius; variety of alpestris Heer; Type locality: Alpes Styriae, Carinthiae, Carnioliae, Montes Bohemiae, Silesiae).

  • — Gridelli, 1925: 142 (Quedius; subgenus Raphirus; variety of alpestris; characters).

  • — Scheerpeltz, 1933: 1430 (Quedius; variety of alpestris).

  • — Scheerpeltz, 1953a: 133 (Quedius; [Note: Misidentification: Smetana, 1958: 382, cited alpestris Scheerpeltz as a synonym of spurius; Scheerpeltz attributed alpestris to Heer]; characters).

  • — Smetana, 1958: 382 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Europe).

  • — Tóth, 1984: 122 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria).

  • — Nowosad, 1990: 147 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Assing, 1999b: 43 (Quedius; synonym of alpestris).

marcuzzii Scheerpeltz, 1953a: 130 (Quedius; Type locality: Massiccio della Marmolada, 2200 m).

  • — Smetana, 1958: 381 (Quedius; synonym of alpestris).

  • — Smetana, 1962b: 140 (Quedius; synonym of alpestris).

  • — Coiffait, 1963a: 382 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Coiffait, 1978: 198 (Quedius; subgenus Microquedius; characters; Dolomites).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; cited as “specie dubbia”; Italy).

  • — Assing, 1999b: 43 (Quedius; synonym of alpestris).

alpinus Fiori, 1894, see: plagiatus Mannerheim, 1843.

alpinus Helliesen, 1892: 74 (Quedius; Type locality: Mjosvand in Telemarken).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1925: 168 (Quedius; characters).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • Distribution: Norway.

altaicus Coiffait, 1967, see: centrasiaticus Coiffait, 1969.

altaicus Korge, 1962: 152 (Quedius; subgenus Quedius; Type locality: Zentral-Altai).

  • — Coiffait, 1978: 194 (Quedius; subgenus Quedius; characters; central Asia).

  • Distribution: “Altai region”.

alticola Smetana, 1971: 172 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Rainier Nat. Pk., Nisqually River, 4000 ft. Also cited from California).

  • — Smetana, 1976c: 175 (Quedius; subgenus Distichalius; Oregon; Washington).

  • — Smetana, 1981b: 640 (Quedius; collecting notes; Oregon).

  • Distribution: USA.

alutaceus Hatch, 1957, see: densiventris Casey, 1915.

amabilis Smetana, 1971: 205 (Quedius; subgenus Raphirus; Type locality: California: nr. Strawberry, El Dorado Co.).

  • — Smetana, 1990d: 100 (Quedius; California).

  • Distribution: USA.

ambiguus Broun, 1894: 423 (Quedius; Type locality: Tarukenga, about 8 miles from Rotorua).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: New Zealand.

amethystinus Roubal, 1948 [Note: Nomen nudum], see: scintillans Gravenhorst, 1806.

amicorum Smetana, 1997d: 470 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4050 m, 27°49 N, 99°34 E).

  • Distribution: China.

amplicollis Scriba 1868, see: crassus Fairmaire, 1860.

amplissimus Bernhauer, 1912k: 262 (Quedius; Type locality: Umgebung von Sebastopol, Krim).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 46 (Quedius; subgenus Microsaurus; characters; Crimea).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Jarrige, 1971: 497 (Quedius; Iran).

  • Distribution: Ukraine, Iran.

analis Fabricius, 1787, see: scitus Gravenhorst, 1806.

analis Stephens, 1835, see: cruentus Olivier, 1795.

analis W. J. MacLeay, 1873, see: manducus Herman, 2001.

analoka Smetana, 1975, see: stevensi Cameron, 1932.

anatolicus Korge, 1964, see: suturalis Kiesenwetter, 1845.

anceps Fairmaire and Laboulbène, 1856: 543 (Quedius; Type locality: H. -Pyr., Aragnouet).

  • — Fauvel, 1874: 294 [= 1874a: 520] (Quedius; synonym of umbrinus).

  • — Mulsant and Rey, 1876: 724 [= 1877: 580] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; Pyrenees; France).

  • — Fauvel, 1900d: 233 (Quedius; notes).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 111 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • — Scheerpeltz, 1933: 1430 (Quedius; catalog).

  • — Jeannel and Jarrige, 1949: 379 (Quedius; subgenus Raphirus; cave species; France).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 230 (Quedius; subgenus Sauridus; characters; Pyrenees).

  • — Tronquet, 1998a: 114 (Quedius; collected from marmot burrow; France).

  • Distribution: Portugal, Spain, France, Italy, Switzerland.

robustus Scriba, 1870: 81 (Quedius; Type locality: Prope Santas Albas, prov. Leon).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; characters; France; Switzerland; Italy; Spain; Portugal).

  • — Mulsant and Rey, 1876: 710 [= 1877: 566] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; western Alps; Spain; Portugal).

  • — Porta, 1907: 136 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; catalog).

  • — Gridelli, 1924d: 111 (Quedius; synonym of anceps).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1431 (Quedius; synonym of anceps).

parviceps Fauvel, 1871b: 359 (Quedius; [Note: Nomen nudum (Articles 11.6, 12.1); species not described but has been listed as a synonym by other authors]; cited from Monte Rosa, Furca, Faulhorn, Alpen von 4000 bis 8000′).

  • — Fauvel, 1874: 292 [= 1874a: 518] (Quedius; synonym of robustus).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of robustus).

  • — Bernhauer and Schubert, 1916: 433 (Quedius; synonym of robustus).

  • — Porta, 1907: 136 (Quedius; synonym of robustus).

  • — Scheerpeltz, 1933: 1430 (Quedius; synonym of anceps).

  • — Coiffait, 1978: 230 (Quedius; synonym of anceps).

andersoni Blackburn, 1888: 6 (Quedius; Type locality: Port Lincoln district).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

andreinii Gridelli, 1924d: 19, 30 (Quedius; subgenus Microsaurus; Type locality: Santo Stefano d'Aveto, Apennino Ligure).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Coiffait, 1978: 173 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy.

angaricus Coiffait, 1975: 34 (Quedius; subgenus Sauridus; Type locality: Kistvianka, région sud-ouest du Lac Baïkal).

  • — Coiffait, 1978: 259 (Quedius; subgenus Sauridus; characters; central Asia).

  • Distribution: Russia.

angnimai Smetana, 1988: 223 (Quedius; subgenus Microsaurus; Type locality: Nepal: Prov. Bagmati, below Thare Pati, 3300 m).

  • — Smetana, 1992b: 4 (Quedius; subgenus Microsaurus; Nepal).

  • Distribution: Nepal.

angularis Cameron, 1948: 41 (Quedius; subgenus Sauridus; Type locality: West of Fort Archambault, Sara-Dai).

  • Distribution: Chad.

angulicollis Fauvel, 1891a: 60 (Quedius; Type locality: Drome; Alpes-Maritimes, Venanson; Apennins).

  • — Ganglbauer, 1895: 408 (Quedius; subgenus Sauridus; characters; Maritime Alps; Apennines).

  • — Porta, 1907: 141 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1922b: 128, 131 (Quedius; subgenus Sauridus; characters; notes; Italy).

  • — Scheerpeltz, 1933: 1433 (Quedius; synonym of bicolor).

  • — Coiffait, 1978: 241 (Quedius; synonym of bicolor).

  • — Herman, 2001: 49 (Quedius; valid name for species).

bicolor Mulsant and Rey, 1876: 738 [= 1877: 594] (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Grande Chartreuse).

  • — Ganglbauer, 1895: 407 (Quedius; synonym of humeralis).

  • — Porta, 1907: 140 (Quedius; synonym of humeralis).

  • — Bernhauer and Schubert, 1916: 424 (Quedius; synonym of humeralis).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1963b: 425 (Quedius; subgenus Sauridus).

  • — Coiffait, 1978: 241 (Quedius; subgenus Sauridus; characters; Alps; Apennines; Massif Central).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Herman, 2001: 49 (Quedius; synonym of angulicollis).

  • Distribution: France, Italy.

angustifrons Wollaston, 1864: 563 (Quedius; Type locality: Gran Canaria; [Note: See lectotype designation by Israelson, 1985]. Originally cited from Grand Canary: Teror; Gomera: Hermigua).

  • — Wollaston, 1865: 486 (Quedius; habitat; Canary Islands).

  • — Fauvel, 1902b: 113 (Quedius; Canaries).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Gridelli, 1924d: 38 (Quedius; subgenus Microsaurus; characters; Canaries).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Israelson, 1985: 152 (Quedius; subgenus Microsaurus; lectotype designation: Gran Canaria; characters; Canary Islands).

  • Distribution: Canary Islands.

angustus Sharp, 1884: 338 (Quedionuchus; Type locality: Guatemala, Totonicapam, 8500 to 10,500 feet; [Note: See lectotype designation by Smetana, 1976a]).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Smetana, 1976a: 232 (Quedius; subgenus Quedionuchus; lectotype designation; characters; Guatemala).

  • Distribution: Guatemala.

annectens Sharp, 1889: 32 (Quedius; Type locality: Miyanoshita; [Note: See lectotype designation by Smetana, 1998a]).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1998a: 119 (Quedius; subgenus Distichalius; lectotype designation; characters; Japan).

  • Distribution: Japan.

anomalus Cameron, 1926b: 370 (Quedius; Type locality: Mussorie district: Dhobi Ghat; [Note: See lectotype designation by Smetana, 1988]).

  • — Cameron, 1932: 295 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 274 (Quedius; subgenus Raphirus; lectotype designation; characters; India).

  • Distribution: India.

antarcticus Bernhauer and Schubert, 1916: 418 (Quedius; replacement name for scutellaris Broun).

  • Distribution: New Zealand.

scutellaris Broun, 1894: 422 (Quedius; [preoccupied]; Type locality: Ashburton).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of antarcticus).

antennalis Cameron, 1932: 285 (Quedius; subgenus Microsaurus; Type locality: Darjeeling: Gopaldhara, alt. 4720 feet).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 201 (Quedius; subgenus Microsaurus; characters; India).

  • Distribution: India.

anthracinus Solsky, 1868a: 125 (Quedius; Type locality: Mexique).

  • — Sharp, 1884: 332 (Quedius; notes; Mexico).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; catalog).

  • Distribution: Mexico.

antipodum Sharp, 1886a: 378 (Quedius; Type locality: New Zealand: Dunedin; Greymouth).

  • — Broun, 1893: 1028 (Quedius; characters; New Zealand).

  • — Broun, 1911: 93 (Quedius; Chatham Islands).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Emberson, 1998: 36 (Quedius; collecting notes; Chatham Islands).

  • Distribution: New Zealand, Chatham Islands.

antoni Smetana, 1995f: 233 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, 3300–3350 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • Distribution: China.

apicalis Cameron, 1941, see: philippinus Cameron, 1941.

apicicornis Eppelsheim, 1895b: 391 (Quedius; subgenus Microsaurus; Type locality: Sikkim im Himalaya).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 286 (Quedius; subgenus Microsaurus; characters; Sikkim: Darjeeling district, Tiger Hill, alt. 8000 feet).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana 1975a: 325 (Quedius; subgenus Microsaurus; characters; Nepal; Sikkim; Bhutan).

  • — Smetana, 1977a: 246 (Quedius; Bhutan).

  • — Coiffait, 1982: 32 (Quedius; characters; notes; Nepal).

  • — Smetana, 1988: 192 (Quedius; subgenus Microsaurus; characters; Bhutan; West Bengal; Sikkim; Nepal).

  • — Smetana, 1992b: 2 (Quedius; subgenus Microsaurus; Nepal; India).

  • Distribution: India, Nepal, Bhutan.

flavoterminatus Cameron, 1932: 290 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Rangirum, alt. 6000 feet).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

cyanopterus Scheerpeltz, 1976a: 150 (Quedius; subgenus Microsaurus; Type locality: Nepal: Umgebung von Kathmandu, 1400 m).

  • — Coiffait, 1982: 32 (Quedius; synonym of apicicornis).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

sherpai Coiffait, 1982: 77 (Quedius; subgenus Microsaurus; Type locality: Nepal: Trisuli Tal, Gosainkund, 3200 m).

  • — Smetana, 1988: 192 (Quedius; synonym of apicicornis).

apiciflavus Lea, 1925: 248 (Quedius; Type locality: Queensland: Mount Tambourine).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

apicipennis Hellén, 1934 [Note: Nomen nudum], see: tenellus Gravenhorst, 1806.

aplodontiae Smetana, 1971, see: spelaeus Horn, 1871.

arabs Lokay, 1921 [Note: Nomen nudum], see: cinctus Paykull, 1790.

arcticus Munster, 1921, see: sublimbatus Mäklin, 1853.

arctifrons Broun, 1880: 112 (Philonthus; Type locality: Parua).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • Distribution: New Zealand.

arestor Tottenham, 1948, see: boops Gravenhorst, 1802.

aridulus Jansson, 1939, see: persimilis Mulsant and Rey, 1876.

armipes Sharp, 1889: 34 (Quedionuchus; Type locality: Shimidzu; Yuyama; Nikko).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1995a: 80 (Quedius; subgenus Quedionuchus; characters; Russia).

  • Distribution: Japan, Russia.

arrogans Sharp, 1884: 333 (Quedius; Type locality: Panama, Volcan de Chiriqui, at elevations between 2000 and 6000 feet).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 173, 174 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • Distribution: Panama.

arrowi Bernhauer, 1941b: 32 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Pudding Hill).

  • Distribution: New Zealand.

arvernicus Mulsant and Rey, 1876, see: mesomelinus Marsham, 1802.

arvernicus Coiffait, 1982, see: obscuripennis Bernhauer, 1901.

arvernus Herman, 2001, see: obscuripennis Bernhauer, 1901.

asiaticus Bernhauer, 1918a: 92 (Quedius; subgenus Ediquus; Type locality: Ost-Buchara, Turkestan).

  • — Gridelli, 1924d: 57 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Coiffait, 1978: 183 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • Distribution: Uzbekistan.

aspromontanus Bernhauer, 1908b: 34 (Quedius; Type locality: Kalabrien, Aspromonte).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 133 (Quedius; subgenus Sauridus; characters; notes; Italy).

  • — Gridelli, 1925: 120 (Quedius; subgenus Raphirus; characters; Italy).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 34 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Fagel, 1968c: 7 (Quedius; subgenus Sauridus; characters; notes; Italy; Corsica; Greece).

  • — Coiffait, 1978: 215 (Quedius; subgenus Sauridus; characters; Italy; Corsica; Greece).

  • — Zanetti, 1992b: 130, 131 (Quedius; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy, Greece.

assamensis Cameron, 1932: 293 (Quedius; subgenus Raphirus; Type locality: Assam: Naga Hills, alt. 4000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Smetana, 1988: 284 (Quedius; subgenus Raphirus; lectotype designation; characters; Nepal; India).

  • — Smetana, 1996c: 225 (Quedius; subgenus Raphirus; China).

  • Distribution: India, Nepal, China.

assecla Mulsant and Rey, 1876, see: ochripennis Ménétriés, 1832.

assimilis Nordmann, 1837, see: fulgidus Fabricius, 1793.

asturicus Bernhauer, 1918a: 96 (Quedius; subgenus Raphirus; Type locality: Asturien: Caboalles, Cancas).

  • — Gridelli, 1925: 165 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1431 (Quedius; catalog).

  • — Fagel, 1960b: 118 (Quedius; characters; Carinthia; Bosnia; Italy).

  • — Smetana, 1962b: 152 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; characters; Pyrenees; Bulgaria).

  • — Drugmand, 1988c: 209 (Quedius; characters).

  • — Tronquet, 1999a: 214 (Quedius; subgenus Raphirus; notes; France).

  • Distribution: Europe.

brevipennis Fairmaire, 1859: 41 (Quedius; [preoccupied]; Type locality: Pyrénées).

  • — Fauvel, 1874: 309 [= 1874a: 535] (Quedius; race of boops; characters; France; Italy; Portugal).

  • — Mulsant and Rey, 1876: 797 [= 1877: 653] (Quedius; subgenus Raphirus; characters; France).

  • — Sharp, 1889: 33 (Quedius; notes; Japan).

  • — Ganglbauer, 1895: 412 (Quedius; variety of boops).

  • — Porta, 1907: 150 (Quedius; variety of boops).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; variety of boops).

  • — Gridelli, 1925: 165 (Quedius; variety of boops).

  • — Scheerpeltz, 1933: 1434 (Quedius; variety of boops).

  • — Fagel, 1960a: 118 (Quedius; synonym of asturicus).

  • — Smetana, 1962b: 153 (Quedius; synonym of asturicus).

  • — Coiffait, 1978: 281 (Quedius; synonym of asturicus).

alatus Coiffait, 1967e: 424 (Quedius; subgenus Raphirus; subspecies of asturicus; Type locality: Vratza, Bulgarie).

  • — Coiffait, 1970e: 63 (Quedius; subspecies of asturicus; Cyprus).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of asturicus; characters).

  • — Tóth, 1984: 129 (Quedius; subspecies of asturicus; characters).

atchala Smetana, 1975a: 341 (Quedius; subgenus Raphirus; Type locality: Central Nepal: Gosaikunde Lakes area 3500–4500 m).

  • — Smetana, 1988: 270 (Quedius; subgenus Raphirus; characters; Nepal).

  • Distribution: Nepal.

ater J. Sahlberg, 1880, see: plagiatus Mannerheim, 1843.

ater Ziegler, 1845, see: capucinus Gravenhorst, 1806.

atl Smetana, 1975: 315 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 37 mi. S. Valle Nacional, 8500 ft.).

  • — Smetana, 1976a: 244 (Quedius; subgenus Quedionuchus; characters; Mexico).

  • Distribution: Mexico.

atricapillus Reitter, 1900: 273 (Quedius; Type locality: Ober-Syrien: Akbes).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 56 (Quedius; subgenus Microsaurus; characters; Syria).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Jarrige, 1952a: 134 (Quedius; subgenus Microsaurus; characters).

  • — Coiffait, 1978: 170 (Quedius; subgenus Microsaurus; characters; Turkey).

  • Distribution: Syria.

atriceps Stephens, 1832, see: scitus Gravenhorst, 1806.

atricillus Gravenhorst, 1806, see: scitus Gravenhorst, 1806.

atripennis Stephens, 1835, see: mesomelinus Marsham, 1802.

atritarsis Roubal, 1949 [Note: Nomen nudum], see: unicolor Kiesenwetter, 1847.

attenuatus Gravenhorst, 1802, see: limbatus Heer, 1839.

attenuatus Gyllenhal, 1810 [Note: Nomen nudum], see: nitipennis Stephens, 1833.

atticus Coiffait, 1967, see: coxalis Kraatz, 1858.

aucklandicus Cameron, 1947d: 707 (Quedius; subgenus Raphirus; Type locality: New Zealand: N. Auckland: Tangihua Range, 1800–2000 feet).

  • Distribution: New Zealand.

aureipilis Cameron, 1932: 294 (Quedius; subgenus Raphirus; Type locality: Assam: Naga Hills, Laimatak; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Smetana, 1988: 288 (Quedius; subgenus Raphirus; lectotype designation; characters; Nagaland).

  • Distribution: India.

aureiventris Bernhauer, 1915b: 56 (Quedius; subgenus Raphirus; Type locality: Zentral-Himalaya: Kumaon, Binsar, 7900 Fuss).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Cameron, 1932: 294 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Coiffait, 1982a: 234 (Quedius; Pakistan).

  • — Smetana, 1988: 250 (Quedius; subgenus Raphirus; characters; Bhutan; Kashmir; Himachal Pradesh; Uttar Pradesh; West Bengal; Nepal; Pakistan).

  • — Smetana, 1992b: 5 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: Pakistan, India, Nepal, Bhutan.

decipiens Cameron, 1944: 14 (Quedius; subgenus Raphirus; Type locality: Darjeeling: Ghum district: Mangpu).

  • — Smetana 1975a: 333 (Quedius; subgenus Raphirus; characters; Nepal; India; Bhutan).

  • — Smetana, 1977a: 246 (Quedius; Bhutan).

  • — Coiffait, 1982: 33 (Quedius; characters; Nepal).

  • — Coiffait, 1982a: 234 (Quedius; Nepal).

  • — Smetana, 1988: 250 (Quedius; synonym of aureiventris).

auricomus Kiesenwetter, 1850: 220 (Quedius; Type locality: Pyrenaei orientales).

  • — Kiesenwetter, 1851a: 420 (Quedius; characters; Pyrenees).

  • — Fairmaire and Laboulbène, 1856: 540 (Quedius; characters; France).

  • — Kraatz, 1857d: 501 (Quedius; characters).

  • — Fauvel, 1874: 303 [= 1874a: 529] (Quedius; characters; France; Britain).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Mulsant and Rey, 1876: 765 [= 1877: 621] (Quedius; subgenus Raphirus; characters; France).

  • — Fowler, 1888: 242 (Quedius; subgenus Raphirus; characters; Britain).

  • — Ganglbauer, 1895: 410 (Quedius; subgenus Raphirus; characters; Germany; France; England).

  • — Porta, 1907: 144 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Johansen, 1914: 369 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1925: 144 (Quedius; subgenus Raphirus; characters; Pyrenees; Germany; Ireland; England; France; Alps; Italy; Belgium; Romania).

  • — Portevin, 1929: 344 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1432 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 48 (Quedius; aedeagal illustration).

  • — Tottenham, 1948a: 245 (Quedius; subgenus Arphirus; characters; Britain).

  • — V. Hansen, 1952: 151 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 383 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Europe).

  • — Coiffait, 1963a: 383 (Quedius; subgenus Sauridus; characters).

  • — Palm, 1963b: 151 (Quedius; subgenus Raphirus; characters).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 290 (Quedius; subgenus Raphirus; Ireland; Britain; Denmark; Holland; Belgium; France; Switzerland; Germany; Austria).

  • — Coiffait, 1967e: 419 (Quedius; subgenus Microquedius).

  • — Bordoni, 1974a: 205, 207 (Quedius; subgenus Microquedius; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 199 (Quedius; subgenus Microquedius; characters; England; Ireland; Belgium; France; Germany; Italy; Romania; Pyrenees).

  • — Tóth, 1984: 122 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic ?).

  • Distribution: Europe.

tomentosomaculatus Cornelius, 1863: 130 (Quedius; Type locality: Elberfeld).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of auricomus).

  • — Fowler, 1888: 242 (Quedius; subgenus Raphirus; synonym of auricomus).

  • — Ganglbauer, 1895: 410 (Quedius; synonym of auricomus).

  • — Porta, 1907: 144 (Quedius; synonym of auricomus).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; synonym of auricomus).

  • — Smetana, 1958: 384 (Quedius; synonym of auricomus).

  • — Coiffait, 1978: 199 (Quedius; synonym of auricomus).

aurofasciatus Bernhauer, 1917h: 91 (Quedius; subgenus Euryquedius; Type locality: Venezuela).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • Distribution: Venezuela.

aurorus Herman, 2001: 49 (Quedius; replacement name for laticollis Sharp).

  • Distribution: Japan.

laticollis Sharp, 1889: 31 (Quedius; [preoccupied]; Type locality: Sapporo).

  • — Bernhauer and Schubert, 1916: 426 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Herman, 2001: 49 (Quedius; synonym of aurorus).

babai Uéno and Watanabe, 1966: 331 (Quedius; subgenus Microsaurus; Type locality: Japan: central Honshu: southwestern Niigata Prefecture: Sekimi-toge, eastern slope of Mt. Myoko).

  • Distribution: Japan.

badius Broun, 1923: 678 (Quedius; Type locality: Gordon's Pyramid, Mount Arthur, 4500 ft.).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • Distribution: New Zealand.

badius Scholz, 1931 [Note: Nomen nudum], see: fuliginosus Gravenhorst, 1802.

bakeri Hatch, 1957: 218 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Baker, Chain L. Also cited from Washington).

  • — Smetana, 1971: 173 (Quedius; subgenus Distichalius; characters; Washington).

  • — Smetana, 1976c: 175 (Quedius; subgenus Distichalius; Oregon; Washington).

  • Distribution: USA.

balianii Gridelli, 1924, see: nigrocaeruleus Fauvel, 1876.

balcanicus Bernhauer, 1908e: 324 (Quedius; Type locality: Duzi in der Herzegowina).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1922b: 128, 133 (Quedius; subgenus Sauridus; characters; Italy; Dalmatia; Hercegovina).

  • — Gridelli, 1925: 136 (Quedius; subgenus Raphirus; characters; Albania).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 41 (Quedius; aedeagal illustration).

  • — Coiffait, 1963a: 386 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 233 (Quedius; subgenus Sauridus; characters; Albania; Dalmatia; Istria).

  • — Tóth, 1984: 137 (Quedius; characters).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy, Bosnia Hercegovina, Croatia, Albania.

baldiensis Blackburn, 1891: 69 (Quedius; variety of cuprinus; Type locality: Mountains of Victoria, about 5,000 feet).

  • — Blackburn, 1903: 94 (Quedius; distinct species, not variety of cuprinus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of cuprinus).

  • — Lea, 1925: 242 (Quedius; cited as valid species; characters; notes; Australia).

  • Distribution: Australia.

balthasari Mařan, 1935, see: levicollis Brullé, 1832.

balticus Korge, 1960: 52 (Quedius; Type locality: Südspitze der Insel Hiddensee bei Rügen).

  • — Last, 1952a (Quedius; cited as molochinus; [Note: Misidentified according to Korge, 1960a: 67]; discussion and characters to separate from pallipes).

  • — Korge, 1960a: 67 (Quedius; characters; notes; England).

  • — Coiffait, 1961a: 50 (Quedius; characters).

  • — Smetana, 1962b: 135 (Quedius; subgenus Quedius; characters).

  • — V. Hansen, 1962a: 319 (Quedius; characters; Denmark).

  • — Last, 1963: 43 (Quedius; characters; notes; Britain).

  • — Palm, 1963: 117 (Quedius; characters; Sweden).

  • — Palm, 1963b: 160 (Quedius; characters; Sweden; Denmark).

  • — Lohse, 1964: 212 (Quedius; subgenus Quedius; characters; central Europe).

  • — Horion, 1965: 277 (Quedius; subgenus Quedius; Sweden; Denmark; Britain; France; Germany; Austria).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; characters; Germany; Denmark; Britain; border of salt lakes in central Europe).

  • — Tóth, 1984: 119 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Austria; Benelux; Denmark; Sweden).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Hodge and Jones, 1995: 41 (Quedius; characters; Britain).

  • Distribution: Europe.

funebris Coiffait, 1971a: 175 (Quedius; subgenus Quedius; variety of balticus; [Note: Nomen nudum (Article 15.2)]; cited from France: La Ferté-Bernard, Sarthe).

  • — Coiffait, 1978: 192 (Quedius; subgenus Quedius; variety of balticus; characters).

  • — Tóth, 1984: 119 (Quedius; variety of balticus; characters).

bardus Melsheimer, 1844, see: capucinus Gravenhorst, 1806.

basiventris Sharp, 1884: 330 (Quedius; Type locality: Guatemala, Volcan de Atitlan about 3000 feet, Mirandilla 1700 feet, Las Mercedes, Pantaleon, San Isidro; Panama, Volcan de Chiriqui between 2500 and 4000 feet).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 174, 175 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala; Panama).

  • — Frank, 1982: 45 (Quedius; list of parasites).

  • Distribution: Guatemala, Panama.

becvari Smetana, 1996f: 119 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan, Daju, 50 km N of Lijiang, 27.21N 100.19E).

  • — Smetana, 1997b: 130 (Quedius; notes; China).

  • — Smetana, 1997d: 455 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

beesoni Cameron, 1932: 285 (Quedius; subgenus Microsaurus; Type locality: Deoban, Chakrata, U.P.; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Chakrata district, alt. 7000–9000 feet. Darjeeling Dist.: Ghum, Rongdong Valley).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; subgenus Microsaurus; lectotype designation: Deoban, Chakrata, U.P.; characters; Uttar Pradesh; West Bengal; Nepal).

  • — Smetana, 1995: 31 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • — Smetana, 1996: 2 (Quedius; subgenus Microsaurus; notes; China).

  • — Smetana, 1999b: 214 (Quedius; subgenus Microsaurus; China).

  • Distribution: India, Nepal, China, Taiwan.

mimeticus Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1450 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

notabilis Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1452 (Quedius; catalog).

  • — Scheerpeltz, 1965: 111 (Quedius; subgenus Microsaurus; Burma).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

peraffinis Cameron, 1932: 286 (Quedius; subgenus Microsaurus; Type locality: Darjeeling Dist.: Ghum, Rongdong Valley; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1456 (Quedius; catalog).

  • — Smetana, 1988: 196 (Quedius; lectotype designation; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

sungkangensis Hayashi, 1992: 11 (Quedius; subgenus Microsaurus; Type locality: Sungkang, Nantou Hsien).

  • — Hayashi, 1992a: 113 (Quedius; synonym of beesoni).

  • — Smetana, 1995: 32 (Quedius; synonym of beesoni).

bohemorum Smetana, 1997d: 461 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4050 m, 27°49 N, 99°34 E).

  • Distribution: China.

belgravensis Oke, 1928: 1 (Quedius; Type locality: Australia: Victoria: Belgrave).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

bellus Lea, 1925: 243 (Quedius; Type locality: Victoria: Belgrave, Ferntree Gully, Beaconsfield).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

bernhaueri Rambousek, 1915a: 27, 29 (Quedius; subgenus Tenebrobius; Type locality: Peristerii-Gebirge bei Monastir in Mazedonien).

  • — Rambousek, 1916: 130 (Quedius; subgenus Tenebrobius; cited as a new species; Macedonia).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 33 (Quedius; subgenus Microsaurus; characters; Macedonia).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Coiffait, 1978: 181 (Quedius; subgenus Microsaurus; characters; Macedonia).

  • Distribution: Macedonia.

bernhauerianus Korge, 1971a: 56 (Quedius; replacement name for reitteri Bernhauer).

  • Distribution: China.

reitteri Bernhauer, 1933a: 51 (Quedius; subgenus Sauridus; [preoccupied]; Type locality: Tatsienlu-Kiulung).

  • — Korge, 1971a: 56 (Quedius; synonym of bernhauerianus).

berytensis Coiffait, 1954a: 160 (Quedius; subgenus Microsaurus; Type locality: Liban: Beyrouth).

  • — Coiffait, 1955e: 427 (Quedius; cave collection; Lebanon).

  • — Coiffait, 1978: 159 (Quedius; subgenus Microsaurus; characters; Lebanon).

  • Distribution: Lebanon.

beryticus Coiffait, 1977 [Note: Nomen nudum], see: quadriceps Fauvel, 1900.

bhari Smetana, 1988: 256 (Quedius; subgenus Raphirus; Type locality: Nepal: Khandbari District, For. above Ahale, 2400 m).

  • Distribution: Nepal.

bicolor Redtenbacher, 1849, see: fulgidus Fabricius, 1793.

bicolor Mulsant and Rey, 1876, see: angulicollis Fauvel, 1891.

bih Smetana, 1995: 101 (Quedius; subgenus Raphirus; Type locality: Taiwan: Chiai Hsien: Yushan N. P., Mun-Li Cliff, 2700 m).

  • — Smetana, 1998b: 110 (Quedius; subgenus Raphirus; characters).

  • Distribution: Taiwan.

bipunctatus Eppelsheim, 1895, see: Indoquedius.

bipustulatus Fiori, 1894, see: scintillans Gravenhorst, 1806.

birmanus Cameron, 1932: 284 (Quedius; subgenus Microsaurus; Type locality: Burma: Ruby Mines).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — Scheerpeltz, 1965: 111 (Quedius; subgenus Microsaurus; Burma).

  • — Smetana, 1988: 202 (Quedius; subgenus Microsaurus; characters; Burma).

  • Distribution: Myanmar.

biseriatus Stierlin, 1875, see: alpestris Heer, 1839.

bithynicus Fagel, 1968, see: gemellus Eppelsheim, 1889.

bito Smetana, 1996: 7 (Quedius; subgenus Microsaurus; Type locality: China: Gansu, Mts. 25 km E. Xiahe, 2805–2925 m).

  • — Smetana, 1997d: 452 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; collecting notes; China).

  • Distribution: China.

blackburni Bernhauer and Schubert, 1916, see: tepperi Blackburn, 1888.

bohemicus Roubal, 1941, see: puncticollis Thomson, 1867.

bolivianus Scheerpeltz, 1955a: 175, 181 (Quedius; subgenus Cyrtoquedius; Type locality: Bolivien: Yungas de la Paz).

  • Distribution: Bolivia.

boluensis Korge, 1971a: 49 (Quedius; subgenus Sauridus; Type locality: Abant-Gebirge bei Bolu).

  • — Coiffait, 1978: 209 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

bonapartei Hatch, 1957, see: densiventris Casey, 1915.

bonvouloirii Brisout, 1863: 31 (Quedius; Type locality: Pyrénées-Orientales, Hautes-Pyrénées).

  • — Fauvel, 1869: 486 (Quedius; synonym of attenuatus).

  • — Fauvel, 1869a: 151 (Quedius; synonym of attenuatus).

  • — Fauvel, 1874: 307 [= 1874a: 533] (Quedius; synonym of acuminatus).

  • — Porta, 1907: 147 (Quedius; synonym of acuminatus).

  • — Bernhauer and Schubert, 1916: 418 (Quedius; synonym of acuminatus).

  • — Scheerpeltz, 1933: 1429 (Quedius; synonym of acuminatus).

  • — Fagel, 1960b: 120 (Quedius; characters).

  • — Coiffait, 1978: 270 (Quedius; subgenus Raphirus; characters; Pyrenees).

  • Distribution: France, Spain.

boopoides Munster, 1923: 201, 203 (Quedius; subgenus Raphirus; variety of fulvicollis; [Note: Name cited as subspecies on page 201 and variety on page 203]; Type locality: Lysaker ved Kristiania; Fokstuen paa Dovre; Sebuhaugen saeter i Torpen; Kongsberg; Fyresdal; Grimstad; Mandal; Bø i Vesteraalen; Lødingen; Tromsø; Matjok i Tana; Vardø; Neiden samt fl. st. ved Pasvikelven i Syd Varanger).

  • — Tottenham, 1948a: 251 (Quedius; subgenus Arphirus; valid species; characters; Britain).

  • — V. Hansen, 1952: 156 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Smetana, 1958: 395 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960b: 116 (Quedius; characters; Norway; Germany; Denmark; England).

  • — Smetana, 1962b: 153 (Quedius; subgenus Raphirus; characters).

  • — Palm, 1963b: 153 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 219 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 80 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Horion, 1965: 305 (Quedius; subgenus Raphirus; Britain; Norway; Sweden; Finland; Russia; Denmark; Holland; Belgium; France; Czech Republic; Poland; Germany).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 280 (Quedius; subgenus Raphirus; characters; Europe; Britain; France).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Tóth, 1984: 130 (Quedius; characters).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 149 (Quedius; collected from mole nest; notes; Poland).

  • — Whitehead, 1991 (Quedius; collecting notes; Britain).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

boops Gravenhorst, 1802: 21 (Staphylinus; Type locality: Not cited, but presumably Brunsvigae).

  • — Latreille, 1804: 311 (Staphylinus; characters; Germany; France).

  • — Gravenhorst, 1806: 71 (Staphylinus; characters; France).

  • — Gyllenhal, 1810: 312 (Staphylinus; characters; Sweden).

  • — Zetterstedt, 1828: 72 (Staphylinus; characters; Lapland).

  • — C. Sahlberg, 1830: 318 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 27 (Staphylinus; Finland).

  • — Mannerheim, 1831: 441 (Staphylinus; Finland).

  • — Stephens, 1833: 242 (Raphirus; characters; England; Scotland).

  • — Lacordaire, 1835: 385 (Emus; characters; France).

  • — Nordmann, 1837: 78 (Philonthus; Europe).

  • — Zetterstedt, 1838: 62 (Staphylinus; characters; habitat; Lapland).

  • — Erichson, 1839a: 494 (Quedius; characters; Germany).

  • — Heer, 1839: 280 (Philonthus; characters; Switzerland).

  • — Stephens, 1839: 393 (Quedius; characters; England; Scotland).

  • — Heer, 1839: 279 (Philonthus; variety b of picipennis described by Heer was cited as a synonym of boops by Scheerpeltz, 1933: 1434; characters; Switzerland).

  • — Erichson, 1840: 548 (Quedius; characters; Germany; Sweden).

  • — Kiesenwetter, 1848: 61 (Quedius; characters; Germany; Sweden).

  • — Gravenhorst, 1847: 229 (Quedius; characters; notes).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 421 (Quedius; Pyrenees).

  • — Hardy, 1851: 34 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 538 (Quedius; characters; France).

  • — Staudinger, 1857: 286 (Quedius; notes; Iceland).

  • — Kraatz, 1857d: 516 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 202 (Quedius; characters; Austria).

  • — Thomson, 1860: 179 (Raphirus; characters; Scandinavia).

  • — Fauvel, 1874: 308 [= 1874a: 534] (Quedius; characters; France; Iceland; Europe; Cyprus).

  • — Redtenbacher, 1874: 199 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 264 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 794 [= 1877: 650] (Quedius; subgenus Raphirus; characters; France).

  • — J. Sahlberg, 1876: 30 (Raphirus; Finland).

  • — Fowler, 1888: 245 (Quedius; subgenus Raphirus; characters; Britain).

  • — Eppelsheim, 1892: 332 (Quedius; Turkestan).

  • — Ganglbauer, 1895: 412 (Quedius; subgenus Raphirus; characters; Europe).

  • — Fauvel, 1902b: 117 (Quedius; Algeria; Tunisia).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Sainte-Claire Deville, 1906: 103 (Quedius; Corsica).

  • — Porta, 1907: 150 (Quedius; subgenus Raphirus; characters; Italy).

  • — Poppius, 1909: 28 (Quedius; subgenus Raphirus; Lena Valley, Siberia).

  • — Reitter, 1909: 115 (Quedius; subgenus Raphirus; characters; Germany).

  • — Poppius, 1913: 10 (Quedius; Iceland).

  • — Johansen, 1914: 371 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Fiori, 1915: 16 (Quedius; Italy).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Henriksen and Lundbeck, 1918: 495 (Quedius; Greenland).

  • — Šulc, 1922 (Quedius; osmeterium; anatomy; histology; physiology).

  • — Gridelli, 1925: 165 (Quedius; subgenus Raphirus; characters).

  • — Portevin, 1929: 345 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1433 (Quedius; catalog).

  • — West, 1937: 38 (Quedius; Faroe Islands).

  • — Wüsthoff, 1938a: fig. 64–68 (Quedius; aedeagal illustration).

  • — Henriksen, 1939: 43 (Quedius; Greenland).

  • — V. Hansen, 1952: 156 (Quedius; subgenus Raphirus; characters; Denmark).

  • — Fagel, 1955a: 85 (Quedius; subgenus Raphirus; Greece).

  • — Larsson and Gígja, 1959: 93 (Quedius; biological notes; Iceland).

  • — Fagel, 1960b: 111 (Quedius; characters; Belgium; France; England; Germany; Sweden; Italy; Austria; Yugoslavia; Spain; Portugal; Corsica; Algeria; Iceland; Norway).

  • — Smetana, 1962b: 153 (Quedius; subgenus Raphirus; characters).

  • — Szujecki, 1963a: 35 (Quedius; Poland).

  • — Palm, 1963b: 154 (Quedius; subgenus Raphirus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Smetana, 1964d: 81 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Scheerpeltz, 1965b: 408 (Quedius; Sicily).

  • — Horion, 1965: 306 (Quedius; subgenus Raphirus; northern and central Europe; Spain; Portugal; Corsica; Italy).

  • — Szujecki, 1968a: 736 (Quedius; Poland).

  • — Kasule, 1970: 64 (Quedius; larval characters).

  • — Bordoni, 1974b: 13 (Quedius; subgenus Raphirus; notes; Italy).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Smetana, 1976d: 28 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 280 (Quedius; subgenus Raphirus; characters; Palaearctic region; Greenland; Iceland).

  • — Coiffait, 1978d: 165 (Quedius; Turkey).

  • — Tóth, 1984: 131 (Quedius; characters; Hungary).

  • — Lucht, 1987: 110 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 175 (Quedius; characters).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — J. Li, 1993: 41 (Quedius; China).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Sadler and Dugmore, 1995: 144 (Quedius; habitat notes; Iceland).

  • — Muona and Viramo, 1995: 9 (Quedius; collected from dung of dog; Finland).

  • — Luff and Owen, 2000: 39 (Quedius; subgenus Raphirus; scutellar variation; separation from Q. aridulus; Britain).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Tunisia, Algeria, Greenland, Iceland, Europe, Turkey, Russia.

fuscipennis Stephens, 1832: 222 (Quedius; [preoccupied]; Type locality: Suffolk; Devonshire).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of boops).

tauricus Nordmann, 1837: 78 (Philonthus; variety of boops Gravenhorst; Type locality: Taurus Chersonesus).

  • — Scheerpeltz, 1933: 1434 (Quedius; variety of boops).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; subspecies of boops; characters; Turkey).

coloratus W. Kolbe, 1912: 12 (Quedius; aberration of boops; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; aberration of boops).

  • — Gridelli, 1925: 165 (Quedius; aberration of boops).

  • — Scheerpeltz, 1933: 1434 (Quedius; aberration of boops).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; variety of boops; characters).

  • — Tóth, 1984: 131 (Quedius; variety of boops; charactes; Hungary).

arestor Tottenham, 1948a: 255 (Quedius; subgenus Arphirus; Type locality: Britain: Chesil Beach).

  • — Smetana, 1958: 397 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Fagel, 1960a: 111 (Quedius; synonym of boops).

  • — Smetana, 1960b: 259 (Quedius; subgenus Raphirus; Czechoslovakia).

  • — Smetana, 1962b: 153 (Quedius; synonym of boops).

  • — Lohse, 1964: 218 (Quedius; synonym of boops).

  • — Pope, 1977: 31 (Quedius; synonym of boops).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of boops; characters).

  • — Smetana, 1993b: 50 (Quedius; synonym of boops).

islandicus Fagel, 1960b: 113 (Quedius; subspecies of boops; Type locality: Island: Reynivellir).

  • — Coiffait, 1978: 281 (Quedius; subgenus Raphirus; subspecies of boops; characters).

borjoniensis Coiffait, 1969 [Note: Nomen nudum], see: grouziacus Coiffait, 1969.

borneensis Cameron, 1933b: 347 (Quedius; Type locality: British North Borneo: Kenokok, 3300 feet).

  • — Korge, 1963a: 87 (Quedius; subgenus Indoquedius).

  • Distribution: Borneo.

bovinus Coquerel, 1860, see: Othius.

brachypterus Coiffait, 1967e: 401 (Quedius; subgenus Sauridus; Type locality: Caucase).

  • — Coiffait, 1978: 220 (Quedius; subgenus Sauridus; characters; Caucasus).

  • Distribution: Caucasus.

brandmayri Zanetti, 1992b: 135 (Quedius; subgenus Sauridus; Type locality: Sicilia: M. Soro).

  • — Sparacio, 1995: 165 (Quedius; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Italy.

breviceps Casey, 1915: 411 (Microsaurus; Type locality: British Columbia: Stikine River).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Hatch, 1957: 227 (Quedius; subgenus Sauridus; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1964: 38 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1965a: 14 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1971: 202 (Quedius; subgenus Raphirus; characters; British Columbia; Alaska; California; Oregon; Washington).

  • — Smetana, 1971a: 1847 (Quedius; subgenus Raphirus; collecting notes; California).

  • — Smetana, 1971b: 4 (Quedius; subgenus Raphirus; Oregon).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; California).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • Distribution: Canada, USA.

brevicornis Thomson, 1860: 175 (Microsaurus; Type locality: Not cited but presumably Sweden).

  • — Sharp, 1867: 451 (Quedius; characters; habitat).

  • — Fauvel, 1874: 277 [= 1874a: 503] (Quedius; characters; France; Scandinavia; Britain; Austria; Russia).

  • — Mulsant and Rey, 1876: 636 [= 1877: 492] (Quedius; characters; France).

  • — Fowler, 1888: 233 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 398 (Quedius; subgenus Microsaurus; characters; north and middle Europe).

  • — Reitter, 1909: 110 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 358 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Gridelli, 1924d: 59 (Quedius; subgenus Microsaurus; characters).

  • — Gridelli, 1929: 35 (Quedius; characters).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 12 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 124, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 134 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 351 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 158 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 134 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 208 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 265 (Quedius; subgenus Microsaurus; Ireland; Britain; Norway; Sweden; Finland; Denmark; Baltics; Russia; Ukraine; Poland; Czechoslovakia; Hungary; Romania; Bosnia; Austria; Germany; France; Belgium; Holland).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 160 (Quedius; subgenus Microsaurus; characters; Europe; Britain; France).

  • — Hoebeke, 1978: 508 (Quedius; parasitized by proctotrupid).

  • — Tóth, 1984: 110 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Benelux; Denmark; Sweden).

  • — Schülke and Uhlig, 1988: 12 (Quedius; notes; Germany).

  • — Drugmand, 1989a: 326 (Quedius; pupal characters; Belgium).

  • — Nowosad, 1990: 136 (Quedius; collected from mole nest; notes; Poland).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Wittwer, 1993: 251 (Quedius; Switzerland).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Fowles, Alexander, and Key, 1999: 129 (Quedius; saproxylic species; status as endangered or threatened species; Britain).

  • Distribution: Europe, Russia.

eppelsheimii Seidlitz, 1891: 398 (Quedius; subgenus Microsaurus; [preoccupied]; Type locality: Eur. bis Curl.).

  • — Ganglbauer, 1895: 398 (Quedius; synonym of brevicornis).

  • — Thomson, 1896: 2389 (Microsaurus; synonym of pectinator).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of brevicornis).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

pectinator Thomson, 1896: 2389 (Microsaurus; replacement name for eppelsheimi Seidlitz; Sweden).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of brevicornis).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

elegans Ericson, 1897: 415 (Microsaurus; Type locality: Gernnebo in Vestergötland, Schweden).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 59 (Quedius; synonym of brevicornis).

  • — Scheerpeltz, 1933: 1434 (Quedius; synonym of brevicornis).

  • — Smetana, 1958: 351 (Quedius; synonym of brevicornis).

  • — Coiffait, 1978: 160 (Quedius; synonym of brevicornis).

brevipennis Motschulsky, 1858, see: sturanyi Ganglbauer, 1895.

brevipennis Fairmaire, 1859, see: asturicus Bernhauer, 1918.

brevipennis Ihssen, 1939, see: haberfelneri Eppelsheim, 1891.

brevis Erichson, 1840: 535 (Quedius; Type locality: Germania septentrionalis).

  • — Märkel, 1841: 217 (Quedius; in ant nest).

  • — Mannerheim, 1843a: 83 (Quedius; characters).

  • — Grimm, 1845: 135 (Philonthus; with Formica rufa; Germany).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Kraatz, 1857d: 500 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Thomson, 1860: 177 (Microsaurus; characters; Scandinavia).

  • — Hochhuth, 1862: 45 (Quedius; Russia).

  • — Fauvel, 1874: 275 [= 1874a: 501] (Quedius; characters; France).

  • — Redtenbacher, 1874: 197 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 625 [= 1877: 481] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 28 (Microsaurus; Finland).

  • — Fowler, 1888: 236 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; north and middle Europe).

  • — Porta, 1907: 111 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Reitter, 1909: 109 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 353 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Munster, 1923: 190 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 23 (Quedius; subgenus Microsaurus; characters; Germany; Bohemia; Silesia; Italy; Istria).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1434 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 1 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 122, 126 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — V. Hansen, 1952: 126 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 338 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 152 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 128 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 207 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 253 (Quedius; subgenus Microsaurus; northern and central Europe; Italy; France; Romania; Hungary; Czechoslovakia; Poland; Ukraine).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Bordoni, 1974: 54 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 165 (Quedius; subgenus Microsaurus; characters; Europe; Scandinavia; Britain).

  • — Frank, 1982: 45 (Quedius; list of parasites).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Drugmand, 1989: 175 (Quedius; characters).

  • — Janák and Vysoký, 1992: 137 (Quedius; collected with Formica rufa and polyctena; Bohemia).

  • — Franc, 1993: 308 (Quedius; synecthrical myrmecophile; Slovakia).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

lokayi M. Dvořák, 1967: 86 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum]; cited from Vlašim).

slovenicus M. Dvořák, 1967: 85 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum (Article 15.2)]; cited from Košice - Bankov).

fraudulentus Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; variety of brevis; [Note: Nomen nudum (Article 15.2)]; cited from Kosternick, Nukwaldy [Note: Correct spelling is Hukvaldy] Moravie, Tchécoslovaquie).

  • — Coiffait, 1978: 165 (Quedius; subgenus Microsaurus; variety of brevis; characters).

breweri† Scudder, 1890: 508 (Quedius; Type locality: Florissant; fossil).

  • — Scudder, 1891: 575 (Quedius; catalog; Oligocene; Colorado).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Miocene; Colorado).

  • Distribution: USA.

brieni Bruge and Drugmand, 1988, see: invreae Gridelli, 1924.

brookesi Cameron, 1947d: 706 (Quedius; subgenus Microsaurus; Type locality: New Zealand: Southland: Hump Ridge).

  • Distribution: New Zealand.

brouni Cameron, 1947d: 706 (Quedius; subgenus Microsaurus; Type locality: New Zealand: Canterbury: Pudding Hill).

  • Distribution: New Zealand.

bruchi Bernhauer, 1934b: 216 (Quedius; subgenus Cyrtoquedius, cited as Cryptoquedius; Type locality: Argentinien: Tucuman).

  • — Scheerpeltz, 1955a: 175 (Quedius; subgenus Cyrtoquedius; characters in key; Argentina).

  • Distribution: Argentina.

brunneorufus Bernhauer, 1941b: 30 (Quedius; subgenus Sauridus; Type locality: Neu Seeland: Greymouth).

  • Distribution: New Zealand.

brunnipennis Mannerheim, 1843b: 232 (Quedius; Type locality: insula Sitkha).

  • — Mannerheim, 1852: 314 (Quedius; characters).

  • — Horn, 1878: 164 (Quedius; characters; Lake Superior region westward to Vancouver and northward to Alaska).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Fall, 1926: 150 (Quedius; cited as brunneipennis; Alaska).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Hatch, 1957: 217 (Quedius; subgenus Distichalius; cited as brunneipennis; characters; British Columbia; Washington; Oregon).

  • — Smetana, 1971: 197 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Labrador; Newfoundland; Northwest Territories; Ontario; Quebec; Yukon Territory; Alaska; Michigan; New Hampshire; New York).

  • — Smetana, 1973: 1429 (Quedius; notes; Alberta; Northwest Territories; Quebec).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Northwest Territories; Ontario).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Labrador; Newfoundland).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: Canada, USA.

bruttius Zanetti, 1977: 307 (Quedius; Type locality: Gariglione, La Sila).

  • — Coiffait, 1978: 174 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; probably subspecies of latialis; Italy).

  • Distribution: Italy.

bryocharis Broun, 1923: 678 (Quedius; Type locality: Ben Lomond, 4500 ft.).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: New Zealand.

bucharensis Bernhauer, 1918a: 93 (Quedius; subgenus Ediquus; Type locality: Karateghingebirge: Baldschuan, 924 m; Buchara; Persien: Kopet-Dagh, Siaret, 1160 m).

  • — Gridelli, 1924d: 56 (Quedius; subgenus Microsaurus; characters; central Asia; Iran).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Coiffait, 1978: 186 (Quedius; subgenus Microsaurus; characters; Transcaspian region).

  • Distribution: “Transcaspia”, Uzbekistan, Iran.

bulgaricus Scheerpeltz, 1937b: 219 (Quedius; Type locality: Bulgarien: Rosalito-polje, Schipka-Balkan, ca. 1900 m).

  • — Coiffait, 1963a: 387 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 262 (Quedius; subgenus Sauridus; characters; Bulgaria).

  • Distribution: Bulgaria.

buphthalmus Erichson, 1840, see: Bolitogyrus.

bureschi Coiffait, 1971b: 283 (Quedius; subgenus Microsaurus; Type locality: Bulgarie: Aleko, 1800 m, Mt. Vitosha, près de Sofia).

  • — Coiffait, 1978: 149 (Quedius; subgenus Microsaurus; characters; Bulgaria).

  • Distribution: Bulgaria.

burrowsi Broun, 1915: 278 (Quedius; Type locality: Pudding Hill, near Methven).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: New Zealand.

cachemiricus Coiffait, 1982, see: fluviatilis Cameron, 1926.

caelebs Rottenberg, 1870: 28 (Quedius; Type locality: Sicily: Nicolosi; Palermo; Girgenti).

  • — Ganglbauer, 1895: 404 (Quedius; synonym of picipes).

  • — Porta, 1907: 135 (Quedius; synonym of picipes).

  • — Bernhauer and Schubert, 1916: 431 (Quedius; cited as coelebs; synonym of picipes).

  • — Gridelli, 1924d: 109 (Quedius; subgenus Raphirus; cited as coelebs characters; Sicily).

  • — Scheerpeltz, 1933: 1436 (Quedius; cited as coelebs; catalog).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; cited as coelebs; characters).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; cited as coelebs; characters; Sicily).

  • — Sparacio, 1995: 165 (Quedius; cited as coelebs; characters; notes; Sicily).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; cited as coelebs; Italy).

  • Distribution: Italy.

caelestis Smetana, 1996a: 54 (Quedius; subgenus Raphirus; Type locality: China: Yunnan, 50 km N Lijiang, Yulongshan Nat. Res.).

  • Distribution: China.

californicus Smetana, 1971, see: desertus Horn, 1878.

calignosus Stephens, 1832 [Note: Stephens, 1839: 392 cited this name as caliginosus], see: Callicerus.

calli Smetana, 1976a: 244 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Nuevo Leon: Cerro Potosi, 12,000 ft.).

  • Distribution: Mexico.

calogaster Lea, 1929: 206 (Quedius; Type locality: Queensland: Ravenshoe and Magnetic Island).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • Distribution: Australia.

campbelli Smetana, 1971: 103 (Quedius; subgenus Microsaurus; Type locality: Quebec: Gatineau Park, Lac Renaud).

  • — Smetana, 1976c: 172 (Quedius; subgenus Microsaurus; Ontario).

  • — Smetana, 1981b: 636 (Quedius; Saskatchewan).

  • — Smetana, 1990d: 97 (Quedius; Maine).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

canadensis Casey, 1915: 409 (Microsaurus; Type locality: Canada: Kazubazua, Province of Quebec).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1971: 53 (Quedius; subgenus Microsaurus; characters; New Brunswick; Ontario; Quebec; Connecticut; Massachusetts; Maine; Michigan; Minnesota; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; New York).

  • — Smetana, 1973: 1425 (Quedius; Ohio; Ontario; Quebec).

  • — Smetana, 1978a: 823 (Quedius; subgenus Microsaurus; collecting notes; North Carolina; Quebec).

  • — Smetana, 1981b: 633 (Quedius; collecting notes; New York; Ohio; Ontario).

  • — Smetana, 1990d: 96 (Quedius; Massachusetts; New Hampshire; New York; Ohio; Pennsylvania).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

candicus Coiffait, 1976b: 90 (Quedius; subgenus Sauridus; Type locality: Crête: Héraklion, Gonie).

  • — Coiffait, 1978: 228 (Quedius; subgenus Sauridus; characters; Crete).

  • Distribution: Greece.

capitalis Eppelsheim, 1892: 329 (Quedius; Type locality: Taschkent).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; characters; Tashkent).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Coiffait, 1978: 148 (Quedius; subgenus Microsaurus; characters; Turkestan).

  • Distribution: Uzbekistan.

capucinus Gravenhorst, 1806: 40 (Staphylinus; Type locality: America septentr.; [Note: See lectotype designation by Smetana, 1971]).

  • — Nordmann, 1837: 79 (Philonthus; America borealis).

  • — Erichson, 1840: 531 (Quedius; characters; America septentrionalis).

  • — Gravenhorst, 1847: 235 (Quedius; characters; notes).

  • — Provancher, 1877: 728 (Quedius; characters; Quebec).

  • — Horn, 1878: 160 (Quedius; characters; California; Atlantic region to Georgia).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Casey, 1915: 405 (Distichalius; characters; New York to Virginia westward to Iowa).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Voris, 1939: 154 (Quedius; larval and pupal characters; Indiana).

  • — Smetana, 1971: 161 (Quedius; subgenus Distichalius; lectotype designation; characters; Ontario; Quebec; Alabama; Arkansas; Connecticut; Georgia; Illinois; Indiana; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; South Carolina; Tennessee; Texas; Virginia; West Virginia; Wisconsin).

  • — Smetana, 1971b: 3 (Quedius; subgenus Distichalius; Maine).

  • — Smetana, 1978a: 827 (Quedius; subgenus Distichalius; collecting notes; Louisiana; Texas).

  • — Smetana, 1981b: 640 (Quedius; Ontario).

  • — Hunter, Kincher, Bay, Beerwinkle, 1991: 169 (Quedius; seasonal distribution and diel flight activity; Texas).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Distichalius; characters; USA).

  • Distribution: Canada, USA.

inversus Say, 1830: 36 [= 1834: 449] (Staphylinus; Type locality: Indiana).

  • — Erichson, 1840: 531 (Quedius; synonym of capucinus).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

bardus Melsheimer, 1844: 39 (Quedius; Type locality: Pennsylvania).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1916: 419 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1435 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

ater Ziegler, 1845: 266 (Philonthus; Type locality: Pennsylvania).

  • — Fauvel, 1869: 488 (Quedius; synonym of capucinus).

  • — Horn, 1878: 160 (Quedius; synonym of capucinus).

  • — Bernhauer and Schubert, 1914: 329 (Philonthus; catalog).

  • — Scheerpeltz, 1933: 1435 (Quedius; synonym of capucinus).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

seeversi Korge, 1963: 122 (Quedius; subgenus Distichalius; Type locality: California: Contra Costa).

  • — Smetana, 1971: 161 (Quedius; synonym of capucinus).

carnicus Fiori, 1894, see: cincticollis Kraatz, 1857.

carpaticola Roubal, 1924, see: cincticollis Kraatz, 1857.

caseyi Scheerpeltz, 1933, see: uteanus Casey, 1915.

castaneipennis Hochhuth, 1872, see: puncticollis Thomson, 1867.

castanopterus Stephens, 1832, see: Philonthus.

cavelli Broun, 1893: 1032 (Quedius; Type locality: Boatman's).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: New Zealand.

centrasiaticus Coiffait, 1969: 54 (Quedius; subgenus Raphirus; replacement name for altaicus Coiffait).

  • — Coiffait, 1978: 267 (Quedius; subgenus Raphirus; characters; central Asia).

  • Distribution: Russia.

altaicus Coiffait, 1967e: 422 (Quedius; subgenus Raphirus; [preoccupied]; Type locality: Telekschk Ozero, 1000 m, Altaï).

  • — Coiffait, 1978: 267 (Quedius; synonym of centrasiaticus).

cephalotes Uéno and Watanabe, 1966: 326 (Quedius; subgenus Microsaurus; Type locality: Japan: on the central massif of Shikoku: Ehime Prefecture: a limestone cave called “Rakan-ana”, at Koya in Sogawa of Nomura-cho).

  • — Uéno and Watanabe, 1970: 18 (Quedius; subgenus Microsaurus; notes; Japan).

  • — Watanabe, 1996a: 13 (Quedius; cave and collections; Japan).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species; Japan).

  • Distribution: Japan.

chalceiventris Fauvel, 1878f: 275 (Quedius; Type locality: Nouvelle-Guinée, Hatam).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: New Guinea.

chalybeipennis W. J. MacLeay, 1873, see: ruficollis Gravenhorst, 1806.

chamberlini† Scudder, 1890: 508 (Quedius; Type locality: Florissant; fossil).

  • — Scudder, 1891: 575 (Quedius; catalog; Oligocene; Colorado).

  • — Handlirsch, 1907: 723 (Quedius; catalog; Miocene; Colorado).

  • — Wickham, 1912: 11 (Quedius; Colorado).

  • — Wickham, 1913: 287 (Quedius; Colorado; doubtful identification).

  • Distribution: USA.

championi Sharp, 1884: 329 (Quedius; Type locality: Guatemala, San Geronimo 3000 feet, Purula).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • Distribution: Guatemala.

chang Smetana, 1995: 83 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien: Anmashan, 2230 m).

  • Distribution: Taiwan.

charactus Hatch, 1957: 222 (Quedius; subgenus Microsaurus; Type locality: Oregon: Curry Co.: Pistol River. Also cited from British Columbia).

  • — Smetana, 1971: 79 (Quedius; subgenus Microsaurus; characters; British Columbia; California; Oregon; Washington).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; British Columbia).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Smetana, 1981b: 634 (Quedius; collecting notes; Washington).

  • Distribution: Canada, USA.

chatterjeei Cameron, 1926b: 370 (Quedius; Type locality: Woodstock Falls, Mussoorie; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Mussorie and Chakrata districts, 6000 to 9000 feet).

  • — Cameron, 1932: 295 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1988: 229 (Quedius; subgenus Distichalius; lectotype designation: Woodstock Falls, Mussoorie; characters; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

chi Smetana, 1995: 76 (Quedius; subgenus Raphirus; Type locality: Taiwan: Taichung Hsien: Hsuehshan, above Shan-Liu-Gieu Hut, 3200 m).

  • Distribution: Taiwan.

chiangi Smetana, 1995: 48 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Nantou Hsien, Yushan N. P., Pai-Yun Hut, 3528 m).

  • Distribution: Taiwan.

chiaw Smetana, 1995: 56 (Quedius; subgenus Microsaurus; Type locality: Taiwan: Meifo Formosa).

  • — Smetana, 1996g: 24 (Quedius; notes; Taiwan).

  • Distribution: Taiwan.

chinensis Bernhauer, 1915c: 74 (Quedius; subgenus Raphirus; Type locality: Kiautschou).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1925: 149 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 52 (Quedius; aedeagal illustration).

  • — Smetana, 1996c: 226 (Quedius; subgenus Raphirus; characters; China).

  • Distribution: China.

chiricahuensis Smetana, 1971, see: leechi Smetana, 1971.

chlorophanus Erichson, 1840: 532 (Quedius; Type locality: Bengala).

  • — Kraatz, 1859: 66 (Quedius; characters; Bengalia).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Cameron, 1932: 283 (Quedius; subgenus Microsaurus; characters; Bengal).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Smetana, 1988: 184 (Quedius; may not be in Quedius; probably not in Himalaya).

  • Distribution: India.

chremes Smetana, 1996: 10 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan Langmusi, 3500–3600 m).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

chrysogonus Smetana, 1997b: 132 (Quedius; subgenus Raphirus; Type locality: China: Mt. Ailao Shan, 2500 m, Jingdong, Yunnan).

  • — Smetana, 1998b: 109 (Quedius; subgenus Raphirus; characters).

  • Distribution: China.

chrysurus Kiesenwetter, 1848, see: microps Gravenhorst, 1847.

cincticollis Kraatz, 1857d: 508 (Quedius; Type locality: Wien).

  • — Redtenbacher, 1857: 985 (Quedius; characters; Austria).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Mulsant and Rey, 1876: 718 [= 1877: 574] (Quedius; characters; Austria; Hungary).

  • — Ganglbauer, 1895: 409 (Quedius; subgenus Raphirus; characters; eastern Alps; Carpathians).

  • — Porta, 1907: 144 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Gridelli, 1925: 141 (Quedius; subgenus Raphirus; characters; Alps; Bosnia).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 45 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 380 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 139 (Quedius; subgenus Raphirus; characters; Alps and Carpathians).

  • — Coiffait, 1963a: 379 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 213 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 288 (Quedius; subgenus Raphirus; Austria; Slovenia; Italy; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Ukraine).

  • — Shilov, 1975a: 377 [= 1975b: 99] (Quedius; Komi region of Russia).

  • — Coiffait, 1978: 225 (Quedius; subgenus Sauridus; characters; Alps; Carpathians; Bosnia; mountains of central Europe).

  • — Tóth, 1984: 135 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Poland; Czechoslovakia; Austria).

  • — Lohse and Lucht, 1989: 172 (Quedius; characters).

  • — Nowosad, 1990: 146 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • Distribution: Europe, Russia.

carnicus Fiori, 1894: 88 (Quedius; Type locality: Visdende, nelle Alpi carniche).

  • — Porta, 1907: 144 (Quedius; synonym of cincticollis).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cincticollis).

  • — Gridelli, 1925: 141 (Quedius; synonym of cincticollis).

  • — Smetana, 1958: 380 (Quedius; synonym of cincticollis).

  • — Coiffait, 1978: 225 (Quedius; synonym of cincticollis).

carpaticola Roubal, 1924: 245 (Quedius; variety of cincticollis; Type locality: Pietros).

  • — Scheerpeltz, 1933: 1435 (Quedius; variety of cincticollis).

  • — Smetana, 1958: 380 (Quedius; cited as carpathicola; variety of cincticollis).

  • — Coiffait, 1978: 225 (Quedius; cited as carpathicola; variety of cincticollis; characters).

  • — Tóth, 1984: 135 (Quedius; cited as carpathicola; subspecies of cincticollis; characters).

cinctipennis Cameron, 1951c: 406 (Quedius; Type locality: Kenya Colony: Kanziko).

  • Distribution: Kenya.

cinctus Paykull, 1790: 137 (Staphylinus; Type locality: Suecia).

  • — Paykull, 1800: 395 (Staphylinus; characters; Sweden).

  • — Gravenhorst, 1802: 35 (Staphylinus; synonym of impressus).

  • — Marsham, 1802: 505 (Staphylinus; synonym of marginellus).

  • — C. Sahlberg, 1830: 317 (Staphylinus; synonym of impressus).

  • — O. Costa, 1839: 47 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 420, cited marginellus Costa as a synonym of cinctus Paykull; Costa did not describe the species as new, he attributed it to Fabricius]; characters; Italy).

  • — Erichson, 1839a: 489 (Quedius; synonym of impressus).

  • — Erichson, 1840: 530 (Quedius; synonym of impressus).

  • — Kiesenwetter, 1848: 50 (Quedius; characters; Europe).

  • — Kraatz, 1857d: 499 (Quedius; synonym of impressus).

  • — Crotch, 1870: 100 (Quedius; valid name).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; characters; France; Algeria; Turkey).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 676 [= 1877: 532] (Quedius; characters; larval characters; France).

  • — J. Sahlberg, 1876: 24 (Microsaurus; Finland).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; Algeria).

  • — Fowler, 1888: 235 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 402 (Quedius; subgenus Quedionuchus; characters; Europe; Mediterranean region).

  • — Fauvel, 1902b: 115 (Quedius; synonym of flavescens).

  • — Poppius, 1905: 9 (Quedius; Shetland Islands).

  • — Porta, 1907: 107 (Quedius; subgenus Quedionuchus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Reitter, 1909: 111 (Quedius; subgenus Quedionuchus; characters; Germany).

  • — Johansen, 1914: 362 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Munster, 1923: 194 (Quedius; subgenus Quedionuchus; Fennoscandia).

  • — Gridelli, 1924d: 75 (Quedius; subgenus Quedionuchus; characters; Albania).

  • — Portevin, 1929: 338 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1435 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 18 (Quedius; aedeagal illustration).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Jeannel and Jarrige, 1949: 38 (Quedius; subgenus Quedionuchus; notes; cave species; Serbia).

  • — V. Hansen, 1952: 139 (Quedius; subgenus Quedionuchus; characters; Denmark).

  • — Coiffait, 1955e: 427 (Quedius; Lebanon).

  • — Scheerpeltz, 1956c: 1102 (Quedius; subgenus Quedionuchus; Cyprus).

  • — Smetana, 1958: 357 (Quedius; subgenus Quedionuchus; characters; ecology; Czechoslovakia).

  • — Smetana, 1959a: 215 (Quedius; Albania).

  • — Smetana, 1960e: 160 (Quedius; subgenus Quedionuchus; characters).

  • — Scheerpeltz, 1960d: 136 (Quedius; subgenus Quedionuchus; Portugal).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Quedionuchus; Italy).

  • — Scheerpeltz, 1961e: 137 (Quedius; subgenus Quedionuchus; Rhodes).

  • — Ferreira, 1962: 19 (Quedius; Portugal).

  • — Scheerpeltz, 1963a: 420 (Quedius; subgenus Quedionuchus; Greece).

  • — Palm, 1963b: 137 (Quedius; subgenus Quedionuchus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 210 (Quedius; subgenus Quedionuchus; characters; central Europe).

  • — Korge, 1964: 118 (Quedius; subgenus Quedionuchus; characters).

  • — Scheerpeltz, 1964a: 300 (Quedius; subgenus Quedionuchus; Crete).

  • — Horion, 1965: 269 (Quedius; subgenus Quedionuchus, cited as Quedionychus; Caucasus; Morocco; Tunisia; Cyprus; Turkey; Scotland; Lapland; Russia; all central and southern Europe).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Scheerpeltz, 1968a: 90 (Quedius; catalog; Austria).

  • — Kasule, 1970: 59 (Quedionuchus; larval characters).

  • — Smetana, 1971: 165 (Quedius; subgenus Distichalius; characters; Massachusetts).

  • — Korge, 1971a: 47 (Quedius; subgenus Quedionuchus; characters; Turkey).

  • — Bordoni, 1973b: 49 (Quedius; subgenus Distichalius; characters; Italy).

  • — Coiffait, 1976b: 82 (Quedius; Greece; Rhodes; Thasos).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 186 (Quedius; subgenus Distichalius; characters; Europe; Scandinavia; Britain; Asia Minor; Middle East; North Africa).

  • — Smetana, 1978: 123 (Quedius; subgenus Distichalius; collecting notes; Israel).

  • — Muona, 1979: 15 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

  • — Burakowski, Mroczkowski, and Stefańska, 1980: 131 (Quedius; catalog; Poland).

  • — Hinton, 1981: 663 (Quedius; characters of egg).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Bordoni, 1984: 342 (Quedius; Lebanon).

  • — Tóth, 1984: 103 (Quedius; characters; Hungary).

  • — Muona and Viramo, 1986: 14 (Quedius; Finland).

  • — H. Franz, 1987: 70 (Quedius; Cyprus).

  • — Ádám, 1987: 152 (Distichalius; collecting notes; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Lohse and Lucht, 1989: 172 (Quedius; subgenus Distichalius).

  • — Smetana, 1990d: 99 (Quedius; New Jersey; New York; Washington).

  • — Whitehead, 1990e: 11 (Quedius; collecting notes; Britain).

  • — Allen, 1990b: 2 (Quedius; collecting notes; Britain).

  • — Angelini, 1991: 193 (Quedius; Italy).

  • — Silfverberg, 1992: 18 (Quedius; Finland; Russia; Sweden; Norway; Denmark; Estonia; Latvia; Lithuania).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 50 (Quedius; subgenus Distichalius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 31 (Quedius; Italy).

  • — Downie and Arnett, 1996: 392 (Quedius; subgenus Distichalius; characters; USA).

  • — M. Hansen, 1996: 107 (Quedius; checklist; Denmark).

  • — Ádám, 1996: 247 (Distichalius; synonym of Staphylinus flavicornis Gmelin; [Note: Until the identity of the Gmelin name has been verified it is best to allow it to remain in Staphylinus as a nomen dubium; furthermore, even if flavicornis and cinctus are synonyms, the latter should be protected via Article 23.9.1]).

  • — Eisinger, 1997: 171 (Quedius; collecting notes; Germany).

  • — T. Wagner, 1997: 235 (Quedius; collecting notes; Germany).

  • — Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Dauphin, Laguerre, Tamisier, and Teissier, 1997: 20 (Quedius; France).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Quedionuchus; facultative cave species; Serbia).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • — Herman, 2001: 29 (Quedius; nomen protectum; discussion).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Georgia, Turkey, Lebanon, Israel, USA.

flavescens Linné, 1758: 422 (Staphylinus; [Note: This name is older than cinctus but has hardly been cited since its original description. In 1775 Fabricius attributed flavescens to Linné, 1767: 684. Most subsequent workers have credited Fabricius with the name]; Type locality: Europa).

  • — Linné, 1760: 232 (Staphylinus; characters; Sweden).

  • — Pontoppidans, 1765: 211 (Staphylinus; Denmark).

  • — Linné, 1767: 684 (Staphylinus; characters; Europe).

  • — Fabricius, 1775: 266 (Staphylinus; characters; Europe).

  • — O. Müller, 1776: 98 (Staphylinus; characters).

  • — Goeze, 1777: 719 (Staphylinus; characters).

  • — Fabricius, 1781: 336 (Staphylinus; characters; Europe).

  • — Fabricius, 1787: 221 (Staphylinus; characters).

  • — Villers, 1789: 415 (Staphylinus; characters).

  • — Gmelin, 1790: 2029 (Staphylinus; characters).

  • — Fabricius, 1793: 526 (Staphylinus; characters).

  • — Panzer, 1795: 355 (Staphylinus; characters; Germany).

  • — Cederhjelm, 1798: 118 (Staphylinus; characters).

  • — Fabricius, 1801: 597 (Staphylinus; characters).

  • — Turton, 1802: 512 (Staphylinus; characters; Europe).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 324 (Staphylinus; synonym of discoideus Gravenhorst).

  • — Westwood, 1845: 49 (Staphylinus; notes; no specimen in Linnean cabinet).

  • — Kraatz, 1857d: 499 (Quedius; synonym of impressus).

  • — Gemminger and Harold, 1868: 570 (Quedius; synonym of impressus).

  • — Fauvel, 1902b: 115 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1436, cited flavescens Fauvel as a synonym of cinctus; Fauvel did not describe the species as new, he attributed it to Linné]; Morocco; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 186 (Quedius; synonym of cinctus).

  • — Herman, 2001: 29 (Quedius; nomen oblitum).

impressus Panzer, 1796d: 21 (Staphylinus; [preoccupied]; Type locality: Neuwaldegg).

  • — Gravenhorst, 1802: 35 (Staphylinus; characters).

  • — Latreille, 1804: 320 (Staphylinus; characters).

  • — Gravenhorst, 1806: 39 (Staphylinus; characters; Germany).

  • — Panzer, 1808a: 21 (Staphylinus; characters; illustrations).

  • — Gyllenhal, 1810: 307 (Staphylinus; characters; Sweden).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 317 (Staphylinus; characters; Finland).

  • — Mannerheim, 1830: 26 (Staphylinus; Finland; Sweden).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland; Sweden).

  • — Stephens, 1832: 219 (Quedius; characters; England).

  • — Brullé, 1832: 131 (Staphylinus; Greece).

  • — Lacordaire, 1835: 381 (Emus; characters; France).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Nordmann, 1837: 79 (Philonthus; Russia; Germany; Sweden).

  • — Westwood, 1838a: 16 (Quedius; type species).

  • — Erichson, 1839a: 489 (Quedius; characters; Germany).

  • — Heer, 1839: 274 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 530 (Quedius; characters; Europe).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — Laporte, 1840: 177 (Staphylinus; characters; France).

  • — Gravenhorst, 1847: 234 (Quedius; characters; notes).

  • — Hochhuth, 1849: 151 (Quedius; characters; Tiflis).

  • — Redtenbacher, 1849: 709 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Hardy, 1851: 32 (Quedius; notes; England).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Kraatz, 1857d: 499 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Thomson, 1860: 174 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1865a: 293 [= 1865b: 43] (Quedius; habitat; France).

  • — Baudi, 1870: 383 (Quedius; characters; Sardinia; Italy).

  • — Crotch, 1870: 100 (Quedius; synonym of cinctus).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; synonym of cinctus).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — J. Sahlberg, 1876: 24 (Microsaurus; synonym of cinctus).

  • — Fowler, 1888: 235 (Quedius; synonym of cinctus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of cinctus).

  • — Porta, 1907: 107 (Quedius; synonym of cinctus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1971: 165 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 186 (Quedius; synonym of cinctus).

rufocinctus Mannerheim, 1830: 26 (Staphylinus; Type locality: Finlandia australis; Sweden).

  • — Mannerheim, 1831: 440 (Staphylinus; Finland).

  • — Nordmann, 1837: 79 (Philonthus; Sweden; Finland).

  • — Erichson, 1840: 551 (Quedius; characters; Finland).

  • — Kraatz, 1857d: 501 (Quedius; Finland).

  • — Fauvel, 1874: 288 [= 1874a: 514] (Quedius; synonym of cinctus).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of cinctus).

  • — Ganglbauer, 1895: 402 (Quedius; synonym of cinctus).

  • — Porta, 1907: 107 (Quedius; synonym of cinctus).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; synonym of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1971: 165 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus).

niger Porta, 1907: 90, 107, 108 (Quedius; subgenus Quedionuchus; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Gridelli, 1924d: 75 (Quedius; aberration of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of cinctus).

  • — Smetana, 1958: 358 (Quedius; aberration of cinctus).

  • — Smetana, 1960e: 160 (Quedius; aberration of cinctus; characters).

  • — Scheerpeltz, 1965b: 407 (Quedius; aberration of cinctus; Sicily).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; subgenus Distichalius; variety of cinctus; characters).

  • — Tóth, 1984: 103 (Quedius; aberration of cinctus).

rufipennis Porta, 1907: 90, 107, 108 (Quedius; subgenus Quedionuchus; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; aberration of cinctus).

  • — Gridelli, 1924d: 75 (Quedius; aberration of cinctus).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of cinctus).

  • — Mařan, 1935: 9 (Quedius; variety of cinctus; Crete).

  • — Méquignon, 1948: 76 (Quedius; variety of cinctus; characters; France).

  • — Smetana, 1958: 358 (Quedius; aberration of cinctus).

  • — Smetana, 1960e: 160 (Quedius; aberration of cinctus; characters).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; subgenus Distichalius; variety of cinctus; characters).

  • — Tóth, 1984: 104 (Quedius; variety of cinctus).

africanus Peyerimhoff, 1917: 123 (Quedius; subgenus Microsaurus; subspecies of flavescens; Type locality: Barbarie).

  • — Gridelli, 1924d: 75 (Quedius; synonym of niger).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus niger).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus variety niger).

lividipennis Lokay, 1921: 71 (Quedius; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5); preoccupied]; cited from Candia, Kreta).

  • — Gridelli, 1924d: 75 (Quedius; synonym of rufipennis).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus rufipennis).

  • — Smetana, 1971: 166 (Quedius; synonym of cinctus).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

arabs Lokay, 1921: 71 (Quedius; aberration of cinctus; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Sidi bel Abbes).

  • — Gridelli, 1924d: 75 (Quedius; synonym of niger).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of cinctus).

  • — Smetana, 1958: 358 (Quedius; synonym of cinctus).

  • — Smetana, 1960e: 160 (Quedius; synonym of cinctus niger).

  • — Bordoni, 1973b: 49 (Quedius; synonym of cinctus).

  • — Coiffait, 1978: 187 (Quedius; synonym of cinctus variety niger).

cipactli Smetana, 1976a: 231 (Quedius; subgenus Quedionuchus; Type locality: Guatemala: Sacatepéquez: 10 km. W. San Miguel Duenas, 8000 ft.).

  • Distribution: Guatemala.

circumductus Mulsant and Rey, 1876, see: muscorum Brisout, 1863.

citelli Kirshenblat, 1933: 103 (Quedius; subgenus Microsaurus; Type locality: Transbaikalien: Adun-Tshelon).

  • — Coiffait, 1978: 185 (Quedius; subgenus Microsaurus; cited as citellus; characters; Transbaikal).

  • — Boháč, 1988: 554 [= 1989: 38] (Quedius; cited as citelli; characters).

  • Distribution: Russia.

clypealis Sharp, 1876: 100 (Quedius; Type locality: Ega).

  • — Bernhauer and Schubert, 1916: 420 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 177, 178 (Quedius; subgenus Cyrtoquedius; characters in key; Brazil).

  • Distribution: Brazil.

coatl Smetana, 1976a: 246 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Jalisco: E. slope Nevado de Colima, 10,000 ft.).

  • Distribution: Mexico.

cobosi Coiffait, 1964d: 293 (Quedius; subgenus Quedius; Type locality: Espagne: Province de Almeria, Seron).

  • — Coiffait, 1978: 197 (Quedius; subgenus Quedius; characters; Spain).

  • Distribution: Spain.

almeriensis Coiffait, 1964d: 294 (Quedius; subgenus Quedius; aberration of cobosi; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Espagne: Prov. de Almeria, Laujar).

  • — Coiffait, 1978: 197 (Quedius; subgenus Quedius; variety of cobosi; characters).

coeruleipennis Fauvel, 1874, see: curtus Erichson, 1840.

coeruleipennis Bernhauer, 1926, see: cyaneipennis Scheerpeltz, 1933.

coeruleus Coiffait, 1977, see: gardneri Cameron, 1932.

cohaesus Eppelsheim, 1888: 60 (Quedius; subgenus Sauridus; Type locality: Turcmenien).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1925: 126 (Quedius; subgenus Raphirus; characters; Turkmenia).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Coiffait, 1963a: 393 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 122 (Quedius; subgenus Sauridus; Turkey).

  • — Smetana, 1967i: 558 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1978: 248 (Quedius; subgenus Sauridus; characters; Turkestan; Caucasus; Transcaucasus; Turkey; Bulgaria).

  • Distribution: Turkmenistan, Caucasus Turkey, Bulgaria.

coiffaitianus Fagel, 1968b: 11 (Quedius; subgenus Sauridus; Type locality: Liban: Nabeh Safa, 1000 m).

  • — Coiffait, 1978: 245 (Quedius; subgenus Sauridus; characters; Lebanon).

  • Distribution: Lebanon.

collaris Erichson, 1840: 545 (Quedius; Type locality: Volhynia).

  • — Kraatz, 1857d: 515 (Quedius; characters).

  • — Redtenbacher, 1857: 986 (Quedius; characters; Austria).

  • — G. Waterhouse, 1858: 23 (Quedius; synonym of fulvicollis).

  • — Redtenbacher, 1874: 202 (Quedius; characters).

  • — Mulsant and Rey, 1876: 783 [= 1877: 639] (Quedius; characters; Volhynia).

  • — Ganglbauer, 1895: 411 (Quedius; subgenus Raphirus; characters; Alps; Sudeten; Carpathians).

  • — Porta, 1907: 148 (Quedius; subgenus Raphirus; characters; Italy).

  • — Reitter, 1909: 114 (Quedius; subgenus Raphirus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1925: 151 (Quedius; subgenus Raphirus; characters).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 54 (Quedius; aedeagal illustration).

  • — Smetana, 1958: 388 (Quedius; subgenus Raphirus; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 149 (Quedius; subgenus Raphirus; characters; Europe).

  • — Lohse, 1964: 218 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; Czechoslovakia; Poland; Romania; Bosnia Hercegovina; Macedonia; Austria).

  • — Coiffait, 1978: 273 (Quedius; subgenus Raphirus; characters; Carpathians; Alps; Apennines).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 126 (Quedius; characters).

  • — Lucht, 1987: 110 (Quedius; checklist; Poland; Czechoslovakia; Austria).

  • — Nowosad, 1990: 149 (Quedius; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; Czech Republic; Slovakia).

  • Distribution: Europe.

maculicollis W. Kolbe, 1909a: 416 (Quedius; aberration of collaris; [Note: Nomen nudum (Articles 1.3.4, 45.5)]).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of collaris).

  • — Scheerpeltz, 1933: 1436 (Quedius; aberration of collaris).

  • — Smetana, 1958: 388 (Quedius; aberration of collaris).

  • — Smetana, 1962b: 149 (Quedius; aberration of collaris; characters).

  • — Coiffait, 1978: 273 (Quedius; subgenus Raphirus; variety of collaris; characters).

  • — Tóth, 1984: 126 (Quedius; variety of collaris).

niger Gridelli, 1922: 25 (Quedius; subgenus Raphirus; variety of collaris; [preoccupied]; Type locality: Gran Sasso, Appennino centrale).

  • — Scheerpeltz, 1933: 1436 (Quedius; synonym of collaris).

  • — Coiffait, 1978: 274 (Quedius; synonym of collaris italicus).

italicus Gridelli, 1925: 151 (Quedius; replacement name for niger Gridelli; subspecies of collaris).

  • — Scheerpeltz, 1933: 1436 (Quedius; subspecies of collaris).

  • — Korge, 1964: 124 (Quedius; subgenus Raphirus; characters; Macedonia).

  • — Horion, 1965: 298 (Quedius; subgenus Raphirus; subspecies of collaris; Italy; France).

  • — Coiffait, 1978: 274 (Quedius; subgenus Raphirus; subspecies of collaris; characters).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of collaris; Italy).

collinus Broun, 1893: 1031 (Quedius; Type locality: Waitakerei Range).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: New Zealand.

coloratus Fauvel, 1875a: XXXIII [= 1875b: 235] (Quedius; Type locality: Palestine Naplouse; [Note: See lectotype designation by Smetana, 1978]. Originally cited from Syrie, Naplouse, Djebel-ech-Cheik).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 109 (Quedius; subgenus Raphirus; characters; Syria; Albania).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • — Scheerpeltz, 1958a: 23 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1963a: 378 (Quedius; subgenus Sauridus; characters).

  • — Korge, 1964: 119 (Quedius; subgenus Sauridus; Turkey).

  • — Smetana, 1967i: 558 (Quedius; subgenus Raphirus; Turkey).

  • — Coiffait, 1978: 212 (Quedius; subgenus Sauridus; characters; Balkans; Syria; Lebanon).

  • — Smetana, 1978: 123 (Quedius; subgenus Raphirus; lectotype designation: Palestine Naplouse; collecting notes; Israel).

  • Distribution: Israel, Lebanon, Syria, Turkey, Albania.

coloratus W. Kolbe, 1912 [Note: Nomen nudum], see: boops Gravenhorst, 1802.

columbinus Bernhauer, 1917: 91 (Quedius; subgenus Euryquedius; Type locality: West-Kolumbien: Pacho, in den Ost-Kordilleren, 2000 m; Venezuela).

  • — Scheerpeltz, 1933: 1436 (Quedius; catalog).

  • Distribution: Colombia, Venezuela.

compransor Fall, 1912: 40 (Quedius; Type locality: Manhattan, Kansas).

  • — Casey, 1915: 421 (Anastictodera).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Smetana, 1971: 61 (Quedius; subgenus Microsaurus; characters; Kansas).

  • — Smetana, 1978a: 823 (Quedius; subgenus Microsaurus; collecting notes; Florida; Kansas).

  • — Smetana, 1981b: 633 (Quedius; characters; collecting notes; Louisiana).

  • Distribution: USA.

concinnus Last, 1987: 52 (Quedius; Type locality: Papua New Guinea NE: Bulldog Road).

  • Distribution: New Guinea.

concolor Sharp, 1884: 332 (Quedius; Type locality: Guatemala, San Geronimo).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 173 (Quedius; subgenus Cyrtoquedius; characters in key; Guatemala).

  • Distribution: Guatemala.

concolor Sharp, 1889, see: samuraicus Bernhauer and Schubert, 1916.

conicus Champion, 1922, see: Indoquedius.

conicus Segers, 1987, see: segersi Herman, 2001.

consors Fauvel, 1866, see: Cheilocolpus.

conspicuellus Broun, 1894: 421 (Quedius; Type locality: Te Aroha).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: New Zealand.

constantini Jarrige, 1965a: 19 (Quedius; subgenus Raphirus; Type locality: Espagne: Pena Rubia, au N.-O. de Ponferrada, Province de Lugo, 1700 m).

  • — Coiffait, 1978: 230 (Quedius; subgenus Sauridus; characters; Spain).

  • Distribution: Spain.

cordatus Lea, 1925: 245 (Quedius; Type locality: Queensland: National Park).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

corion Tottenham, 1948a: 258 (Quedius; subgenus Arphirus; Type locality: Haxey).

  • — Smetana, 1958: 397 (Quedius; subgenus Raphirus; characters; ecology).

  • — Fagel, 1960b: 114 (Quedius; notes).

  • — Smetana, 1962b: 152 (Quedius; subgenus Raphirus; characters).

  • — Pope, 1977: 31 (Quedius; possibly synonym of aridulus).

  • — Coiffait, 1978: 277 (Quedius; subgenus Raphirus; characters; England).

  • Distribution: Britain, Czechoslovakia.

coronatus Delabie, 1953: 33 (Quedius; Type locality: versant N. du massif du Canigou Pyrénées Orientales, 1900 m).

  • — Coiffait, 1978: 204 (Quedius; subgenus Quediops; characters; Pyrenees).

  • Distribution: France.

delabiei Coiffait, 1977c: 136 (Quedius; subgenus Quediops; variety of coronatus; [Note: Nomen nudum (Article 15.2)]).

  • — Coiffait, 1978: 204 (Quedius; subgenus Quediops; variety of coronatus; characters).

coxalis Kraatz, 1858a: 59 (Quedius; Type locality: Athen).

  • — Fauvel, 1874: 297 [= 1874a: 523] (Quedius; characters; Italy; Greece; Syria; Palestine).

  • — Mulsant and Rey, 1876: 744 [= 1877: 600] (Quedius; characters; Italy; Greece).

  • — Sainte-Claire Deville, 1906: 102 (Quedius; Corsica).

  • — Porta, 1907: 139 (Quedius; subgenus Sauridus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1922b: 130, 134 (Quedius; characters; notes; Hungary; Transylvania; Italy; Balkan Peninsula; Greece; Syria; Palestine).

  • — Gridelli, 1925: 130 (Quedius; synonym of oblitteratus).

  • — Scheerpeltz, 1933: 1453 (Quedius; synonym of oblitteratus).

  • — Smetana, 1958: 378 (Quedius; synonym of oblitteratus).

  • — Fagel, 1968c: 2 (Quedius; subgenus Sauridus; valid species; characters; notes; Greece; Turkey; Israel; Romania).

  • — Coiffait, 1978: 213 (Quedius; subgenus Sauridus; characters; Greece).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Europe, Turkey, Syria, Lebanon, Israel.

atticus Coiffait, 1967e: 408 (Quedius; subgenus Sauridus; Type locality: Mont Pantélique près d'Athènes).

  • — Coiffait, 1977c: 140 (Quedius; synonym of coxalis).

  • — Coiffait, 1978: 214 (Quedius; synonym of coxalis).

crassus Fairmaire, 1860b: 153 (Quedius; Type locality: Mers-el-Kébir; Médéah).

  • — Fauvel, 1866c: 247 [= 1866d: 105] (Quedius; synonym of curtus).

  • — Fauvel, 1869: 486 (Quedius; valid species).

  • — Fauvel, 1874: 276 [= 1874a: 502] (Quedius; characters; France; Spain; Algeria).

  • — Mulsant and Rey, 1876: 634 [= 1877: 490] (Quedius; characters; France).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Tunisia; Algeria).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; France; Algeria).

  • — Fauvel, 1902b: 114 (Quedius; Algeria; Tunisia).

  • — Porta, 1907: 112 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; characters; France; Spain; Portugal; Algeria).

  • — Portevin, 1929: 339 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Roubal, 1941b: 123, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Bordoni, 1974: 72 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; characters; western Mediterranean region; North Africa; Iberian Peninsula; Italy; France).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Outerelo, Gamarra, and Salgado, 1998: 129 (Quedius; subgenus Microsaurus; cited as crassus rigidicornis Gridelli, 1924 [Note: I have found no use of rigidicornis by Gridelli]; collected from cave; Spain; Portugal).

  • Distribution: Tunisia, Algeria, Portugal, Spain, France, Italy.

amplicollis Scriba, 1868: 155 (Quedius; Type locality: Spanien).

  • — Fauvel, 1874: 276 [= 1874a: 502] (Quedius; synonym of crassus).

  • — Ganglbauer, 1895: 397 (Quedius; synonym of crassus).

  • — Porta, 1907: 112 (Quedius; synonym of crassus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of crassus).

  • — Gridelli, 1924d: 40 (Quedius; synonym of crassus).

  • — Scheerpeltz, 1933: 1437 (Quedius; synonym of crassus).

  • — Bordoni, 1974: 72 (Quedius; synonym of crassus).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; subspecies of crassus; characters).

nigricans Gridelli, 1924d: 40 (Quedius; subgenus Microsaurus; variety of crassus; Type locality: Monchique, Portogallo).

  • — Scheerpeltz, 1933: 1437 (Quedius; variety of crassus).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; variety of crassus; characters).

  • — Bordoni, 1974: 72 (Quedius; variety of crassus).

  • — Coiffait, 1978: 145 (Quedius; subgenus Microsaurus; variety of crassus; characters).

crescenti Hatch, 1957: 220 (Quedius; subgenus Microsaurus; Type locality: Washington: L. Crescent. Also cited from Oregon).

  • — Smetana, 1971: 100 (Quedius; subgenus Microsaurus; characters; Oregon; Washington).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; British Columbia).

  • — Smetana, 1981b: 634 (Quedius; collecting notes; Oregon).

  • Distribution: Canada, USA.

creticus Mařan, 1935, see: fulgidus Fabricius, 1793.

criddlei Casey, 1915: 410 (Microsaurus; Type locality: Manitoba: Aweme).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Smetana, 1971: 107 (Quedius; subgenus Microsaurus; characters; Alberta; British Columbia; Manitoba; Northwest Territories; Yukon Territories; Colorado; Idaho; Montana; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; collecting notes; Washington).

  • — Smetana, 1981b: 636 (Quedius; collecting notes; Alaska; Saskatchewan).

  • — Smetana, 1990d: 97 (Quedius; Alaska; Ontario).

  • Distribution: Canada, USA.

hoodi Hatch, 1957: 221 (Quedius; subgenus Microsaurus; Type locality: Oregon: Mt. Hood, Timberline. Also cited from Washington, Idaho, and British Columbia).

  • — Smetana, 1971: 107 (Quedius; synonym of criddlei).

crius Tottenham, 1948, see: reitteri Gridelli, 1925.

crnagoricus Coiffait, 1980a: 38 (Quedius; subgenus Raphirus; Type locality: Monténégro, Col de Cakor, 1850 m).

  • — Drugmand, 1988c: 204 (Quedius; characters).

  • Distribution: Yugoslavia.

cruentus Olivier, 1795:(42): 27 (Staphylinus; Type locality: Paris).

  • — Gravenhorst, 1802: 25 (Staphylinus; synonym of fulvipes).

  • — Marsham, 1802: 503 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 421 listed fulgidus Marsham as a synonym of cruentus; Marsham did not describe the species as new, he attributed it to Fabricius]; characters; Britain).

  • — Gyllenhal, 1810: 343 (Staphylinus; synonym of fulvipes).

  • — C. Sahlberg, 1830: 329 (Staphylinus; synonym of fulvipes).

  • — Erichson, 1840: 527 (Quedius; characters; Germany; France; Italy; Volhynia).

  • — Hochhuth, 1849: 150 (Quedius; characters; Iberia).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Fairmaire and Laboulbène, 1856: 540 (Quedius; characters; France).

  • — Kraatz, 1857d: 495 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Kraatz, 1858a: 58 (Quedius; Greece).

  • — Kraatz, 1859: 67 (Quedius; characters; India orientalis).

  • — Thomson, 1867: 162 (Microsaurus; characters; Scandinavia).

  • — Fauvel, 1874: 282 [= 1874a: 508] (Quedius; characters; France; Caucasus; East Indies).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 661 [= 1877: 517] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 25 (Microsaurus; Finland).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; notes; Algeria).

  • — Fowler, 1888: 234 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; characters; Europe; Caucasus; East Indies).

  • — Fauvel, 1902b: 114 (Quedius; Morocco; Algeria; Tunisia).

  • — Porta, 1907: 115 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Johansen, 1914: 354 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Munster, 1923: 192 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 53 (Quedius; subgenus Microsaurus; characters; Belgium; Styria; Italy; Albania; Dalmatia).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Cameron, 1932: 289 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Wüsthoff, 1938a: fig. 10 (Quedius; aedeagal illustration).

  • — Roubal, 1941b: 123, 128 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Microsaurus; cave species; France).

  • — V. Hansen, 1952: 135 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Smetana, 1958: 350 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; characters).

  • — Scheerpeltz, 1961d: 132 (Quedius; subgenus Microsaurus; Italy).

  • — Palm, 1963b: 135 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Horion, 1965: 263 (Quedius; subgenus Microsaurus; Morocco; Algeria; Turkey; all southern and central Europe; Denmark; Britain; Ireland; Norway; Sweden; Finland; Russia).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Bordoni, 1974: 40 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Legner and Moore, 1977: 176 (Quedius; found under bark).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; characters; Europe; Britain; Scandinavia; North Africa; Asia Minor; India).

  • — Hinton, 1981: 662 (Quedius; characters of egg).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 113 (Quedius; characters; Hungary).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Welch, 1993: 229 (Quedius; ovariole number and ovary structure).

  • — Smetana, 1993b: 49 (Quedius; subgenus Microsaurus; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 1999b: 61 (Quedius; captured underground at roots of old trees; Britain).

  • — Owen, 2000: 251 (Quedius; pitfall trap collections from underground at roots of trees; Britain).

  • Distribution: Tunisia, Algeria, Morocco, Europe, Russia, Caucasus, Turkey, India, “East Indies”.

haemorrhous Stephens, 1832: 217 (Quedius; Type locality: London; Norfolk; Suffolk).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of nitidus; characters; England).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

erythropterus Stephens, 1832: 226 (Quedius; Type locality: London).

  • — Stephens, 1839: 393 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

analis Stephens, 1835: 435 (Quedius; [preoccupied]; Type locality: London; Yorkshire).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of cruentus).

  • — Gemminger and Harold, 1868: 568 (Quedius; synonym of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; synonym of cruentus).

  • — Cameron, 1932: 289 (Quedius; synonym of cruentus).

  • — Smetana, 1958: 350 (Quedius; synonym of cruentus).

  • — Bordoni, 1974: 41 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; synonym of cruentus).

virens Rottenberg, 1870: 28 (Quedius; variety of fulgidus; Type locality: Sicily).

  • — Fauvel, 1874: [= 1874a:] (Quedius; race of cruentus; characters; Scandinavia; Russia; Tyrol; Italy; Corsica; Sardinia; Sicily).

  • — Duvivier, 1883: 138 (Quedius; catalog; synonym of cruentus).

  • — Ganglbauer, 1895: 399 (Quedius; variety of cruentus).

  • — Porta, 1907: 115 (Quedius; variety of cruentus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of cruentus).

  • — Gridelli, 1924d: 53 (Quedius; subgenus Microsaurus; variety of cruentus; Styria; Italy; Croatia; Dalmatia).

  • — Scheerpeltz, 1933: 1437 (Quedius; aberration of cruentus).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; aberration of cruentus; characters).

  • — Smetana, 1958: 350 (Quedius; aberration of cruentus).

  • — Smetana, 1960e: 157 (Quedius; aberration of cruentus; characters).

  • — Bordoni, 1974: 41 (Quedius; aberration of cruentus).

  • — Pope, 1977: 31 (Quedius; synonym of cruentus).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; variety of cruentus; characters).

  • — Tóth, 1984: 114 (Quedius; aberration of cruentus; characters; Hungary).

obscurus Lokay, 1913: 138, 140 (Quedius; variety of cruentus; Type locality: Kreta).

  • — Scheerpeltz, 1933: 1437 (Quedius; variety of cruentus).

  • — Mařan, 1935: 8 (Quedius; variety of cruentus; Crete).

  • — Roubal, 1941b: 123 (Quedius; subgenus Microsaurus; aberration of cruentus; characters).

  • — Smetana, 1958: 350 (Quedius; aberration of cruentus).

  • — Smetana, 1960e: 157 (Quedius; aberration of cruentus; characters).

  • — Scheerpeltz, 1961d: 133 (Quedius; subgenus Microsaurus; aberration of cruentus; Italy).

  • — Bordoni, 1974: 41 (Quedius; aberration of cruentus).

  • — Coiffait, 1978: 152 (Quedius; subgenus Microsaurus; variety of cruentus; characters).

  • — Tóth, 1984: 114 (Quedius; aberration of cruentus).

cuprinus Fauvel, 1877: 274 (Quedius; Type locality: Australie, Nouvelle Galles du Sud, Sydney).

  • — Lea, 1912: 40 (Quedius; with Camponotus nigriceps; Victoria).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Scheerpeltz, 1935a: 655 (Quedius; Australia).

  • Distribution: Australia.

cursor Smetana, 1971: 99 (Quedius; subgenus Microsaurus; Type locality: California: Luther Pass).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes).

  • Distribution: USA.

curtidens Smetana, 1967i: 559 (Quedius; subgenus Raphirus; Type locality: Türkei: Suluhan, Toros).

  • — Coiffait, 1978: 279 (Quedius; subgenus Raphirus; characters; Turkey).

  • Distribution: Turkey.

curtipennis Bernhauer, 1908e: 335 (Quedius; form of fuliginosus; Type locality: Buchara; Böhmen, Wran a. Moldau).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; variety of fuliginosus).

  • — Gridelli, 1924d: 79 (Quedius; variety of fuliginosus).

  • — Scheerpeltz, 1933: 1441 (Quedius; variety of fuliginosus).

  • — V. Hansen, 1952: 141 (Quedius; subgenus Quedius; characters; Denmark).

  • — Smetana, 1958: 362 (Quedius; subgenus Quedius; characters; ecology; Czechoslovakia).

  • — Korge, 1962b: 332 (Quedius; subgenus Quedius; synonymic notes).

  • — Smetana, 1962b: 133 (Quedius; subgenus Quedius; characters; Czechoslovakia).

  • — Palm, 1963b: 141 (Quedius; subgenus Quedius; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 211 (Quedius; subgenus Quedius; characters; central Europe).

  • — Korge, 1964: 119 (Quedius; subgenus Quedius; Turkey).

  • — Smetana, 1965: 38 (Quedius; subgenus Quedius; characters).

  • — Smetana, 1965a: 12 (Quedius; British Columbia).

  • — M. Dvořák, 1965: 89 (Quedius; Slovakia).

  • — Horion, 1965: 272 (Quedius; subgenus Quedius; Turkey; Faroes; Britain; Norway; Sweden; Finland; Russia; Denmark; all central Europe; Spain; Azores; Corsica; Italy; Bosnia; Romania).

  • — Szujecki, 1968a: 735 (Quedius; Poland).

  • — Frank, 1969: 267 (Quedius; biological notes; England).

  • — Smetana, 1970: 62 (Quedius; distributional summary; Azores).

  • — Smetana, 1971: 129 (Quedius; subgenus Quedius; characters; British Columbia; Oregon; Washington).

  • — Smetana, 1971b: 3 (Quedius; subgenus Quedius; British Columbia).

  • — Coiffait, 1973a: 275 (Quedius; Morocco).

  • — Bordoni, 1974b: 10 (Quedius; subgenus Quedius; characters; Italy).

  • — Bordoni, 1976b: 92 (Quedius; characters; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Coiffait, 1978: 191 (Quedius; subgenus Quedius; characters; Europe; Britain).

  • — Outerelo, 1978a: 281 (Quedius; pupal characters; Spain).

  • — Smetana, 1978a: 826 (Quedius; subgenus Quedius; collecting notes; Oregon).

  • — Tóth, 1984: 118 (Quedius; characters; Hungary).

  • — Drugmand, 1987a: 312 (Quedius; Belgium).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

  • — Nowosad, 1990: 145 (Quedius; collected from mole nest; notes; Poland).

  • — Borges, 1990: Table II (Quedius; Azores).

  • — Smetana, 1990d: 98 (Quedius; New Hampshire).

  • — Smetana, 1993b: 50 (Quedius; subgenus Quedius; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • — Owen, 1997a: 149 (Quedius; collecting sites; Ireland).

  • — Outerelo, Gamarra, and Salgado, 1998: 129 (Quedius; subgenus Quedius; collected from cave; Spain; Portugal).

  • — Assing, 2001: 75 (Quedius; collected from xerothermous site; Germany).

  • Distribution: Morocco, Azores, Europe, Turkey, Uzbekistan, Canada, USA.

parallelus Hatch, 1957: 216 (Quedius; subgenus Quedius; Type locality: Washington: Seattle, U[niversity of] W[ashington] Campus. Also cited from British Columbia).

  • — Korge, 1962b: 332 (Quedius; synonym of curtipennis).

  • — Smetana, 1965a: 12 (Quedius; synonym of curtipennis).

  • — Smetana, 1971: 130 (Quedius; synonym of curtipennis).

curtipennis Casey, 1915, see: uteanus Casey, 1915.

curtus Erichson, 1840: 534 (Quedius; Type locality: Sardinia).

  • — Fairmaire and Laboulbène, 1856: 535 (Quedius; characters; France).

  • — Redtenbacher, 1857: 200 (Quedius; characters; Austria).

  • — Fauvel, 1874: 274 [= 1874a: 500] (Quedius; characters; France; Italy; Sicily; Sardinia).

  • — Redtenbacher, 1874: 197 (Quedius; characters; Austria).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; Corsica).

  • — Mulsant and Rey, 1876: 623 [= 1877: 479] (Quedius; characters; France).

  • — Ganglbauer, 1895: 397 (Quedius; subgenus Microsaurus; characters; France; Italy; Sardinia; Sicily).

  • — Porta, 1907: 110 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • — Gridelli, 1924d: 17 (Quedius; subgenus Euryquedius; characters; Italy).

  • — Portevin, 1929: 337 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1437 (Quedius; catalog).

  • — Bordoni, 1973b: 43 (Quedius; subgenus Euryquedius; characters; Italy).

  • — Coiffait, 1978: 143 (Quedius; subgenus Euryquedius; characters; western Mediterranean; France; Corsica).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: France, Italy.

coeruleipennis Fauvel, 1874: 275 [= 1874a: 501] (Quedius; race of curtus; Type locality: Mont Rosa; Dijon; Hautes-Alpes; rochers de Royan; Dordogne; Nyons; Toulon; St Maximin; Provence, Languedoc; Gênes).

  • — Duvivier, 1883: 138 (Quedius; catalog; variety of curtus).

  • — Ganglbauer, 1895: 388 (Quedius; variety of curtus).

  • — Porta, 1907: 110 (Quedius; variety of curtus).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; variety of curtus).

  • — Scheerpeltz, 1933: 1438 (Quedius; variety of curtus).

  • — Bordoni, 1973b: 43 (Quedius; variety of curtus).

  • — Coiffait, 1978: 143 (Quedius; variety of curtus; characters).

cyaneipennis Scheerpeltz, 1933: 1438 (Quedius; replacement name for coeruleipennis Bernhauer).

  • Distribution: Indonesia.

coeruleipennis Bernhauer, 1926f: 319 (Quedius; [preoccupied]; Type locality: Buru).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of cyaneipennis).

cyanellus Fauvel, 1878f: 274 (Quedius; Type locality: Ternate, Acquiconora).

  • — Bernhauer and Schubert, 1916: 421 (Quedius; catalog).

  • Distribution: Indonesia.

cyaneorufus Fauvel, 1878f: 274 (Quedius; Type locality: Iles Arou, Wokan).

  • — Fauvel, 1879: 107 (Quedius; Dorei; Amberbaki).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Cameron, 1931: 363 (Quedius; New Guinea).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Cameron, 1937a: 104 (Quedius; New Guinea).

  • — Last, 1975: 439 (Quedius; subgenus Sauridus; New Guinea).

  • — Last, 1987: 54 (Quedius; Papua New Guinea; Irian).

  • Distribution: Indonesia, New Guinea.

gazellensis Last, 1987: 54 (Quedius; variety of cyaneorufus; [Note: Nomen nudum (Article 15.2)]; cited from Papua New Guinea: New Britain, Gazelle Pen., Mt. Sinewit, 900 m; Morobe Dist., Wau, 1200–1300 m; Iongai, 10 km E Mt. Albert Edward, 1450 m; Umboi I., ca. 8 km WNW Lablab, 300 m; New Ireland, ridge above “Camp Bishop”, 15 km up Kait R., 250–750 m).

cyanescens Mulsant and Rey, 1876: 727 [= 1877: 583] (Quedius; subgenus Sauridus; Type locality: environs d'Hyères).

  • — Ganglbauer, 1895: 406 (Quedius; subgenus Sauridus; characters; France).

  • — Bernhauer and Schubert, 1916: 436 (Quedius; variety of umbrinus).

  • — Gridelli, 1925: 113 (Quedius; variety of umbrinus).

  • — Scheerpeltz, 1933: 1464 (Quedius; variety of umbrinus).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; valid species; characters).

  • — Coiffait, 1963b: 423 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 237 (Quedius; subgenus Sauridus; characters; Pyrenees; Mediterranean region of western Europe).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: France.

cyanopterus Scheerpeltz, 1976, see: apicicornis Eppelsheim, 1895.

cyprinus H. Franz, 1987, see: scheerpeltzi Gridelli, 1938.

cyprusensis Last, 1955: 251 (Quedius; Type locality: Cyprus: Mount Troodus, 5600–6000 ft.).

  • — Coiffait, 1963a: 385 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1970e: 62 (Quedius; Cyprus).

  • — Coiffait, 1978: 236 (Quedius; subgenus Sauridus; characters; Cyprus; Lebanon).

  • — Bordoni, 1984: 342 (Quedius; Lebanon).

  • — Assing and Wunderle, 2001: 37 (Quedius; Cyprus).

  • Distribution: Cyprus, Lebanon.

freyi Scheerpeltz, 1956c: 1102 (Quedius; subgenus Raphirus; Type locality: Cypern: Pedhoulas, 3600 feet).

  • — Coiffait, 1978: 236 (Quedius; synonym of cyprusensis).

daibosatsunis Watanabe, 1986: 19 (Quedius; subgenus Microsaurus; Type locality: Japan: Honshu: Yamanashi Pref.: Hikawa Valley, 1400 m alt., Enzan-shi).

  • Distribution: Japan.

daksumensis Coiffait, 1982a: 279 (Quedius; subgenus Raphirus; Type locality: Cachemire: Daksum, 2400–2700 m).

  • — Smetana, 1988: 247 (Quedius; subgenus Raphirus; characters; India; Nepal).

  • — Smetana, 1992b: 4 (Quedius; subgenus Raphirus; Nepal).

  • Distribution: India, Nepal.

dalmatinus Korge, 1960, see: meridiocarpathicus Smetana, 1958.

debilis Horn, 1878: 165 (Quedius; Type locality: California; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Clear Lake, California and Vancouver).

  • — Casey, 1915: 403 (Quediellus; characters; California).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 215 (Quedius; subgenus Raphirus; lectotype designtion: Cala [California]; characters; California).

  • — Smetana, 1971a: 1848 (Quedius; subgenus Raphirus; collecting notes; California).

  • Distribution: USA.

deceptor Cameron, 1944: 14 (Quedius; subgenus Sauridus; Type locality: Darjeeling: Ghum district).

  • — Smetana, 1988: 231 (Quedius; subgenus Distichalius; characters; Uttar Pradesh; West Bengal).

  • Distribution: India.

decipiens Cameron, 1944, see: aureiventris Bernhauer, 1915.

decius Smetana, 1996: 12 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan prov., Liziping env. near Shimien, 200 km SW of Ya'an).

  • — Smetana, 1999b: 215 (Quedius; subgenus Microsaurus; notes; China).

  • Distribution: China.

declivus Fauvel, 1878: 128 [= 1878a: 48] (Quedius; Type locality: Bône).

  • — Fauvel, 1886: 58 [= 1886a: 50] (Quedius; notes; Algeria).

  • — Fauvel, 1902b: 116 (Quedius; Algeria; Tunisia).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 100 (Quedius; subgenus Raphirus; characters; Tunisia; Algeria).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Jeannel and Jarrige, 1949: 378 (Quedius; subgenus Raphirus; cave species; Algeria).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 206 (Quedius; subgenus Sauridus; characters; Algeria; Tunisia).

  • — Maachi, 1991: 251 (Quedius; Morocco).

  • Distribution: Morocco, Algeria, Tunisia.

decolor Coiffait, 1976 [Note: Nomen nudum], see: fissus Gridelli, 1938.

decoratus Petri 1902, see: haberfelneri Eppelsheim, 1891.

decoui Coiffait, 1963a: 381, 406 (Quedius; subgenus Sauridus; Type locality: Roumanie, Olténie).

  • — Coiffait, 1978: 221 (Quedius; subgenus Sauridus; characters; Romania).

  • — Tóth, 1984: 135 (Quedius; characters).

  • Distribution: Romania.

deficiens Cameron, 1945, see: quadripunctus Bernhauer, 1941.

delabiei Coiffait, 1977 [Note: Nomen nudum], see: coronatus Delabie, 1953.

delahoni H. Wagner, 1916 [Note: Nomen nudum], see: mesomelinus Marsham, 1802.

densiventris Casey, 1915: 404 (Quediellus; Type locality: Montana: Mullan).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 199 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Newfoundland; Yukon Territory; Alaska; Colorado; Idaho; New Hampshire; New Mexico; New York; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971a: 1846 (Quedius; subgenus Raphirus; collecting notes; New Mexico; California).

  • — Smetana, 1973: 1429 (Quedius; notes; Quebec).

  • — Smetana, 1976c: 181 (Quedius; subgenus Raphirus; Colorado; New Mexico).

  • — Smetana, 1976d: 26 (Quedius; characters).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; Arizona; Utah).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: USA, Canada.

alutaceus Hatch, 1957: 219 (Quedius; subgenus Distichalius; Type locality: Washington: Mt. Rainier, Sunrise Park. Also cited from British Columbia).

  • — Smetana, 1971: 199 (Quedius; synonym of densiventris).

bonapartei Hatch, 1957: 226 (Quedius; subgenus Sauridus; Type locality: Washington: Mt. Bonaparte. Also cited from Idaho, Oregon, and British Columbia).

  • — Smetana, 1965: 46 (Quedius; subgenus Raphirus; characters; Newfoundland).

  • — Smetana, 1965a: 14 (Quedius; subgenus Raphirus; British Columbia).

  • — Smetana, 1971: 199 (Quedius; synonym of densiventris).

denudatus Stephens, 1832, see: molochinus Gravenhorst, 1806.

depauperatus Wollaston, 1871: 299 (Quedius; variety of fulgidus; Type locality: Teneriffe).

  • — Wollaston, 1864: 563 (Quedius; [Note: Misidentification: Bernhauer and Schubert, 1916: 422, cited fulgidus Wollaston as a synonym of depauperatus; Wollaston did not describe the species as new, he attributed it to Fabricius]; notes; habitat; Canary Islands).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; variety of fulgidus).

  • — Fauvel, 1902b: 115s (Quedius; Canaries).

  • — Porta, 1907: 114 (Quedius; variety of fulgidus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 64 (Quedius; subgenus Microsaurus; characters; Canaries).

  • — Gridelli, 1929: 39 (Quedius; characters; Canaries).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Coiffait, 1978: 155 (Quedius; subgenus Microsaurus; characters; Canary Islands).

  • — Assing and Wunderle, 1999: 435 (Quedius; collecting notes; Canary Islands).

  • — Assing and Wunderle, 1999: 437 (Quedius; Canary Islands).

  • Distribution: Canary Islands.

robustus Wollaston, 1871: 299 (Quedius; variety of fulgidus; [preoccupied]; Type locality: Gomera).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of maurus).

  • — Eppelsheim, 1878b: 175 (Quedius; [Note: Scheerpeltz, 1933: 1438, cited robustus Eppelsheim and robustus Wollaston as synonyms of depauperatus; Eppelsheim attributed the name to Wollaston]).

  • — Ganglbauer, 1895: 400 (Quedius; variety of maurus).

  • — Porta, 1907: 125 (Quedius; synonym of maurus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of depauperatus).

  • — Gridelli, 1929: 39 (Quedius; synonym of depauperatus).

  • — Scheerpeltz, 1933: 1438 (Quedius; synonym of depauperatus).

  • — Coiffait, 1978: 155 (Quedius; synonym of depauperatus).

deperditus† Scudder, 1900a: 82 (Quedius; pleistocene fossil; Type locality: Scarborough).

  • Distribution: Canada.

deschutesi Hatch, 1957: 220 (Quedius; subgenus Microsaurus; Type locality: Oregon: Deschutes Co.: Camp Abbott).

  • — Smetana, 1971: 102 (Quedius; subgenus Microsaurus; characters; Oregon).

  • — Smetana, 1981b: 634 (Quedius; characters; collecting notes; Arizona).

  • Distribution: USA.

desertus Horn, 1878: 161 (Quedius; Type locality: Pinal Mts., Arizona; [Note: See lectotype designation by Smetana, 1971]. Originally cited from Colorado River in California; Arizona; southwestern Utah).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 412 (Microsaurus; characters; Arizona).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; subgenus Microsaurus; nominate subspecies; lectotype designation: Pinal Mts., Ariz.; characters; Arizona; Colorado; New Mexico; Utah).

  • — Smetana, 1971a: 1840 (Quedius; subgenus Microsaurus; collecting notes; New Mexico; Arizona).

  • — Smetana, 1973: 1427 (Quedius; Texas).

  • Distribution: USA.

fontinalis Casey, 1915: 414 (Microsaurus; Type locality: New Mexico: Jemez Springs).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; synonym of desertus).

pinalicus Casey, 1915: 413 (Microsaurus; Type locality: Arizona: Pinal Mts.; [Note: See lectotype designation by Smetana, 1971]).

  • — Scheerpeltz, 1933: 1458 (Quedius; catalog).

  • — Smetana, 1971: 123 (Quedius; lectotype designation; synonym of desertus).

californicus Smetana, 1971: 125 (Quedius; subgenus Microsaurus; subspecies of desertus; Type locality: California: Volcan Mt., San Diego Co.).

exulans Smetana, 1971: 125 (Quedius; subgenus Microsaurus; subspecies of desertus; Type locality: California: Niles Cñ., Alameda Co.).

  • — Smetana, 1978a: 825 (Quedius; subgenus Microsaurus; subspecies of desertus; collecting notes; California).

deuvei Coiffait, 1978d: 170 (Quedius; subgenus Sauridus; Type locality: Drannaz Dag).

  • — Coiffait, 1978: 226 (Quedius; subgenus Sauridus; characters; Anatolia).

  • — Coiffait, 1984e: 374 (Quedius; Nepal).

  • Distribution: Turkey, Nepal ?.

deuvei Coiffait, 1983, see: kashmirensis Cameron, 1944.

deuveianus Coiffait, 1983, see: kashmirensis Cameron, 1944.

dewar Smetana, 1988: 265 (Quedius; subgenus Raphirus; Type locality: Nepal: Prov. Bagmati: Mere Dara, 3200 m).

  • Distribution: Nepal.

dhaulagirensis Coiffait, 1982, see: kanyasa Smetana, 1975.

dichrous Fauvel, 1878g: 553 (Quedius; Type locality: Australie méridionale: Victoria, Melbourne).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: Australia.

diemenensis Blackburn, 1894a: 91 (Quedius; Type locality: Tasmania).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Lea, 1925: 242 (Quedius; characters; Australia).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • Distribution: Australia.

dih Smetana, 1996g: 24 (Quedius; subgenus Raphirus; Type locality: Taiwan: Ilan Hsien: Taipingshan, 1820 m).

  • Distribution: Taiwan.

discrepans Broun, 1893: 1030 (Quedius; Type locality: Wellington).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: New Zealand.

dispersepunctatus Scheerpeltz, 1965: 293 (Quedius; subgenus Indoquedius; Type locality: N.E. Burma, Kambaiti, 7000 ft.).

  • Distribution: Myanmar.

distans Smetana, 1971: 91 (Quedius; subgenus Microsaurus; Type locality: California: Haypress Mdws., Eldorado Co. Also cited from Nevada).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Microsaurus; collecting notes).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; California).

  • — Smetana, 1973: 1426 (Quedius; Nevada).

  • — Smetana, 1976c: 170 (Quedius; subgenus Microsaurus; California).

  • Distribution: USA.

distincticolor Roubal, 1914 [Note: Nomen nudum], see: minor Hochhuth, 1849.

distinctus Runde, 1835, see: fulgidus Fabricius, 1793.

divergens Casey, 1915, see: utenaus Casey, 1915.

diversicollis Cameron, 1945d: 789 (Quedius; subgenus Sauridus; Type locality: New Zealand: Reefton).

  • Distribution: New Zealand.

diversipennis Fauvel, 1878g: 554 (Quedius; Type locality: Australie occidentale, Swan River).

  • — Bernhauer, 1908: 19 (Quedius; Australia).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Bernhauer, 1916a: 7 (Quedius; Australia).

  • Distribution: Australia.

doderoi Gridelli, 1922: 22 (Quedius; subgenus Microsaurus; Type locality: Monte Misurasca [Apennino ligure]).

  • — Gridelli, 1924d: 32 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Bordoni, 1974: 18 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Coiffait, 1978: 182 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Italy.

dohertyi Cameron, 1932, see: muscicola Cameron, 1932.

dolosus Coiffait, 1961 [Note: Nomen nudum], see: unicolor Kiesenwetter, 1847.

dorsalis Stephens, 1835: 436 (Quedius; [Note: Nomen dubium]; Type locality: Coombe Wood).

  • Distribution: Britain.

dorni Korge, 1969, see: ochropterus Erichson, 1840.

douglasi Bernhauer, 1929: 2 (Quedius; Type locality: Nord-China: Shensi, Yung-ting-Shan Mts, 90 m. W of Taiyan-fu, 7000 ft.).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog).

  • — Smetana, 1996f: 113 (Quedius; subgenus Microsaurus; characters; China).

  • Distribution: China.

downiei Hatch, 1957, see: montanicus Casey, 1915.

drannazensis Coiffait, 1978d: 171 (Quedius; subgenus Sauridus; Type locality: Drannaz Dag).

  • — Coiffait, 1978: 252 (Quedius; subgenus Sauridus; characters; Anatolia).

  • Distribution: Turkey.

dryadum Peyerimhoff, 1917: 121 (Quedius; subgenus Microsaurus; Type locality: Boucharen, au sud de Larache, Maroc occidental; Massif des Mouzaia, Alger, 1200 m).

  • — Gridelli, 1924d: 46 (Quedius; subgenus Microsaurus; characters; Algeria).

  • — Gridelli, 1925: 170 (Quedius; characters).

  • — Scheerpeltz, 1933: 1438 (Quedius; catalog; Mauritania).

  • — Coiffait, 1978: 172 (Quedius; subgenus Microsaurus; characters; North Africa).

  • Distribution: Mauritania, Morocco, Algeria.

dubius Heer, 1839: 278 (Philonthus; variety b of montanus; [Note: Preoccupied as a secondary homonym in Philonthus; Heer presented characters for montanus variety b; following those characters he equated the variety with Staphylinus dubius Heer. I take the characterization and link established between variety b and dubius to be sufficient to validate the name which Heer first published in 1834 as an unavailable name]; Type locality: Paradies).

  • — Heer, 1834a: 156 (Staphylinus; subgenus Microsaurus; [Note: Unavailable name]; elevational listing).

  • — Gemminger and Harold, 1868: 569 (Quedius; synonym of fimbriatus).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; characters; France; Switzerland; Italy; Austria).

  • — Mulsant and Rey, 1876: 719 [= 1877: 575] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; subgenus Sauridus; characters; Germany; Alps; Carpathians).

  • — Porta, 1907: 135 (Quedius; subgenus Sauridus; characters; Italy).

  • — Reitter, 1909: 112 (Quedius; subgenus Sauridus; characters; Germany).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 98 (Quedius; subgenus Raphirus; characters; Italy).

  • — Portevin, 1929: 342 (Quedius; characters; France).

  • — Scheerpeltz, 1933: 1438 (Quedius; [Note: Scheerpeltz, 1933: 1439 cited rubromaculatus Gridelli, 1924: 99 as an aberration of dubius. Gridelli, in the reference cited by Scheerpeltz, described an aberration but did not propose a new name. Gridelli began the description with “ab. elytris rubromaculatis” and he did not cite the name in the index to his article]; catalog).

  • — Wüsthoff, 1938a: fig. 25 (Quedius; aedeagal illustration).

  • — Smetana, 1962b: 140 (Quedius; subgenus Raphirus; characters).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; characters).

  • — Lohse, 1964: 215 (Quedius; subgenus Raphirus; characters; central Europe).

  • — Horion, 1965: 277 (Quedius; subgenus Raphirus; Germany; Austria; Switzerland; Italy; France; Slovenia; Croatia; Bosnia Hercegovina; Czechoslovakia; Poland; Romania).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; characters; Alps; Apennines).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 131 (Quedius; characters).

  • — Lucht, 1987: 109 (Quedius; nominate subspecies; checklist; Germany; Austria; Switzerland; France).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; Italy).

  • Distribution: Europe.

montanus Heer, 1839: 277 (Philonthus; Type locality: Matt; alle Glarneralpen; Paradies an der Zaportalp; Urserenthal; Furca; am Mt. Rosa).

  • — Küster, 1853a: 63 (Quedius; characters; Switzerland).

  • — Kraatz, 1857d: 506 (Quedius; synonym of fimbriatus).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; synonym of dubius).

  • — Mulsant and Rey, 1876: 721 [= 1877: 577] (Quedius; subgenus Sauridus; characters; France).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; subgenus Raphirus; variety of dubius; characters; Italy).

  • — Scheerpeltz, 1933: 1439 (Quedius; variety of dubius).

  • — Wüsthoff, 1938a: fig. 26 (Quedius; form of dubius; aedeagal illustration).

  • — Smetana, 1958: 366 (Quedius; synonym of dubius fimbriatus).

  • — Smetana, 1962b: 141 (Quedius; aberration of dubius).

  • — Lohse, 1964: 214 (Quedius; subgenus Raphirus; aberration of dubius; characters; central Europe).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; variety of dubius; characters).

fimbriatus Erichson, 1840: 539 (Quedius; Type locality: Austria).

  • — Märkel and Kiesenwetter, 1848a: 325 (Quedius; notes; Carpathian Alps).

  • — Redtenbacher, 1849: 710 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; France).

  • — Fairmaire and Laboulbène, 1856: 544 (Quedius; characters; France).

  • — Kraatz, 1857d: 506 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 203 (Quedius; characters; Austria).

  • — Fauvel, 1874: 293 [= 1874a: 519] (Quedius; synonym of dubius).

  • — Redtenbacher, 1874: 200 (Quedius; characters; Austria).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; subspecies of dubius).

  • — Scheerpeltz, 1933: 1439 (Quedius; subspecies of dubius).

  • — Wüsthoff, 1938a: fig. 27 (Quedius; form of dubius; aedeagal illustration).

  • — Smetana, 1958: 366 (Quedius; subgenus Raphirus; subspecies of dubius; characters; ecology; Czechoslovakia).

  • — Smetana, 1962b: 141 (Quedius; subgenus Raphirus; subspecies of dubius; characters).

  • — Coiffait, 1963a: 375 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; variety of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; subspecies of dubius; characters).

  • — Lucht, 1987: 110 (Quedius; subspecies of dubius; checklist; Czechoslovakia; Austria).

  • — Smetana, 1993b: 50 (Quedius; subgenus Raphirus; subspecies of dubius; Czech Republic).

  • — Ciceroni and Zanetti, 1995: 32 (Quedius; subspecies of dubius; Italy).

speculator Kiesenwetter, 1848: 57 (Quedius; Type locality: Kärthner Alpen).

  • — Kraatz, 1857d: 506 (Quedius; synonym of fimbriatus).

  • — Ganglbauer, 1895: 405 (Quedius; synonym of dubius).

  • — Porta, 1907: 135 (Quedius; synonym of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; synonym of dubius).

  • — Gridelli, 1924d: 98 (Quedius; synonym of dubius fimbriatus).

  • — Scheerpeltz, 1933: 1439 (Quedius; synonym of dubius).

  • — Smetana, 1958: 366 (Quedius; synonym of dubius fimbriatus).

flavolineatus Bernhauer, 1899a: 433 (Quedius; variety of dubius; Type locality: Schweiz).

  • — Porta, 1907: 135 (Quedius; variety of dubius).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; variety of dubius).

  • — Gridelli, 1924d: 98 (Quedius; synonym of dubius variety montanus).

  • — Scheerpeltz, 1933: 1439 (Quedius; synonym of dubius).

  • — Smetana, 1962b: 141 (Quedius; synonym of dubius).

ermischi Korge, 1969: 44 (Quedius; subspecies of dubius; Type locality: Südbulgarien: Musalla, Rhodope-Gebirge, 1600–2400 m).

  • — Coiffait, 1978: 207 (Quedius; subgenus Sauridus; subspecies of dubius; characters).

  • — Tóth, 1984: 132 (Quedius; subspecies of dubius; characters).

dui Smetana, 1988: 204 (Quedius; subgenus Microsaurus; Type locality: Punjab: Kulu Div.).

  • Distribution: India.

duplepunctatus Gridelli, 1922: 21 (Quedius; subgenus Sauridus; Type locality: Francia, Pirenei orient.: Montlouis).

  • — Gridelli, 1925: 138 (Quedius; subgenus Raphirus; probably identical to muscorum; characters; Pyrenees).

  • — Scheerpeltz, 1933: 1450 (Quedius; synonym of muscorum).

  • — Coiffait, 1963a: 381 (Quedius; subgenus Sauridus; characters).

  • — Coiffait, 1978: 226 (Quedius; subgenus Sauridus; cited as valid species; characters; Pyrenees).

  • Distribution: France.

duplex Fauvel, 1886, see: iridicolor Quedenfeldt, 1882.

duplopunctatus Lea, 1925, see: leai Scheerpeltz, 1933.

durgaa Smetana, 1988: 273 (Quedius; subgenus Raphirus; Type locality: India: W. Bengal, Darjeeling dist., 13 km N Ghoom, 1500).

  • Distribution: India.

dzambulensis Coiffait, 1967e: 403 (Quedius; subgenus Sauridus; Type locality: Aulie-Ata, Dzambul, Kazakstan).

  • — Coiffait, 1978: 229 (Quedius; subgenus Sauridus; characters; Turkestan).

  • — Boháč, 1988: 556 [= 1989: 40] (Quedius; characters; Uzbekistan; Turkestan).

  • Distribution: Kazakhstan, Uzbekistan.

echion Smetana, 1997: 65 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Langmusi, 3500 m).

  • — Smetana, 1999b: 217 (Quedius; subgenus Microsaurus; notes on holotype; characters of male; China).

  • Distribution: China.

edmundi Coiffait, 1969: 54 (Quedius; subgenus Microsaurus; replacement name for reitterianus Coiffait).

  • — Coiffait, 1978: 164 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • Distribution: Caucasus.

microphthalmus Reitter, 1909: 109 (Quedius; subgenus Ediquus; [preoccupied]; Type locality: Kaukasus: Meskisches Gebirge).

  • — Gridelli, 1924d: 71 (Quedius; subgenus Microsaurus; characters; Caucasus).

  • — Coiffait, 1967e: 394 (Quedius; synonym of reitterianus).

  • — Coiffait, 1978: 164 (Quedius; synonym of edmundi).

reitterianus Coiffait, 1967e: 394 (Quedius; subgenus Microsaurus; replacement name for microphthalmus Reitter; [preoccupied]).

  • — Coiffait, 1978: 164 (Quedius; synonym of edmundi).

edwardsi Bernhauer, 1935, see: edwarsianus Korge, 1963.

edwardsi Sharp, 1886a: 378 (Quedius; Type locality: New Zealand: Picton).

  • — Broun, 1893: 1029 (Quedius; characters; New Zealand).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: New Zealand.

edwarsianus Korge, 1963a: 84 (Quedius; replacement name for edwardsi, cited as edwarsi for both the Sharp and Bernhauer names).

  • — Coiffait and Saiz, 1966: 412 (Quedius; characters; Chile).

  • Distribution: Chile.

edwardsi Bernhauer, 1935c: 92 (Quedius; [preoccupied]; Type locality: Süd-Chile: Llanquihue Prov., Casa Pangue).

  • — Korge, 1963a: 84 (Quedius; cited as edwarsi; synonym of edwarsianus).

  • — Coiffait and Saiz, 1966: 413 (Quedius; synonym of edwardsianus).

ehecatl Smetana, 1976a: 240 (Quedius; subgenus Quedionuchus; Type locality: Mexico: Oaxaca: 4 mi. W. jct. Mex. 175 Yuvila Turnoff, 9300 ft.).

  • Distribution: Mexico.

eklai Smetana, 1988: 257 (Quedius; subgenus Raphirus; Type locality: Nepal: 172 Parbat Dist., zwischen Chitre und Ghandrung, Chitre-Seite des Passes, 2800–2900 m).

  • Distribution: Nepal.

elegans Ericson, 1897, see: brevicornis Thomson, 1860.

elevatus Hatch, 1957: 228 (Quedius; subgenus Sauridus; Type locality: Washington: Snoqualmie. Also cited from Oregon and British Columbia).

  • — Smetana, 1971: 224 (Quedius; subgenus Raphirus; characters; British Columbia; California; Nevada; Oregon; Washington).

  • — Smetana, 1971a: 1848 (Quedius; subgenus Raphirus; collecting notes; California).

  • Distribution: Canada, USA.

emei Smetana, 1997: 67 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Emei Shan, 3000 m, 29°32′N, 103°21′E).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; characters of female; China).

  • Distribution: China.

ennius Smetana, 1996: 16 (Quedius; subgenus Microsaurus; Type locality: China: Yunnan prov., Heishui 35 km N. Lijiang, 27°13′N, 100°19′E).

  • Distribution: China.

ephialtes Smetana, 1997: 60 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, abv. Camp 3, 3300–3350 m).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; collecting notes; China).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; notes; China).

  • Distribution: China.

eppelsheimi Quedenfeldt, 1882, see: plancus Erichson, 1840.

eppelsheimii Seidlitz, 1891, see: brevicornis Thomson, 1860.

epytus Smetana, 1995f: 236 (Quedius; subgenus Microsaurus; Type locality: China: Gansu Mts., 25 km E. Xiahe, 3000 m).

  • Distribution: China.

erdciyasicus Korge, 1971, see: limbatus Heer, 1839.

erinci Korge, 1971, see: nemoralis Baudi, 1848.

ermischi Korge, 1969, see: dubius Heer, 1839.

ernestini Fauvel, 1872, see: praecox Gravenhorst, 1802.

erratus Coiffait, 1977 [Note: Nomen nudum], see: fissus Gridelli, 1938.

eruensis Broun, 1912: 398 (Quedius; Type locality: Erua).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • Distribution: New Zealand.

erythras Smetana, 1997: 58 (Quedius; subgenus Microsaurus; Type locality: China: Gansu Mts., 25 km E Xiahe, 3000 m).

  • — Smetana, 1999b: 216 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

erythroderes Lea, 1925: 247 (Quedius; Type locality: South Australia: Mount Lofty, Wallaroo. Victoria: Forrest).

  • — Scheerpeltz, 1933: 1439 (Quedius; cited as erythroderus; catalog).

  • — Oke, 1933: 123, 124 (Quedius; tarsal characters; Australia).

  • Distribution: Australia.

erythrogaster Mannerheim, 1852: 314 (Quedius; Type locality: insula Sitkha; [Note: See lectotype designation by Smetana, 1971]).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Horn, 1878: 159 (Quedius; variety of fulgidus; characters).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Casey, 1915: 410 (Microsaurus; characters; Queen Charolotte Island; California; New York).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1929: 24 (Quedius; characters).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Hatch, 1957: 222 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 80 (Quedius; subgenus Microsaurus; lectotype designation; characters; Alberta; British Columbia; Quebec; Alabama; Arkansas; California; Connecticut; Georgia; Illinois; Kentucky; Maine; Michigan; Nevada; New York; Ohio; Oregon; Pennsylvania; Tennessee; Virginia; Washington; West Virginia).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; Alaska).

  • — Smetana, 1973: 1426 (Quedius; Quebec).

  • — Smetana, 1976c: 169 (Quedius; subgenus Microsaurus; Ontario; Quebec).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; New York; Oklahoma).

  • — Smetana, 1981b: 634 (Quedius; Saskatchewan).

  • — Dearborn and Donahue, 1994: 15 (Quedius; Maine).

  • — Downie and Arnett, 1996: 390 (Quedius; subgenus Microsaurus; characters; USA).

  • Distribution: Canada, USA.

neotomae Hatch, 1957: 224 (Quedius; subgenus Microsaurus; Type locality: Oregon: Medford. Also cited from Washington).

  • — Graham, 1963: 37 (Quedius; collected from cave; California).

  • — Smetana, 1971: 80 (Quedius; synonym of erythrogaster).

erythropterus Stephens, 1832, see: cruentus Olivier, 1795.

erythropygus Gridelli, 1929, see: fulgidus Fabricius, 1793.

etruscus Mulsant and Rey, 1876, see: scintillans Gravenhorst, 1806.

euander Smetana, 1997: 63 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, above Camp 3, 3300–3350 m).

  • — Smetana, 1997d: 454 (Quedius; subgenus Microsaurus; characters; collecting notes; China).

  • — Smetana, 1999b: 217 (Quedius; subgenus Microsaurus; China).

  • Distribution: China.

eumerus Smetana, 1971: 182 (Quedius; subgenus Distichalius; Type locality: Oregon: Mt. Hood. Also cited from Washington).

  • — Smetana, 1976c: 178 (Quedius; subgenus Distichalius; collecting notes; characters; Oregon; Washington).

  • Distribution: USA.

euryalus Smetana, 1997: 52 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan, Gongga Shan, Lake abv. Camp 2, 2750 m).

  • — Smetana, 1997d: 453 (Quedius; subgenus Microsaurus; collecting notes; China).

  • Distribution: China.

excessus Last, 1975 [Note: Nomen nudum], see: splendidus Fauvel, 1878.

expectatus Israelson, 1985: 145 (Quedius; subgenus Microsaurus; [Note: Described for some specimens misidentified as angustifrons by Wollaston, 1864: 563 and 1865: 486 and by Coiffait(see following citations)]; Type locality: Gomera: Bosque del Cedro).

  • — Coiffait, 1954: 167 (Quedius; subgenus Microsaurus; [Note: Misidentified as angustifrons according to Israelson, 1985: 145]; Canary Islands).

  • — Coiffait, 1978: 175 (Quedius; subgenus Microsaurus; [Note: Misidentified as angustifrons according to Israelson, 1985: 145]; characters; Canary Islands).

  • — Assing, 1999a: 346 (Quedius; Canary Islands).

  • Distribution: Canary Islands.

explanatus LeConte, 1858: 61 (Quedius; Type locality: San Diego, California; [Note: See lectotype designation by Smetana, 1971]).

  • — Horn, 1878: 157 (Quedius; characters; California; Montana).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Casey, 1915: 423 (Megaquedius; characters; California).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1932: 15 (Quedius; subgenus Microsaurus; characters; California; Montana).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Hatch, 1957: 230 (Quedius; subgenus Megaquedius; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1971: 42 (Quedius; subgenus Megaquedius; lectotype designation; characters; British Columbia; Arizona; California; Colorado; Idaho; kansas; Missouri; Montana; New Mexico; Oregon; Utah; Washington; Wyoming).

  • — Smetana, 1971a: 1839 (Quedius; subgenus Megaquedius; collecting notes; New Mexico; Arizona).

  • — Smetana, 1978a: 823 (Quedius; subgenus Megaquedius; collecting notes; Kansas; Nevada).

  • Distribution: USA, Canada.

laxatus Casey, 1915: 422 (Megaquedius; Type locality: California, probably northern).

  • — Scheerpeltz, 1933: 1446 (Quedius; catalog).

  • — Smetana, 1971: 43 (Quedius; synonym of explanatus).

exulans Smetana, 1971, see: desertus Horn, 1878.

faang Smetana, 1999e: 538 (Quedius; subgenus Microsaurus; Type locality: China: Sichuan Pv., Meigu Xian, Wahei, Dafengding, 2320 m).

  • Distribution: China.

fagelianus Coiffait, 1967e: 391 (Quedius; subgenus Microsaurus; Type locality: Safa, Liban).

  • — Coiffait, 1978: 180 (Quedius; subgenus Microsaurus; characters; Lebanon).

  • — Smetana, 1995a: 80 (Quedius; subgenus Distichalius; notes; Lebanon).

  • Distribution: Israel, Lebanon.

loebli Smetana, 1978: 121 (Quedius; subgenus Distichalius; Type locality: Israel: Galilee au-dessus Safad, 500 m).

  • — Smetana, 1995a: 80 (Quedius; synonym of fagelianus).

fageti Thomson, 1867, see: maurus C. Sahlberg, 1830.

fallaciosus Kraatz, 1862, see: nitipennis Stephens, 1833.

fangi Smetana, 1995: 80 (Quedius; subgenus Raphirus; Type locality: Taiwan: Hualien Hsien: Taroko N. P., Nanhushi Hut, 2200 m).

  • Distribution: Taiwan.

farkaci Smetana, 1997d: 464 (Quedius; subgenus Microsaurus; Type locality: China: N. Yunnan, Xue Shan, nr. Zhongdian, 4000-4100 m, 27°49 N, 99°34 E).

  • Distribution: China.

fasciculatus Eppelsheim, 1886: 39 (Quedius; Type locality: Nicolajewsk; Chabarofka).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Gridelli, 1924d: 25 (Quedius; subgenus Microsaurus; characters; Amur region).

  • — Scheerpeltz, 1933: 1439 (Quedius; catalog).

  • — Coiffait, 1978: 159 (Quedius; subgenus Microsaurus; characters; Siberia).

  • — Smetana, 1978b: 84 (Quedius; subgenus Microsaurus; characters; Siberia).

  • Distribution: Russia.

klimai Roubal, 1914a: 164 (Quedius; Type locality: Irkutsk: Narcinck).

  • — Bernhauer and Schubert, 1916: 425 (Quedius; catalog).

  • — Gridelli, 1924d: 73 (Quedius; subgenus Microsaurus; characters; Siberia).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of fasciculatus).

  • — Coiffait, 1978: 159 (Quedius; synonym of fasciculatus).

feihuensis Smetana, 1999d: 530 (Quedius; subgenus Microsaurus; Type locality: China: Hunan Pv., Longshan X., Huoyan, Feihu Dong Cave, 260 m).

  • Distribution: China.

fellmani Zetterstedt, 1838: 62 (Staphylinus; Type locality: Lapponia Umensis: Wilhelmina).

  • — Erichson, 1840: 547 (Quedius; synonym of hyperboreus).

  • — Thomson, 1860: 179 (Raphirus; synonym of attenuatus).

  • — Fauvel, 1869: 486 (Quedius; synonym of attenuatus).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — A. Strand, 1937: 110 (Quedius; characters; notes).

  • — Renkonen, 1939: 65 (Quedius; notes; Finland).

  • — Smetana, 1963c: 300 (Quedius; subgenus Raphirus; Mongolia).

  • — Palm, 1963b: 153 (Quedius; subgenus Raphirus; characters; Sweden; Norway; Finland).

  • — Smetana, 1964: 39 (Quedius; subgenus Raphirus; Alaska).

  • — Smetana, 1965: 47 (Quedius; subgenus Raphirus; characters; Labrador; Newfoundland).

  • — Smetana, 1965a: 16 (Quedius; subgenus Raphirus; notes; Alaska).

  • — Smetana, 1967d: 217 (Quedius; subgenus Raphirus; Mongolia).

  • — Smetana, 1995a: 86 (Quedius; subgenus Raphirus; notes; Russia).

  • — Smetana, 1971: 233 (Quedius; subgenus Raphirus; characters; Alberta; British Columbia; Labrador; Manitoba; Newfoundland; Northwest Territories; Quebec; Yukon Territory; Alaska; Colorado; Idaho; New Hampshire).

  • — Smetana, 1973: 1431 (Quedius; notes; Northwest Territories; Alaska).

  • — Smetana 1975c: 176 (Quedius; Mongolia).

  • — Smetana, 1976c: 183 (Quedius; subgenus Raphirus; Colorado).

  • — Smetana, 1976d: 28 (Quedius; subgenus Raphirus; characters; Siberia).

  • — Coiffait, 1978: 268 (Quedius; subgenus Raphirus; characters; Scandinavia).

  • — Smetana, 1978b: 86 (Quedius; subgenus Raphirus; Siberia).

  • — Smetana, 1981b: 643 (Quedius; collecting notes; Manitoba; Saskatchewan).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • — Ryabukhin, 1999: 74 (Quedius; subgenus Raphirus; catalog; habitat; distribution; northeastern Russia).

  • Distribution: Europe, Russia, Mongolia, Canada, USA.

hyperboreus Erichson, 1840: 547 (Quedius; Type locality: Unalaschka; [Note: See lectotype designation by Smetana, 1971]).

  • — Mannerheim, 1843b: 233 (Quedius; characters; Unalaska).

  • — Fauvel, 1874: 306 [= 1874a: 532] (Quedius; synonym of attenuatus).

  • — Porta, 1907: 151 (Quedius; synonym of fulvicollis).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulvicollis).

  • — Notman, 1919a: 95 (Quedius; Ontario).

  • — Gridelli, 1925: 161 (Quedius; synonym of attenuatus).

  • — Fall, 1926: 150 (Quedius; tentative identification; Alaska).

  • — Scheerpeltz, 1933: 1432 (Quedius; synonym of attenuatus).

  • — Smetana, 1958: 393 (Quedius; synonym of nitipennis).

  • — Smetana, 1971: 233 (Quedius; lectotype designation; synonym of fellmani).

  • — Coiffait, 1978: 270 (Quedius; [Note: I assume Coiffait overlooked the change of synonymy by Smetana, 1971: 233]; synonym of nitipennis).

femoralis Sharp, 1884: 338 (Quedionuchus; Type locality: Guatemala, Totonicapam, 8500 to 10,500 feet; [Note: See lectotype designation by Smetana, 1976a]. Also cited from Quiche Mountains 7000 to 9000 feet).

  • — Bernhauer and Schubert, 1916: 422 (Quedius; catalog).

  • — Smetana, 1976a: 234 (Quedius; subgenus Quedionuchus; lectotype designation: Totonicapam, 8500–10,500 ft., Guatemala; characters; Guatemala).

  • Distribution: Guatemala.

fen Smetana, 1996c: 229 (Quedius; subgenus Raphirus; Type locality: China: Sichuan: Emei Shan).

  • Distribution: China.

fenderi Hatch, 1957: 228 (Quedius; subgenus Sauridus; Type locality: Oregon: Lake of Woods, Ashland Rd.).

  • — Smetana, 1971: 178 (Quedius; subgenus Distichalius; characters; Oregon).

  • — Smetana, 1978a: 828 (Quedius; subgenus Distichalius; collecting notes; Oregon).

  • Distribution: USA.

fenestratus Scholz, 1928 [Note: Nomen nudum], see: paradisianus Heer, 1839.

ferox Blackburn, 1888, see: tepperi Blackburn, 1888.

ferox LeConte, 1878, see: Hemiquedius.

filicornis Eppelsheim, 1895, see: Indoquedius.

fimbriatus Erichson, 1840, see: dubius Heer, 1839.

fiorii Korge, 1961, see: mesomelinus Marsham, 1802.

fissus Gridelli, 1938: 13 (Quedius; subgenus Microsaurus; Type locality: Asia Minore meridionale: Monti Salbak nel “Karisch. Taurus. Isola di Lesbo. Asia Minore, a Smirne).

  • — Coiffait, 1970e: 62 (Quedius; Lesbos).

  • — Coiffait, 1976b: 90 (Quedius; subgenus Microsarus; Rhodes).

  • — Bordoni, 1976: 236 (Quedius; characters; notes; Turkey).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; characters; Turkey; Rhodes; Lesbos).

  • Distribution: Turkey, Greece.

decolor Coiffait, 1976b: 90 (Quedius; subgenus Microsarus; variety of fissus; [Note: Nomen nudum (Article 15.2)]; cited from Rhodes: Mont Profitis Ilias).

  • — Coiffait, 1978: 178 (Quedius; subgenus Microsaurus; variety of fissus; characters).

erratus Coiffait, 1977: 136 (Quedius; subgenus Microsaurus; variety of fissus; [Note: Nomen nudum (Article 15.2)]; cited from Rhodes).

  • — Coiffait, 1978: 178 (Quedius; synonym of fissus).

flavescens Linné, 1758, see: cinctus Paykull, 1790.

flavicaudus Sharp, 1884: 333 (Quedius; Type locality: Panama, Volcan de Chiriqui between 2000 and 4000 feet, Bugaba).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog; Bolivia; Argentina).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1955a: 177 (Quedius; subgenus Cyrtoquedius; characters in key; Panama).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • Distribution: Panama, Bolivia, Argentina.

flavicornis Sharp, 1889: 30 (Quedius; Type locality: Ishiuchi; Hitoyoshi).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Nakane, 1963a: 94 (Quedius; characters; habitus photograph; Japan).

  • — Smetana, 1995: 55 (Quedius; subgenus of Microsaurus; characters; bionomics; Taiwan).

  • Distribution: Japan, Taiwan.

flavinasus Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius; Type locality: Iatahy, Goyaz).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 178 (Quedius; subgenus Cyrtoquedius; characters in key; Argentina; Paraguay; Brazil).

  • Distribution: Brazil, Paraguay, Argentina.

nigrifrons Bernhauer, 1934b: 217 (Quedius; variety of rufinasus; [Note: Cited as rufinasus Bernhauer but that species was described by Sharp. Bernhauer probably intended flavinasus Bernhauer]; Type locality: Argentinien: Misiones: Loreto).

  • — Scheerpeltz, 1955a: 176 (Quedius; subgenus Cyrtoquedius; variety of flavinasus; characters in key; Argentina).

flavipennis Baudi, 1848, see: virgulatus Erichson, 1840.

flavocaudatus Cameron, 1944: 13 (Quedius; subgenus Microsaurus; Type locality: United Provinces: Naini Tal, altitude 6000–8000 feet).

  • — Smetana, 1988: 200 (Quedius; subgenus Microsaurus; characters; Uttar Pradesh; Nepal).

  • Distribution: India, Nepal.

flavocaudatus Hayashi, 1992, see: noboruitoi Hayashi, 1992.

flavolineatus Bernhauer, 1899, see: dubius Heer, 1839.

flavomarginatus Gridelli, 1925, see: fumatus Stephens, 1833.

flavopilus Gerhardt, 1899, see: paradisianus Heer, 1839.

flavopterus Geoffroy, 1785, see: plagiatus Mannerheim, 1843.

flavoterminatus Cameron, 1932, see: apicicornis Eppelsheim, 1895.

floralis Lacordaire, 1835, see: ochripennis Ménétriés, 1832.

fluviatilis Cameron, 1926b: 369 (Quedius; subgenus Raphirus; Type locality: Chakrata Dist., Dodora Khud, 8000′; [Note: See lectotype designation by Smetana, 1988]. Originally cited from Chakrata district, 6000 to 9300 feet).

  • — Cameron, 1932: 297 (Quedius; subgenus Raphirus; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Smetana, 1988: 239 (Quedius; subgenus Raphirus; lectotype designation: Chakrata Dist., Dodora Khud, 8000′; characters; Kashmir; Himachal Pradesh; Uttar Pradesh).

  • Distribution: India.

kashmiricus Cameron, 1932: 296 (Quedius; subgenus Raphirus; Type locality: Kashmir: Gulmarg, alt. 8000 feet; [Note: See lectotype designation by Smetana, 1988]).

  • — Scheerpeltz, 1933: 1445 (Quedius; catalog).

  • — Smetana, 1988: 239 (Quedius; lectotype designation; synonym of fluviatilis).

cachemiricus Coiffait, 1982a: 277 (Quedius; subgenus Sauridus; Type locality: Cachemire: Sonamarg, 2600–2750 m).

  • — Smetana, 1988: 239 (Quedius; synonym of fluviatilis).

nichinaiensis Coiffait, 1982: 79 (Quedius; subgenus Sauridus; Type locality: Inde: Cachemire, Sonamarg, Himalaya-Hauptkette, Nichinai Tal, 3100–3200 m).

  • — Smetana, 1988: 239 (Quedius; synonym of fluviatilis).

fodinarum Uéno and Watanabe, 1966: 323 (Quedius; subgenus Microsaurus; Type locality: Japan: central Kyushu: Oita Prefecture: an abandoned mine called “Otani-ko”, at Kiura-Kozan in Onoichi of Umé-mura).

  • — Watanabe, 1996a: 13 (Quedius; mine collection; Japan).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • Distribution: Japan.

fonteius Smetana, 1995f: 247 (Quedius; subgenus Microsaurus; Type locality: China E Quinghai prov. pass 30 km S of Huizo, 3600 m).

  • Distribution: China.

fontinalis Casey, 1915, see: desertus Horn, 1878.

formosae Cameron, 1949, see: Indoquedius.

forsteri Scheerpeltz, 1960c: 129 (Quedius; subgenus Microsaurus; Type locality: Bolivien: Songtal, St. Rosa, 2800 m).

  • Distribution: Bolivia.

franzi Smetana, 1975a: 326 (Quedius; subgenus Microsaurus; Type locality: Central Nepal: Trail from Gosaikunde to Fulung monastery).

  • — Smetana, 1988: 217 (Quedius; subgenus Microsaurus; characters; Nepal).

  • Distribution: Nepal, India.

trisulensis Coiffait, 1982: 79 (Quedius; subgenus Microsaurus; Type locality: Nepal: Trisuli Tal, Gosainkund, 3200 m).

  • — Smetana, 1988: 217 (Quedius; synonym of franzi).

najik Smetana, 1992b: 4 (Quedius; subgenus Microsaurus; subspecies of franzi; Type locality: India: Sikkim, “Dzongri, 3970 m, Kangchendzonga Area”).

frater Smetana, 1971: 182 (Quedius; subgenus Distichalius; Type locality: Washington: Olympic Hot Springs).

  • — Smetana, 1976c: 178 (Quedius; subgenus Distichalius; Washington).

  • — Smetana, 1978a: 829 (Quedius; subgenus Distichalius; collecting notes; Alberta; British Columbia).

  • Distribution: Canada, USA.

fraternus Bernhauer, 1914, see: suturalis Kiesenwetter, 1845.

fraudulentum Coiffait, 1967 [Note: Nomen nudum], see: brevis Erichson, 1840.

frenatus Erichson, 1840: 533 (Quedius; Type locality: La Guayram, Columbia).

  • — Fauvel, 1891: 119 (Quedius; Venezuela).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Bernhauer, 1917h: 93 (Quedius; subgenus Cyrtoquedius).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Scheerpeltz, 1955a: 179 (Quedius; subgenus Cyrtoquedius; characters in key; Venezuela; Ecuador; Colombia).

  • Distribution: Venezuela, Colombia, Ecuador.

freyanus Smetana, 1999: 79 (Quedius; subgenus Raphirus; replacement name for freyi Smetana).

  • Distribution: China.

freyi Smetana, 1998b: 108 (Quedius; subgenus Raphirus; [preoccupied]. Type locality: China: Lungai, 2000 m, Wassuland, W. Szechuan, Sankiangkou).

  • — Smetana, 1999: 79 (Quedius; synonym of freyanus).

  • Distribution: China.

freyi Scheerpeltz, 1956, see: cyprusensis Last, 1955.

freyi Smetana, 1998, see: freyanus Smetana, 1999.

frigidus Smetana, 1971: 230 (Quedius; subgenus Raphirus; Type locality: New York: Mt. Whiteface. Also cited from Maine, New Hampshire, and Alberta).

  • — Smetana, 1973: 1431 (Quedius; notes; Newfoundland; Northwest Territories).

  • — Smetana, 1976c: 183 (Quedius; subgenus

  • — Smetana, 1976c: 183 (Quedius; subgenus Raphirus; collecting notes; Ontario).

  • — Smetana, 1978a: 830 (Quedius; subgenus Raphirus; collecting notes; British Columbia; Newfoundland; New Hampshire).

  • — Smetana, 1981b: 642 (Quedius; collecting notes; Alaska).

  • — Smetana, 1990d: 101 (Quedius; Alberta; New Hampshire).

  • — Downie and Arnett, 1996: 393 (Quedius; subgenus Raphirus; characters; USA).

  • Distribution: Canada, USA.

frontalis Nordmann, 1837, see: levicollis Brullé, 1832.

fuchsii Scriba, 1866, see: mesomelinus Marsham, 1802.

fulgidus Fabricius, 1793: 525 (Staphylinus; [preoccupied]; [Note: Nearly all writers (see for example Erichson, 1839a: 486, 1840: 525; Bernhauer and Schubert, 1916: 423; Smetana, 1958: 349; Smetana, 1971: 66; Coiffait, 1978: 146) have cited Fabricius, 1787: 220 as the original reference for Quedius fulgidus. Tottenham (1939d: 237) in his discussion of fulgidus Fabricius, 1787: 220, and fulgidus Fabricius, 1793: 525, concluded that the earlier description referred to the xantholine, Gauropterus fulgidus, and the later to the quediine, Quedius fulgidus]; Type locality: Germania).

  • — Panzer, 1795: 355 (Staphylinus; characters; Germany).

  • — Fabricius, 1801: 596 (Staphylinus; characters; Germany).

  • — Turton, 1802: 512 (Staphylinus; characters; Germany).

  • — Gyllenhal, 1810: 304 (Staphylinus; [Note: Misidentification: Bernhauer and Schubert, 1916: 423 cited variabilis variety e Gyllenhal as a synonym of fulgidus; there is no available name]; characters; Sweden).

  • — Erichson, 1839a: 486 (Quedius; characters; Germany).

  • — Heer, 1839: 273 (Philonthus; characters; Switzerland).

  • — Erichson, 1840: 525 (Quedius; characters; Europe; America septentrionalis).

  • — Redtenbacher, 1849: 708 (Philonthus; characters; Austria).

  • — Kiesenwetter, 1851a: 419 (Quedius; Pyrenees).

  • — Küster, 1853a: 60 (Quedius; characters; Europe).

  • — Hardy, 1851: 31 (Quedius; notes; England).

  • — Rosenhauer, 1856: 71 (Quedius; Granada; Sierra Nevada).

  • — Fairmaire and Laboulbène, 1856: 539 (Quedius; characters; France).

  • — Staudinger, 1857: 285 (Quedius; notes; Iceland).

  • — Kraatz, 1857d: 492 (Quedius; characters; Germany).

  • — Redtenbacher, 1857: 201 (Quedius; characters; Austria).

  • — Peyron, 1858: 428 (Quedius; Turkey).

  • — Kraatz, 1859: 67 (Quedius; characters; India).

  • — Thomson, 1860: 174 (Microsaurus; characters; Scandinavia).

  • — Hochhuth, 1862: 44 (Quedius; notes; Russia).

  • — Wollaston, 1865: 486 (Quedius; habitat; Canary Islands).

  • — Rottenberg, 1870: 28 (Quedius; notes; Sicily).

  • — Wollaston, 1871: 299 (Quedius; notes).

  • — Hochhuth, 1872: 131 (Quedius; notes; characters; Ukraine).

  • — Fauvel, 1874: 279 [= 1874a: 505] (Quedius; characters; France; Madeira and Canary Islands; Algeria; Caramania; Indonesia; Australia; USA).

  • — Redtenbacher, 1874: 198 (Quedius; characters; Austria).

  • — Seidlitz, 1875: 266 (Quedius; characters; Baltic region).

  • — Mulsant and Rey, 1876: 657 [= 1877: 513] (Quedius; characters; France).

  • — J. Sahlberg, 1876: 25 (Microsaurus; Finland).

  • — Fauvel, 1877: 268 (Quedius; characters; Australie, Melbourne, Rockhampton; Tasmanie).

  • — Provancher, 1877: 728 (Quedius; characters; Quebec).

  • — Fauvel, 1878: 127 [= 1878a: 47] (Quedius; Algeria).

  • — Fauvel, 1878g: 552 (Quedius; Australia).

  • — Weise, 1884: 423 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1440 cited erythrogaster Weise as a variety of fulgidus; Weise did not describe the species as new, he attributed it to Mannerheim]; variety of fulgidus; notes; characters).

  • — Fauvel, 1886: 57 [= 1886a: 49] (Quedius; Algeria).

  • — Fowler, 1888: 232 (Quedius; characters; Britain).

  • — Ganglbauer, 1895: 399 (Quedius; subgenus Microsaurus; characters; Europe; Mediterranean region; Atlantic Islands; India; Australia; New Zealand; North America).

  • — Fauvel, 1902b: 114 (Quedius; Algeria; Tunisia).

  • — Germain, 1903: 437 (Quedius; introduced species; notes; Chile).

  • — Sainte-Claire Deville, 1906: 101 (Quedius; Corsica).

  • — Porta, 1907: 114 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Fall and Cockerell, 1907: 166 (Quedius; New Mexico).

  • — Montandon, 1908: 80 (Quedius; Romania).

  • — Reitter, 1909: 111 (Quedius; subgenus Microsaurus; characters; Germany).

  • — Blatchley, 1910: 371 (Quedius; characters; Indiana).

  • — Johansen, 1914: 355 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; catalog).

  • — Notman, 1919a: 95 (Quedius; Ontario).

  • — Munster, 1923: 192 (Quedius; subgenus Microsaurus; Fennoscandia).

  • — Gridelli, 1924d: 51 (Quedius; subgenus Microsaurus; characters; Germany; Styria; Transylvania; Dalmatia; Italy; Portugal).

  • — Gridelli, 1929: 22 (Quedius; characters; Germany, France, Portugal, Bohemia, Russia).

  • — Portevin, 1929: 341 (Quedius; characters; France).

  • — Cameron, 1932: 288 (Quedius; subgenus Microsaurus; characters; India).

  • — Scheerpeltz, 1933: 1440 (Quedius; catalog).

  • — Oke, 1933: 123 (Quedius; tarsal characters).

  • — Scheerpeltz, 1935a: 655 (Philonthus; Australia).

  • — Wüsthoff, 1938a: fig. 9 (Quedius; aedeagal illustration).

  • — Tottenham, 1939d: 237 (Quedius; synonym of assimilis).

  • — Hellén, 1939: 32 (Quedius; characters; Finland).

  • — Roubal, 1941b: 124, 127 (Quedius; subgenus Microsaurus; characters; collecting notes).

  • — Paulian, 1941: 264 (Quedius; larval characters).

  • — Brinck, 1948: 34 (Quedius; collecting and ecological notes; Tristan da Cunha).

  • — Jeannel and Jarrige, 1949: 383 (Quedius; subgenus Microsaurus; notes; cave species; France; Spain; Algeria).

  • — Tottenham, 1949: 422 (Quedius; synonym of assimilis).

  • — V. Hansen, 1952: 133 (Quedius; subgenus Microsaurus; characters; Denmark).

  • — Hatch, 1957: 221 (Quedius; subgenus Microsaurus; characters; British Columbia; Washington; Oregon; Idaho).

  • — Smetana, 1958: 349 (Quedius; subgenus Microsaurus; characters; ecology; Czechoslovakia).

  • — Smetana, 1960e: 157 (Quedius; subgenus Microsaurus; characters).

  • — Palm, 1963b: 134 (Quedius; subgenus Microsaurus; characters; Sweden; Denmark; Norway; Finland).

  • — Lohse, 1964: 209 (Quedius; subgenus Microsaurus; characters; central Europe).

  • — Raianu, 1964: 102 (Quedius; Romania).

  • — Horion, 1965: 262 (Quedius; subgenus Microsaurus; all central and southern Europe; Finland; Sweden; Norway; Britain; Denmark; Ukraine).

  • — Coiffait and Saiz, 1966: 412 (Quedius; subgenus Microsaurus; characters; Chile).

  • — Pototskaia, 1967: 83 (Quedius; larval characters).

  • — Korge, 1968: 51 (Quedius; notes).

  • — Szujecki, 1968a: 734 (Quedius; Poland).

  • — Smetana, 1971: 66 (Quedius; subgenus Microsaurus; characters; British Columbia; Manitoba; Arizona; California; Colorado; Georgia; Idaho; Illinois; Indiana; Iowa; Kansas; Kentucky; Maryland; Massachusetts; Michigan; Minnesota; Missouri; New Jersey; New Mexico; New York; Oregon; Pennsylvania; Texas; Virginia; Washington; West Virginia; Wisconsin).

  • — Smetana, 1971b: 2 (Quedius; subgenus Microsaurus; California).

  • — Saiz, 1971: 385 (Quedius; Chile).

  • — Tikhomirova, 1973a: 188 (Quedius; checklist USSR).

  • — Bordoni, 1973d (Quedius; subgenus Microsaurus; aedeagal anomaly).

  • — Bordoni, 1974: 65 (Quedius; subgenus Microsaurus; characters; Italy).

  • — Bordoni, 1974b: 10 (Quedius; Italy).

  • — Pope, 1977: 31 (Quedius; Britain).

  • — Bordoni, 1977a: 18 (Quedius; subgenus Microsaurus; Spain).

  • — Coiffait, 1978: 146 (Quedius; subgenus Microsaurus; characters; Palaearctic region).

  • — Smetana, 1978a: 824 (Quedius; subgenus Microsaurus; collecting notes; Tennessee).

  • — Bordoni, 1978a: 63 (Quedius; cave collection; Turkey).

  • — Muona, 1979: 15 (Quedius; synonym of assimilis).

  • — Bordoni, 1981a: 14 (Quedius; pupal characters; Italy).

  • — Uhlig and Vogel, 1981: 123 (Quedius; notes; Germany).

  • — Bordoni, 1982a: 140 (Quedius; cave collection; Sardinia).

  • — Frank, 1982: 46 (Quedius; list of parasites).

  • — Tóth, 1984: 115 (Quedius; characters; Hungary).

  • — Segers, 1986: 49 (Quedius; Belgium).

  • — Lucht, 1987: 109 (Quedius; checklist; Germany; Poland; Czechoslovakia; Austria; France; Benelux; Denmark; Sweden).

  • — Smetana, 1988: 209 (Quedius; subgenus Microsaurus; characters).

  • — Lohse and Lucht, 1989: 172 (Quedius; synonym of assimilis).

  • — Schülke and Uhlig, 1988: 12 (Quedius; Germany).

  • — M. Hansen, 1991: 107 (Quedius; synonym of assimilis).

  • — Elias, 1991: 557 (Quedius; Quaternary fossil collection; Montana).

  • — Campbell and Davies, 1991: 121 (Quedius; Canada).

  • — Silfverberg, 1992: 17 (Quedius; synonym of assimilis).

  • — Smetana, 1993b: 50 (Quedius; synonym of rufitarsis).

  • — Dearborn and Donahue, 1994: 16 (Quedius; Maine).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (Quedius; synonym of assimilis).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; synonym of rufitarsis).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (Quedius; synonym of assimilis).

  • — M. Hansen, 1996: 107 (Quedius; synonym of assimilis).

  • — Downie and Arnett, 1996: 391 (Quedius; subgenus Microsaurus; characters; USA).

  • — Ádám, 1996: 247 (Microsaurus; synonym of rufitarsis).

  • — Emberson, 1998: 36 (Quedius; collecting notes; Chatham Islands).

  • — Bordoni and Oromi, 1998: 1157 (Quedius; subgenus Microsaurus; facultative cave species).

  • — Zerche, 2000: 52 (Quedius; synonym of assimilis).

  • — Herman, 2001: 29, 83 (Quedius; case presented for citing assimilis as valid in application to Commission under Article 23.9.3, but opinion reversed and petition will request that fulgidus be cited as name of species).

  • Distribution: Libya, Tunisia, Algeria, Tristan da Cunha, Canary Islands, Madeira, Iceland, Europe, Russia, Turkey, India, Indonesia, Australia, New Zealand, Chatham Islands, Canada, USA, Chile.

fuscipennis Block, 1799: 116 (Staphylinus; [Note: Type locality: Tharand, Plauischer Grund bei Dresden).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 428 (Quedius; synonym of mesomelinus).

  • — Smetana, 1958: 353 (Quedius; synonym of mesomelinus).

  • — Bordoni, 1974: 33 (Quedius; synonym of mesomelinus).

  • — Coiffait, 1978: 150 (Quedius; synonym of mesomelinus).

  • — Ádám, 1996: 247 (Microsaurus; valid name; Hungary).

  • — Zerche, 2000: 52, 53 (Quedius; synonym of assimilis; discussion; nomen oblitum).

rufitarsis Marsham, 1802: 512 (Staphylinus; Type locality: Britain).

  • — Gyllenhal, 1810: 304 (Staphylinus; synonym of variabilis Gyllenhal).

  • — Curtis, 1829: 25 (Quedius; catalog; Britain).

  • — C. Sahlberg, 1830: 316 (Staphylinus; variety of variabilis Gyllenhal; characters; Finland).

  • — Stephens, 1832: 220 (Quedius; characters; England).

  • — Curtis, 1837: pl. 638 (Quedius; characters; Britain).

  • — Stephens, 1839: 391 (Quedius; characters; England).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

  • — Nowosad, 1990: 136 (Quedius; cited as valid species; collected from mole nest; notes; Poland).

  • — Smetana, 1993b: 50 (Quedius; subgenus Microsaurus; cited as valid species; Czech Republic; Slovakia).

  • — Ciceroni and Zanetti, 1995: 33 (Quedius; cited as valid species; Italy).

  • — Ádám, 1996: 247 (Microsaurus; valid species; Hungary).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

iracundus Say, 1830: 35 [= 1834: 449] (Staphylinus; Type locality: Indiana).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — LeConte, 1859: 564 (Quedius).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Horn, 1878: 158 (Quedius; [Note: Misidentification: Scheerpeltz, 1933: 1444 cited fulgidus Horn as a synonym of iracundus; Horn did not describe the species as new, he attributed it to Say]; characters; California; Georgia).

  • — Horn, 1878: 158 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Casey, 1915: 411 (Microsaurus; characters; Iowa; Indiana).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1444 (Quedius; catalog).

  • — Voris, 1939: 188 (Quedius; larval characters; Indiana).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

haemopterus Stephens, 1832: 217 (Quedius; Type locality: London; Suffolk; Devonshire).

  • — Curtis, 1829: 25 (Quedius; nomen nudum; catalog; Britain).

  • — Stephens, 1839: 390 (Quedius; variety of nitidus; characters; England).

  • — Hardy, 1851: 32 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Tottenham, 1939d: 237 (Quedius; possible synonym of picipes [Note: Species not specifically listed as a synonym of picipes by Tottenham, 1949, Kloet and Hincks, 1945, or Pope, 1977]).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

haemorrhoidalis Stephens, 1832: 218 (Quedius; Type locality: London; Devonshire).

  • — Stephens, 1839: 390 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

nigricornis Stephens, 1832: 222 (Quedius; Type locality: London).

  • — Stephens, 1839: 392 (Quedius; characters; England).

  • — G. Waterhouse, 1858: 22 (Quedius; synonym of fulgidus).

  • — Rye, 1870a: 82 (Quedius; synonymn of fulgidus; see last sentence under notes on nigricornis Holme which begins on page 81).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

distinctus Runde, 1835: 6 (Staphylinus; Type locality: Halae).

  • — Erichson, 1839a: 486 (Quedius; synonym of fulgidus).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Herman, 2001: 29 (Quedius; synonym of assimilis).

assimilis Nordmann, 1837: 78 (Philonthus; name given to variabilis variety f of Gyllenhal; Type locality: Europa).

  • — Erichson, 1839a: 486 (Quedius; synonym of fulgidus).

  • — Heer, 1839: 273 (Philonthus; synonym of fulgidus).

  • — Erichson, 1840: 526 (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1453 (Quedius; synonym of ochripennis).

  • — Tottenham, 1939d: 237 (Quedius; valid name to replace fulgidus).

  • — Kloet and Hincks, 1945: 167 (Quedius; cited as valid name to replace fulgidus; Britain).

  • — Hinton, 1945: 70 (Quedius; characters).

  • — Tottenham, 1949: 422 (Quedius; checklist; Britain).

  • — Scheerpeltz, 1968a: 89 (Quedius; synonym of ochripennis).

  • — Pope, 1977: 31 (Quedius; synonym of fulgidus).

  • — Muona, 1979: 15 (Quedius; cited as valid name; Denmark; Norway; Sweden; Finland; Russia; Baltic States).

  • — Lohse and Lucht, 1989: 172 (Quedius; note).

  • — M. Hansen, Kristensen, Mahler, and Pedersen, 1991: 107 (Quedius; Denmark).

  • — Silfverberg, 1992: 17 (Quedius; valid species; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

  • — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (Quedius; cited as valid; Denmark).

  • — M. Hansen, Mahler, Palm, and Pedersen, 1996: 245 (Quedius; cited as valid name; Denmark).

  • — M. Hansen, 1996: 107 (Quedius; cited as valid name; Denmark).

  • — Telnov, Barsevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 39 (Quedius; valid species; Latvia).

  • — Anderson, Nash, and O'Connor, 1997: 19 (Quedius; Ireland).

  • — Assing, 1998e: 140 (Quedius; Canary Islands).

  • — Zerche, 2000: 52, 53 (Quedius; discussion; nomen protectum).

  • — Herman, 2001: 29, 83 (Quedius; case presented for citing name as valid in petition to Commission under Article 23.9.3, but opinion reversed and petition will request that fulgidus be cited as name of species).

bicolor Redtenbacher, 1849: 710 (Philonthus; [Note: Bernhauer and Schubert, 1916: 423 cited Redtenbacher, 1872 (= 1874): 201 as the first reference to this name]; Type locality: Wien).

  • — Kraatz, 1857d: 492 (Quedius; synonym of fulgidus).

  • — Redtenbacher, 1857: 204 (Quedius; characters; Austria).

  • — Fauvel, 1866c: 247 [= 1866d: 105] (Quedius; synonym of fulgidus).

  • — Fauvel, 1869: 486 (Quedius; synonym of fulgidus).

  • — Fauvel, 1874: 280 [= 1874a: 506] (Quedius; synonym of fulgidus).

  • — Redtenbacher, 1874: 201 (Quedius; characters; Austria).

  • — Fowler, 1888: 232 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Scheerpeltz, 1933: 1440 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

quadripunctatus Thomson, 1867: 159 (Microsaurus; Type locality: Scandinavia).

  • — Fauvel, 1876a: 76 [= 1876: 261] (Quedius; synonym of fulgidus).

  • — J. Sahlberg, 1876: 25 (Microsaurus; synonym of fulgidus).

  • — Fowler, 1888: 232 (Quedius; synonym of fulgidus).

  • — Ganglbauer, 1895: 399 (Quedius; synonym of fulgidus).

  • — Porta, 1907: 114 (Quedius; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

  • — Smetana, 1958: 349 (Quedius; synonym of fulgidus).

  • — Smetana, 1971: 66 (Quedius; synonym of fulgidus).

  • — Bordoni, 1974: 65 (Quedius; synonym of fulgidus).

  • — Coiffait, 1978: 146 (Quedius; synonym of fulgidus).

peranxius Mulsant and Rey, 1876: 661 [= 1877: 517] (Quedius; Type locality: Not cited).

  • — Duvivier, 1883: 138 (Quedius; catalog; synonym of fulgidus).

  • — Bernhauer and Schubert, 1916: 423 (Quedius; synonym of fulgidus).

  • — Cameron, 1932: 288 (Quedius; synonym of fulgidus).

erythropygus Gridelli, 1929: 24 (Quedius; variety of fulgidus; Type locality: Cirenaica: Golfo di Bomba; Bengasi).

  • — Gridelli, 1930: 77 (Quedius; variety of fulgidus; [Note: Listed as a new variety but it has the same locality as the variety of the same name described in 1929 so I list this citation as a subsequent reference]; Libya).

  • — Scheerpeltz, 1933: 1440 (Quedius; variety of fulgidus).

  • — Roubal, 1941b: 124 (Quedius; subgenus Microsaurus; variety of fulgidus; char