Rakotondrainibe, F. & A. Jouy (2015). Novelties in the genus Huperzia Bernh. (Lycophytes, Lycopodiaceae) from Madagascar. Candollea 70:49–56. In French, English and French abstracts.
Within the context of the revision of Lycopodiaceae from Madagascar, two new species, Huperzia ambrensis Rakotondr. and Huperzia teretirigida Rakotondr., are described and illustrated. Two new synonyms are also proposed: Huperzia hildebrandtii (Herter) Tardieu is a synonym of Huperzia squarrosa (G. Forst.) Trevis. and Huperzia rubrica (Herter) Tardieu is a synonym of Huperzia ophioglossoides (Lam.) Rothm. A presumed hybrid, Huperzia megastachya (Baker) Tardieu × Huperzia squarrosa (G. Forst.) Trevis., is described and compared with its putative parents.