DYMYTROVA, L., NADYEINA, O., NAUMOVYCH, A., KELLER, C. & SCHEIDEGGER, C. 2013. Primeval beech forests of Ukrainian Carpathians are sanctuaries for rare and endangered epiphytic lichens. — Herzogia 26: 73– 89.
A total of 203 epiphytic lichens and lichen-allied fungi were recorded in the Uholsko-Shyrokoluzhanskyi primeval beech forests, seven of them (Buellia chloroleuca, Lecanora strobilina, Ramonia luteola, Rinodina capensis, Thelopsis flaveola, Usnea wasmuthii and Wadeana dendrographa) are listed for the first time for Ukraine. 25 lichen species, including Arthonia vinosa, Arthopyrenia punctiformis, A. rhyponta, Bacidia incompta, Biatora carneoalbida, Buellia insignis, B. schaereri, Caloplaca cerina var. chloroleuca, C. monacensis, Megalaria laureri, Piccolia ochrophora, Porina hibernica, P. leptalea, Reichlingia leopoldii, Thelocarpon laureri, Verrucaria viridigrana, Xanthoria fulva, X. ulophyllodes are reported as new to the Carpathian Biosphere Reserve. The high species richness of rare and threatened lichens recorded in the Uholsko-Shyrokoluzhanskyi primeval forest stands underlines a high value of this area for the conservation of epiphytic lichens, i.e. an important part of forest biodiversity.