New Data on Homocladus grandis, A Permian Stem-Mantodean (Polyneoptera: Dictyoptera)

Olivier Béthoux; Roy J. Beckemeyer; Michael S. Engel; Joseph D. Hall

doi:10.1666/09-121.1

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1 July 2010 New Data on Homocladus grandis, A Permian Stem-Mantodean (Polyneoptera: Dictyoptera)

Olivier Béthoux, Roy J. Beckemeyer, Michael S. Engel, Joseph D. Hall

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J. of Paleontology, 84(4):746-753 (2010). https://doi.org/10.1666/09-121.1

Abstract

Representatives of the family Strephocladidae have been considered as fossil relatives (i.e., stem-group) of Mantodea (mantises) based on characters of the forewing morphology. Here we describe new specimens from the Wellington Formation that we assign to the strephocladid species Homocladus grandis Carpenter, 1966. The range of morphological variation exhibited by the new material, in addition to wing morphology variability documented in extant mantises and roaches, suggest that H. ornatus Carpenter, 1966 and Paracladus retardatus Carpenter, 1966, reported from the same formation, are new junior subjective synonyms of H. grandis. We describe the first hind wing for this species based on a well-preserved specimen. It exhibits a combination of character states unique to dictyopteran insects.

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Olivier Béthoux, Roy J. Beckemeyer, Michael S. Engel, and Joseph D. Hall "New Data on Homocladus grandis, A Permian Stem-Mantodean (Polyneoptera: Dictyoptera)," Journal of Paleontology 84(4), 746-753, (1 July 2010). https://doi.org/10.1666/09-121.1

Accepted: 1 March 2010; Published: 1 July 2010

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