A newly discovered trilobite fauna from the Cambrian Honey Creek Formation marks a distinct interval that follows an extinction event. Dominated by the genus Monocheilus in association with Ptychaspis, it resembles faunas from Alberta, Canada, and the Upper Mississippi Valley region of the United States. Ptychaspis bullasa Lochman and Hu, 1959 is a species that has been reported widely in North America. However, restudy of various museum collections shows that the various occurrences record a set of more narrowly distributed species. The pattern of distribution is similar to groups of modern “pseudocryptic species” identified by a combination of genetic and anatomical data.
The Cambrian (Jiangshanian, Sunwaptan) Honey Creek Formation in the Wichita Mountains region of Oklahoma yielded a new fauna dominated by Monocheilus Resser, 1937 (senior synonym of Stigmacephalus Resser, 1937) in association with Ptychaspis Hall, 1863. It occupies the same stratigraphic position as similar faunas in the Upper Mississippi Valley and Alberta, lying a little above an interval characterized by species of Taenicephalus Ulrich and Resser in Walcott, 1924 and Orygmaspis Resser, 1937. Revision of Ptychaspis bullasa Lochman and Hu, 1959 from type material from Idaho and sclerites attributed to the species from Texas reveals a plexus of pseudocryptic species that share tuberculate sculpture on the cranidium. New species are Monocheilus reginae, Monocheilus richardi, Ptychaspis occulta, and Ptychaspis matuszaki.