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The narrowing of the Florida Peninsula into the Florida Keys concentrates large flights of migrating raptors en route to southern wintering grounds each fall. Between 1999 and 2004, HawkWatch International (HWI) conducted standardized daily counts of migrant raptors from 15 September to 13 November at Curry Hammock State Park in the middle Keys. Average annual counts for the eight most common species were: Broad-winged Hawk (Buteo platypterus; 3893); Sharp-shinned Hawk (Accipiter striatus; 3300); American Kestrel (Falco sparverius; 2800); Peregrine Falcon (F. peregrinus; 1908); Osprey (Pandion haliaetus; 1171); Northern Harrier (Circus cyaneus; 568); Merlin (F. columbarius; 554); and Cooper's Hawk (A. cooperii; 545). Counts were comparable to or greater than totals from each of nine sites representative of the major raptor migration corridors within the Nearctic-Neotropical migration system. Given the unique geographic position of the Florida Keys in the southeastern U.S. where no other major concentrations of migratory raptors occur, long-term counts at Curry Hammock will be important for monitoring North American raptor population trends. Counts of migrating Peregrine Falcons at this site may be of particular importance, because more costly monitoring efforts at breeding sites will decrease over time now that peregrines are no longer listed as an Endangered Species.

Each fall, millions of diurnal raptors migrate south from temperate breeding areas to wintering areas in the southern United States, Mexico, the Caribbean, Central America, and South America (Heintzelman 1986, Kerlinger 1989). Many of these raptors are counted each fall at hawkwatches situated along the major migration corridors across North America. During the breeding season and winter, many raptor species occupy remote habitats and large home ranges, making their populations difficult and costly to monitor (Fuller and Mosher 1987, Dunn 2005). During fall migration, however, raptors become conspicuous and numerous at locations where their flights are concentrated due to topographic or geographic features (Mueller and Berger 1967, Hoffman 1985, Heintzelman 1986). Consequently, long-term counts at a network of hawkwatch sites across North America have been proposed as an efficient and cost-effective means for monitoring the status and trends of raptor populations (Bednarz and Kerlinger 1989, Titus et al. 1990, Smith and Hoffman 2000) and raptor migration count data have figured prominently in comprehensive reviews of the status of raptor populations in Canada (Kirk and Hyslop 1998) and for all of North America (Fuller 1996).

The continental status of Peregrine Falcon (Falco peregrinus) populations has been reviewed several times in the past forty years (Hickey 1969, Cade et al. 1988, Enderson et al. 1995), most recently in the context of the proposed removal of the anatum subspecies from the United States Fish and Wildlife Service (USFWS) list of Endangered and Threatened Species (Millsap et al. 1998). The tundrius subspecies was removed from the Endangered Species list in 1994 (Swem and Ambrose 1994) and the anatum subspecies was removed in 1999 (Mesta 1999). Now that North American Peregrine Falcons will no longer receive Endangered Species level research attention, monitoring of peregrine populations will be critical to ensure that populations remain stable or continue to show increasing trends (Millsap et al. 1998, U.S. Fish and Wildlife Service 2003), particularly if falconry take of migrant peregrines is approved by the USFWS (B. Millsap pers. comm.).

Migration monitoring data are particularly important for species where more than a third of the species' breeding range or most of a species' wintering range occurs outside the coverage of major monitoring programs such as the Breeding Bird Survey or Christmas Bird Count (Dunn et al. 2005, Hussell and Ralph 2005). This is true for a number of migrant raptor species that have large breeding populations in boreal or arctic regions where regular survey efforts would be cost-prohibitive for long-term monitoring (Bart and Ralph 2005, Dunn 2005). Migration count data were used to document Peregrine Falcon population declines during the DDT era (Carson 1962, Mueller et al. 1988) and Enderson (1969) suggested that peregrine populations could be effectively monitored by a network of count sites at locations where flights of migrant peregrines concentrate along the Atlantic and Gulf of Mexico coasts. Migration count data from the Florida Keys and elsewhere should provide valuable data to evaluate the overall long-term status and trends of Peregrine Falcon populations at a fraction of the cost of short-term breeding season monitoring programs (U.S. Fish and Wildlife Service 2003).

Some of the major raptor migration concentration points in the Nearctic-Neotropical migration system, such as the coastlines of the Great Lakes, northern Appalachian ridges, and major peninsulas along the North Atlantic coast, have been known for many years (Heintzelman 1986, Kerlinger 1989); however, other major sites have been documented more recently (Smith and Hoffman 2000). Long-term counts at well-established sites in the upper Midwest and the northeastern U.S.A. have produced information on population trends for raptors breeding to the north of these sites in the U.S. and Canada (Bednarz et al. 1990, Hussell and Brown 1992) and major raptor migration count sites in western North America have documented population trends for western raptor populations since the mid-1980s (Hoffman and Smith 2003). In the mid-1990s, standardized counts were initiated at several major migration concentration points along the western Gulf of Mexico, spanning from Smith Point, Texas to Veracruz, Mexico, where flights of more than five million raptors have been observed in a single season (Smith and Hoffman 2000, Ruelas Inzunza et al. 2000, Smith et al. 2001). Raptor migration counts in Costa Rica that began in 2000 have begun to quantify the enormous concentration of migrant raptors that has long been known to occur across the Mesoamerican land corridor (Smith 1980, Porras-Penaranda et al. 2004). Using data from single-day counts in October over several years, Hoffman and Darrow (1992) suggested that the Florida Peninsula may be a raptor migration corridor of continental significance and that the Florida Keys represent the largest concentration point along this corridor. This was confirmed by two years of full season counts by Brashear and Stoddard (2001), which documented high count totals for Peregrine Falcons and seven other species. These short-term counts were conducted from private property, which was not conducive to the development of a long-term raptor migration monitoring site (Brashear and Stoddard 2001).

In autumn 1999, HawkWatch International (HWI) initiated its first full-season count of migrating raptors in the middle Florida Keys at Curry Hammock State Park. This count was designed to document long-term trends in Peregrine Falcon populations as well as population trends for the seven other common diurnal raptor species. Because the Florida Keys are situated near the southern end of the breeding distributions of most North American raptor populations, potential source areas for Florida Keys migrants are vast, and mostly unknown. I summarize 6 yr of raptor migration count data from Curry Hammock State Park to document the magnitude and timing of raptor migration through the Florida Keys. I also compare Curry Hammock count totals with nine other raptor migration count sites across North and Central America to place raptor migration through the Florida Keys in the context of the major raptor migration concentration points within the Nearctic-Neotropical migration system and to demonstrate the particular importance of these counts for monitoring long-term trends in Peregrine Falcon populations.


Study Site

The Florida Keys are an approximately 170-km-long island chain that trends northeast to the southwest from the southern tip of the Florida peninsula (Fig. 1). The Curry Hammock hawkwatch was located in the middle Florida Keys at Curry Hammock State Park, on Little Crawl Key, just northeast of the town of Marathon at 24°44.50′N, 80°59.00′W. Earlier, simultaneous counts at multiple locations throughout the Florida Keys demonstrated that the flight path of migrant raptors tends to be most concentrated in the middle Keys due to the relatively narrow landmass and the lack of alternative overland routes that are formed by strings of offshore islands in both the Upper and Lower Keys (Hoffman and Darrow 1992). HWI chose Curry Hammock State Park as the hawkwatch location based on the combination of its favorable location in the middle Keys and open public access, which has allowed us to develop a complementary environmental education program.

Figure 1

Map of south Florida and the Florida Keys. Detailed inset map at bottom shows the Middle Keys near the Curry Hammock hawkwatch.


For 6 km in either direction from the Curry Hammock hawkwatch, the middle Keys are typically less than 1 km wide (range 0.1–2 km in width). Birds that travel over land or slightly offshore within this area are often visible to the naked eye. We assumed that nearly all birds flying in the area of Curry Hammock State Park were detectable by binoculars, regardless of distance or direction from the hawkwatch. To qualitatively test this assumption, pairs of observers made simultaneous observations from both the north end of Long Point Key and the hawkwatch platform on Little Crawl Key on two different days when the primary flight line was well offshore of Long Point Key to the north (Fig. 1). This was the greatest distance from the Curry Hammock hawkwatch platform that we ever observed migrating raptors in this part of the Florida Keys. Observers at the hawkwatch communicated numbers and species of migrating birds they saw to observers on Long Point Key by radio in real time. Although many of these birds were nearly 2.5 km from the hawkwatch, birds were still easily detectable with binoculars and identifiable with a 30× spotting scope, suggesting that regardless of the daily flight path (which shifts with local weather conditions), migrant raptors were detectable from the hawkwatch platform.

The hawkwatch was on the elevated deck of a park restroom. The area around the hawkwatch had been cleared of vegetation in preparation for a development project that never came to fruition and observers had a relatively unobscured view of the sky in all directions. The vegetation surrounding the hawkwatch was mostly native grasses and mixed small native trees; primarily Red Mangrove (Rhizophora mangle) or Buttonwood (Conocarpus erectus). Observers standing on the deck at the hawkwatch could see over most of these trees; however, some trees to the north and west of the site made detection of distant low-flying birds difficult. Consequently, observers scanned this area more frequently, particularly on windy days when birds were most likely to fly low. This “blind spot” may have resulted in undercounting some species that often migrate at low altitudes through the Florida Keys, particularly Merlins (F. columbarius).

Count Period

Previous full-season counts of migrating raptors in the Florida Keys by Brashear and Stoddard (2001) began as early as 5 August. Data from these counts indicated limited passage of most species before 15 September or after 13 November. Exceptions included small numbers of Swallow-tailed Kites (Elanoides forficatus) and Ospreys (Pandion haliaetus) in August and early September. In this study, two official observers conducted standardized daily counts of migrating raptors at Curry Hammock State Park between 15 September and 30 October, 1999, and between 15 September and 13 November each year from 2000–04. The shorter count period in 1999 was due to limited start-up funds.

Daily counts were conducted from 0800–1600 Eastern Standard Time (EST). Previous counts by Brashear and Stoddard (2001) indicated low numbers of observations after 1600 EST on most days. However, occasional, large, late-afternoon flights (mostly by American Kestrels [F. sparverius], Ospreys, and Northern Harriers [Circus cyaneus]), were observed. To reduce the chance of missing these late-afternoon flights, observers employed the “15-min rule,” and stayed open for an additional 15 min if more than 10 southbound birds were counted during the last 15 min of the count. Thus, if more than 10 southbound birds were counted between 1545–1600, observers continued observations until the first 15-min period went by with fewer than 10 southbound migrants observed.


Two primary observers conducted the daily count each year. All observers had at least one full season of experience as a primary hawk counter at another migration site in North America and were capable of identifying migrant raptors in flight prior to being hired in the Florida Keys. Each primary observer worked 5–6 d per wk. On their days off, another similarly qualified observer filled in as a primary observer. At the beginning of each season, new observers were trained in count data collection protocols, ensuring that scanning routines, flight-line coverage, and counting, recording, and data entry protocols were consistent among years. Observer experience and quality remained fairly consistent across all 6 yr of the count.

Count and Data Collection Protocols

Observers used high quality 8–10× binoculars to spot and identify migrating raptors. Wide-angle 30× spotting scopes were used to assist in the identification of distant birds. Dunne et al. (1988), Wheeler and Clark (1995), and Clark and Wheeler (2001) served as primary identification references. Both observers scanned the sky regularly with naked eyes, dividing the sky into quadrants of responsibility depending on the flight patterns of the day. Observers systematically scanned more distant areas with binoculars once every 1–2 min.

Observers recorded the species and flight direction of each migrant raptor seen from the watch site. Migrants following the Keys from the northeast to southwest were classified as southbound and migrants traveling southwest to northeast were classified as northbound (Fig. 1). Multiple-tab clickers were used to tally sightings for common species on busy days, minimizing the amount of time spent looking away from the sky while recording data. Hourly totals for each species were transferred from the clickers to the datasheets at the end of each hour. Mechanical hand clickers were used to count individuals in large flocks (200–800 individuals). Observers were conscious of the potential to count resident raptors as migrants and care was taken to follow the flight paths of common resident or wintering species well to the south or north of the count site before each bird was recorded as a migrant. Resident Ospreys were common around the hawkwatch in all years and Bald Eagles (Haliaeetus leucocephalus) were regular around the hawkwatch in some years. Individual resident Bald Eagles were identifiable by unique plumages. Resident Ospreys frequently vocalized as migrants passed through, simplifying their identification as residents; however, because they were so common as residents, observers spent additional time ensuring that migrant Ospreys continued to fly well past the hawkwatch and out of site to the south. Nonetheless, some resident Ospreys may have been counted as migrants. After the fall 2000 season, observers stopped counting the large flocks of Turkey Vultures (Cathartes aura) that move throughout the Florida Keys in late fall and early winter (Darrow 1983), because the flight direction of these flocks was often difficult to determine and observers frequently observed the same flock of vultures heading north and south several times on the same day.

Count Totals

Count totals are represented in tables and figures as southbound, net, or percent northbound sightings. Southbound totals best represent the total number of birds that migrate into the Middle Keys each fall, regardless of their final wintering destination. Southbound totals include both the birds that continue to the south of the hawkwatch and the birds that head south past the watch site and later reverse the direction of their flight, passing by the hawkwatch again on their way north. Net totals represent southbound observations minus northbound observations for each species. Therefore, net totals reflect the number of individuals that continue past the count site to either winter in the lower Keys or continue their migration through the Caribbean. Some species with a high percentage of northbound sightings (such as Broad-winged Hawks [Buteo platypterus] and Swainson's Hawks [B. swainsoni]) may move back and forth through the Keys more than once during the migration period and are thus likely to be counted more than once as they repeatedly pass the hawkwatch. For these two species, net totals may be closer to the true number of birds that pass through the Middle Keys than southbound totals.

I compared southbound totals from the Florida Keys for the 6 yr between 1999–2004 with count totals from other major raptor migration sites in North America for this same range of years (unless otherwise noted in Table 3). I selected nine raptor migration sites for comparison that are representative of the major regions where flights of migrating raptors are known to concentrate during fall in North America (Heintzelman 1986, Kerlinger 1989, Smith and Hoffman 2000). Each of these sites met the four following criteria: (1) count data were available for 1999–2004 (see Table 3 for one exception); (2) standard data collection protocols were employed for this range of years; (3) the count period at each of these sites encompassed the full migration period for each of the eight common species included in comparisons; and (4) permission was obtained to present count totals in this paper for this comparison. In some cases, counts from multiple sites within a single region (e.g., the Appalachian Mountains, the Great Lakes, or the North Atlantic Coast) were available, but data from only a single representative site for each region were included in this comparison. Other sites within these same regions typically have similar species compositions and relatively similar count totals. Annual count totals from count sites were obtained from the Hawk Migration Association of North America's online database (, websites for individual hawk migration sites, or directly from count site coordinators (see Table 3).

Passage Dates

Average median and bulk passage dates of raptors through the Florida Keys were calculated for all species with more than 20 southbound sightings per year using daily totals for each of the years between 2000–04. Data from 1999 were not included in these calculations because this season ended 14 d earlier than the other five seasons. Bulk passage dates were the range of dates between which the central 80% of the entire season's southbound total for each species passed through the Florida Keys (i.e., the first date is the date by which 10% of the season's cumulative sightings have been made and the last is the date by which 90% of the season's cumulative sightings have been made).


Observation Effort

Observation dates varied slightly across the six years of this study (CV  =  8%). Some of this variation is related to different start dates among years (see Methods) and the rest is due to different amounts of full-day closures among years due to mandatory hurricane evacuations. In 2004, the start date for the count was delayed 4 d due to a mandatory evacuation of the Florida Keys for Hurricane Ivan. Observations were cancelled for one full day five different times between 1999–2004. Other than these five within-season full-day closures due to hurricanes or extreme weather, daily counts were rarely suspended for more than 1 hr due to rain and observers continued to count migrants during light to moderate rains. The number of hours of observation per day varied only slightly among years (range 6.8–8.4 hr/d).

Count Totals

Between 1999–2004, observers counted an average southbound total (excluding Turkey Vultures) of 15 036 raptors (CV  =  10%) of 16 different species (Table 1). Eight raptor species comprised 98% of all southbound observations. Broad-winged Hawks were the most abundant raptor observed, followed by Sharp-shinned Hawks (Accipiter striatus), American Kestrels, Peregrine Falcons, Ospreys, Northern Harriers, Merlins, and Cooper's Hawks (A. cooperii; Table 1). Six species were uncommon during fall migration, averaging 11–84 observations per yr. These were: Swainson's Hawks, Short-tailed Hawks (B. brachyurus), Mississippi Kites (Ictinia mississippiensis), Red-shouldered Hawks (B. lineatus), Bald Eagles, and Swallow-tailed Kites. Two species averaged fewer than two sightings per year: Red-tailed Hawks (B. jamaicensis), and Black Vultures (Coragyps atratus). Observers counted 11 932 southbound Turkey Vultures in 2000, the only year when this species was counted during their peak migration into the Florida Keys.

Table 1

Southbound and net count totals (and percentage of northbound sightings) for the raptor migration count at Curry Hammock State Park, Florida Keys, FL, 1999–2004.


Species abundance ranks for net totals differed from those for southbound totals (Table 1). Most notably, net totals for Broad-winged Hawks were much lower than southbound totals, reflecting the high proportion of northbound sightings for this species. Sharp-shinned Hawks and American Kestrels had the highest net totals of all species (Table 1). Net totals were also high for Peregrine Falcons, Broad-winged Hawks, Ospreys, Northern Harriers, Merlins, and Cooper's Hawks. A net total of 3720 Turkey Vultures were counted in 2000, suggesting that only a small percentage of the 11 932 Turkey Vultures observed heading south at Curry Hammock continued their migration to the south of the Florida Keys. Northbound birds comprised 21% of all observations across all years of the count (Table 1). For species with more than 500 observations per year, this ranged from a high of 35% in Broad-winged Hawks to a low of 4% for American Kestrels and Northern Harriers.

Seasonal Timing of Migration

Fall raptor migration through the Florida Keys took place within a relatively short time frame from mid-September through early November. However, the timing of migration within this window varied among years and daily counts varied considerably within each season. Each fall, there was a large number of days (47–70% of d per yr) with very low counts, <1% of the entire season's total. Conversely, infrequent and large, single-day flights often accounted for a large percentage of the entire season's total. In the 6 yr between 1999–2004, the 5 d with the highest count totals within each season accounted for between 32–47% of the total annual count for all species combined.

The seasonal timing of raptor migration through the Florida Keys varied among species (Table 2). Mississippi Kites, Ospreys, and Merlins were early season migrants and Short-tailed Hawks and Swainson's Hawks were late season migrants. Among the common species, annual passage dates were fairly predictable, and median passage dates showed less annual variation than peak passage dates (Table 2). Bulk (80%) passage took place over a relatively small number of days for some species (e.g., Peregrine Falcons, Sharp-shinned Hawks, and American Kestrels; Table 2) and was more protracted for other species (e.g., Cooper's Hawks and Northern Harriers; Table 2).

Table 2

Median, peak, 80% bulk passage dates, and high single-day southbound counts for all diurnal migrant raptor species with more than 20 southbound sightings per yr at Curry Hammock State Park, Florida Keys, FL, 2000–04.


Comparisons with other Migration Count Sites

Count totals for many species in the Florida Keys were comparable to totals from nine other hawk migration count sites representative of the major known hawk migration concentration points in North America (Table 3). Mean annual Florida Keys count totals for Peregrine Falcons were considerably higher than counts in any other region of North America. Totals in the Florida Keys for Ospreys, American Kestrels, and Merlins were second only to north Atlantic coast hawkwatches (represented here by Cape May) in the United States and Canada. Florida Keys count totals for Northern Harriers were similar to totals from several of the other major concentration points for this species in North America. In the southern U.S., count totals for Sharp-shinned Hawks were higher than the other major coastal concentration point at Smith Point on the upper Texas coast. Count totals for Cooper's Hawks in the Florida Keys were lower than totals from many other sites, reflecting the position of this site near the southern limit of their wintering range in the eastern U.S. Count totals for Broad-winged Hawks in the Florida Keys, although substantial, did not come close to rivaling the large concentrations of this species counted at many other locations, particularly those along the west side of the Gulf of Mexico.

Table 3

Mean count totals (1999–2004 unless otherwise specified in footnotes) for 10 major raptor migration count sites within the Nearctic-Neotropical migration system for the eight species of diurnal raptors that are common during fall migration in the Florida Keys.



The Florida Keys are a migration concentration point of similar magnitude to other major, well-known raptor migration sites in North America, such as Cape May, New Jersey, Hawk Mountain, Pennsylvania, or the Goshute Mountains, Nevada. Count totals for Peregrine Falcons in the Florida Keys were considerably higher than totals from any other site in North America. If continued for the long term, raptor migration counts in the Florida Keys will provide high-quality, cost-effective data for monitoring Peregrine Falcon population trends and population trends for seven other common species of diurnal migrant raptors. The value of any given raptor migration count site for monitoring long-term population trends may be evaluated relative to: (1) the number of raptors counted at that site compared with the size of the breeding population that is to be monitored (Bart and Ralph 2005); (2) the coefficient of variation of annual counts, which contributes to statistical power to estimate long-term trends (Lewis and Gould 2000); and (3) the geographic representation (i.e., source areas) of migrants that are counted each year at that site. In each of these categories, long-term counts at Curry Hammock State Park can provide important data for monitoring North American raptor population trends.

White et al. (2002) summarized estimates of the number of breeding Peregrine Falcon pairs in North America as 5650–7000. This estimate included the U.S.A., Canada, Greenland, and Mexico. Previous migration banding and telemetry data suggested that individuals of the pealei subspecies and individuals of the anatum subspecies in the Pacific Northwest, Rocky Mountains, Southwestern U.S., and Mexico most likely do not migrate through the Florida Keys (White et al. 2002). Thus, the potential source population for Florida Keys migrant Peregrine Falcons may be defined as all individuals of the tundius or anatum subspecies from Alaska, Canada, Greenland, or the U.S.A. east of the Mississippi River (referred to hereafter as the “boreal and eastern U.S. population”). The boreal and eastern U.S. population was estimated as 3434–4360 pairs by White et al. (White et al. 2002). Given these breeding population estimates and recent productivity estimates of 1.3–2.5 young per pair (White et al. 2002), I calculated that approximately 7345–17 500 young peregrines are produced in North America each year (which includes 4464–10 900 within the boreal and eastern U.S. population). White et al. (2002) suggested that there was approximately one nonbreeding adult “floater” per breeding pair. Therefore, by combining breeding adults, adult floaters, and young-of-the-year, I estimated that total Peregrine Falcon population size in North America peaks at 24 295–38 500 individuals at the end of each breeding season and the boreal and eastern U.S. population peaks at 14 765–23 982. From the population estimates above and our mean annual count of 1908 peregrines, it appears that approximately 5.0–7.9% of all North American Peregrine Falcons (and 8.0–12.9% of all boreal and eastern U.S. peregrines) were counted in the Florida Keys each fall. Similarly robust population estimates do not exist for other species of diurnal raptors.

For the eight common migrant species in the Florida Keys, annual coefficients of variation for counts ranged from 26%–37% (with the exception of Cooper's Hawks, which had a CV of 57%). These CVs were lower than those for many of the raptor migration site and species combinations reported in Lewis and Gould (2000) and counts for seven of the eight common migrant species in the Florida Keys also met the statistical power criteria for monitoring long-term trends (80% power to detect a 50% decline over 20 years with a level of significance of 0.10, using a two-tailed test) that was suggested by Bart et al. (2004). One possible reason for the relatively low CVs of annual counts observed in the Florida Keys is that raptors in the Florida Keys may remain detectable regardless of wind direction at this site because the landmass is so narrow and raptors that are following the island chain can be seen offshore in either direction, regardless of which direction they are drifting offshore due to local winds. In contrast, coastal sites in areas with alternate inland routes may only detect raptors on days where wind conditions concentrate the flight of migrating raptors along the shore. On days with on-shore winds at other locations, many other sites may not detect raptors as their flight-lines shift inland. Thus, a reduced effect of wind direction on detectability may play a part in the relatively low CVs of annual counts in the Florida Keys.

Data from the Florida Keys will contribute to the Raptor Population Index (RPI), a collaborative effort by HWI, Hawk Migration Association of North America, Hawk Mountain Sanctuary, and several other partners that is designed to pool data from a network of raptor migration count sites to estimate population trends for North American raptors (MacLeod 2004). Given the large volume of raptor migration and the unique geographic position of the Florida Keys (far to the south and east of the other major hawkwatch locations in North America), count data from Curry Hammock State Park will provide important data to RPI. The far southern location of this site relative to the breeding distributions of all eight common migrant species ensures that counts in the Florida Keys will contribute population trend data for at least some individuals that are not counted at northern hawkwatch sites during fall migration, thus improving the spatial coverage of this continental network of raptor migration count sites designed to document North American raptor population trends.


I would like to thank each of the outstanding full-time observers that worked at the Curry Hammock hawkwatch between 1999–2004: Corrie Borgman, Stijn Brand, Karen Johnson, Derek Lovitch, Jose Antonio Martinez Climent, Mark Hedden, Frank Mayer, Tiffany Musgrove, Jen Ottinger, Gerard Phillips, and Paula Shannon. Funding for the migration count was provided by the Florida Fish and Wildlife Conservation Commission (FWC). HWI would like to specifically thank Robin Boughton, Tim Breen, Lara Coburn, David Cook, Stuart Cumberbatch, Michael Evans, Alex Kropp, Karl Miller, Brian Millsap, Laura Phillips, Keith Singleton, George Wallace, and Ricardo Zambrano of FWC. Additional funding for the count was provided by the Florida Keys Audubon Society, the Broward County Audubon Society, and donations from the members of HWI. Curry Hammock State Park has been a great partner for this project, particularly rangers David Huntt and Craig Johnson, park manager Kathy Close, and regional biologist Renate Skinner. Thanks to Audubon of Florida's Tavernier Science Center staff, particularly Jerry Lorenz. I would like to thank several past and present staff of HWI: Paul Grindrod, Howard Gross, Evelyn Johnson, and Jeff Smith. I am grateful to Steve Hoffman, the founder of HWI, for inspiring my interest in hawk migration and encouraging me to develop a long-term raptor migration monitoring program in the Florida Keys.

Literature Cited


J. Bart and C. J. Ralph . 2005. Suggestions for establishing a network of landbird migration monitoring sites. 701–704. in C. J. Ralph and T. R. Rich , editors. eds. Bird conservation implementation and integration in the Americas: proceedings of the third international Partners in Flight conference, 2002. Volume 1. Gen. Tech. Rep. PSW-GTR-191. Pacific Southwest Research Station, USDA Forest Service. Albany, CA U.S.A. Google Scholar


J. Bart, K. P. Burnham, E. A. Dunn, C. M. Francis, and C. J. Ralph . 2004. Goals and strategies for estimating trends in landbird abundance. J. Wildl. Manage 68:611–626. Google Scholar


J. C. Bednarz and P. Kerlinger . 1989. Monitoring hawk populations by counting migrants. 328–342. in B. G. Pendleton, M. N. LeFranc Jr, M. B. Moss, C. E. Ruibal, M. A. Knighton, and D. L. Krahe , editors. eds. Proceedings of the northeast raptor management symposium and workshop. Natl. Wildl. Fed. Washington, DC U.S.A. Google Scholar


J. C. Bednarz, D. Klem, L. J. Goodrich, and S. E. Senner . 1990. Migration counts of raptors at Hawk Mountain, Pennsylvania, as indicators of population trends, 1934–1991. Auk 113:329–338. Google Scholar


C. B. Brashear and P. K. Stoddard . 2001. Fall raptor migration through the Florida Keys with special focus on the Peregrine Falcon. Final Report. Florida Fish and Wildlife Conservation Commission. Tallahassee, FL U.S.A. Google Scholar


T. J. Cade, J. H. Enderson, C. G. Thelander, and C. M. White [EDs.] . 1988. Peregrine Falcon populations: their management and recovery. The Peregrine Fund. Boise, ID U.S.A. Google Scholar


R. Carson 1962. Silent spring. Houghton Mifflin. Boston, MA U.S.A. Google Scholar


W. S. Clark and B. Wheeler . 2001. Hawks of North America, 2nd edition. Houghton Mifflin. Boston, MA U.S.A. Google Scholar


H. N. Darrow 1983. Late fall movements of Turkey Vultures and hawks in the Florida Keys. Fla. Field Nat 11:35–39. Google Scholar


E. H. Dunn 2005. Counting migrants to monitor bird populations: state of the art. 712–717. in C. J. Ralph and T. R. Rich , editors. eds. Bird conservation implementation and integration in the Americas: proceedings of the third international Partners in Flight conference, 2002. Volume 1. Gen. Tech. Rep. PSW-GTR-191. Pacific Southwest Research Station, USDA Forest Service. Albany, CA U.S.A. Google Scholar


E. H. Dunn, B. L. Altman, J. Bart, C. J. Beardmore, H. Berlanga, P. J. Blancher, G. S. Butcher, D. W. Demarest, R. Dettmers, W. C. Hunter, E. E. Iñigo-Elias, A. O. Panjabi, D. N. Pashley, C. J. Ralph, T. D. Rich, K. V. Rosenberg, C. M. Rustay, J. M. Ruth, and T. C. Will . 2005. High priority needs for range-wide monitoring of North American landbirds. Partners in Flight Technical Series No. 2. Partners in Flight. (last accessed 15 June, 2006).  Google Scholar


P. D. Dunne, D. Sibley, and C. Sutton . 1988. Hawks in flight. Houghton Mifflin. Boston, MA U.S.A. Google Scholar


J. H. Enderson 1969. Peregrine and prairie falcon life tables based on band-recovery data. 505–509. in J. J. Hickey , editor. ed. Peregrine falcon populations: their biology and decline. University of Wisconsin Press. Madison, WI U.S.A. Google Scholar


J. H. Enderson, W. Heinrich, L. Kiff, and C. M. White . 1995. Population changes in North American peregrines. Trans. N. Am. Wildl. Nat. Resour. Conf 60:142–161. Google Scholar


M. R. Fuller 1996. Forest raptor population trends in North America. 167–208. in R. M. DeGraaf and R. I. Miller , editors. eds. Conservation of faunal diversity in forested landscapes. Chapman Hall. New York, NY U.S.A. Google Scholar


M. R. Fuller and A. Mosher . 1987. Raptor survey techniques. 37–65. in B. A. Giron Pendleton, B. A. Millsap, K. W. Cline, and D. M. Bird , editors. eds. Raptor management techniques manual. Natl. Wildl. Fed. Washington, DC U.S.A. Google Scholar


D. S. Heintzelman 1986. The migration of hawks. Indiana University Press. Indianapolis, IN U.S.A. Google Scholar


J. J. Hickey 1969. Peregrine Falcon populations: their biology and decline. University of Wisconsin Press. Madison, WI U.S.A. Google Scholar


S. W. Hoffman 1985. Raptor movements in inland western North America: a synthesis. 325–338. in M. Harwood , editor. ed. Proceedings of Hawk Migration Conference IV, Rochester, NY. Hawk Migration Association of North America. Kempton, PA U.S.A. Google Scholar


S. W. Hoffman and J. P. Smith . 2003. Population trends of migratory raptors in western North America, 1977–2001. Condor 105:397–419. Google Scholar


W. Hoffman and H. Darrow . 1992. Migration of diurnal raptors from the Florida Keys into the West Indies. Hawk Migration Studies 17:7–14. Google Scholar


D. J. T. Hussell and L. Brown . 1992. Population changes in diurnally-migrating raptors at Duluth, Minnesota (1974–1989) and Grimsby, Ontario (1975–1990). Ontario Ministry of Natural Resources. Maple, Ontario, Canada. Google Scholar


D. J. T. Hussell and C. J. Ralph . 2005. Recommended methods for monitoring change in bird populations by counting and capture of migrants. N. Am. Bird Bander 30:7–20. Google Scholar


P. Kerlinger 1989. Flight strategies of migrating hawks. University of Chicago Press. Chicago, IL U.S.A. Google Scholar


D. A. Kirk and C. Hyslop . 1998. Population status and recent trends in Canadian raptors: a review. Biol. Conserv 83:91–118. Google Scholar


S. A. Lewis and W. R. Gould . 2000. Survey effects on power to detect trends in raptor migration counts. Wildl. Soc. Bull 28:317–329. Google Scholar


I. MacLeod 2004. The raptor population index: a scientific system to monitor North American raptor populations. Hawk Migration Studies 24:7–9. Google Scholar


R. Mesta 1999. Final rule to remove the American Peregrine Falcon from the federal list of endangered and threatened wildlife, and to remove the similarity of appearance provision for free-flying peregrines in the conterminous United States. Federal Register 64:46541–46558. Google Scholar


B. A. Millsap, P. L. Kennedy, M. A. Byrd, G. Court, J. H. Enderson, and R. N. Rosenfield . 1998. Review of the proposal to de-list the American Peregrine Falcon. Wildl. Soc. Bull 26:522–538. Google Scholar


H. C. Mueller and D. D. Berger . 1967. Wind drift, leading lines, and diurnal migrations. Wilson Bull 79:50–63. Google Scholar


H. C. Mueller, D. D. Berger, and G. Allez . 1988. Population trends in migrating peregrines at Cedar Grove, Wisconsin, 1936–1985. in T. J. Cade, J. H. Enderson, C. G. Thelander, and C. M. White , editors. eds. Peregrine Falcon populations: their management and recovery. The Peregrine Fund. Boise, ID U.S.A. Google Scholar


P. Porras-Penaranda, L. Robichaud, and F. Branch . 2004. New full season count sites for raptor migration in Talamanca, Costa Rica. Ornithol. Neotrop 15:Supplement267–278. Google Scholar


E. Ruelas Inzunza, S. W. Hoffman, L. J. Goodrich, and R. Tingay . 2000. Conservation strategies for the world's largest known raptor migration flyway: Veracruz the river of raptors. 591–596. in R. D. Chancellor and B-U. Meyburg , editors. eds. Raptors at risk: proceedings of the V world conference on birds of prey and owls. World Working Group on Birds of Prey and Owls. Berlin, Germany. Google Scholar


N. G. Smith 1980. Hawk and vulture migration in the Neotropics. 51–65. in A. Keast and E. S. Morton , editors. eds. Migrant birds in the neotropics. Smithsonian Institute Press. Washington, DC U.S.A. Google Scholar


J. P. Smith and S. W. Hoffman . 2000. The value of extensive raptor migration monitoring in western North America. 597–615. in R. D. Chancellor and B-U. Meyburg , editors. eds. Raptors at risk: proceedings of the V world conference on birds of prey and owls. World Working Group on Birds of Prey and Owls. Berlin, Germany. Google Scholar


J. P. Smith, J. Simon, S. W. Hoffman, and C. Riley . 2001. New full-season fall hawkwatches in coastal Texas. 67–91. in K. L. Bildstein and D. Klem , editors. eds. Hawkwatching in the Americas. Hawk Migration Association of North America. North Wales, PA U.S.A. Google Scholar


T. Swem and R. Ambrose . 1994. Notice of the removal of the arctic Peregrine Falcon from the list of threatened and endangered species. Federal Register 59:50794. Google Scholar


K. Titus, M. R. Fuller, and D. Jacobs . 1990. Detecting trends in hawk migration count data. 105–113. in J. R. Sauer and S. Droege , editors. eds. Proceedings for a workshop on analysis of avian population trends. U.S. Fish and Wildlife Service Biological Report 90. Google Scholar


U.S. Fish and Wildlife Service 2003. Monitoring plan for the American Peregrine Falcon, a species recovered under the Endangered Species Act. U.S. Fish and Wildlife Service, Divisions of Endangered Species and Migratory Birds and State Programs. Pacific Region, Portland, OR U.S.A. Google Scholar


B. K. Wheeler and W. S. Clark . 1995. A photographic guide to North American raptors. Academic Press. San Diego, CA U.S.A. Google Scholar


C. M. White, N. J. Klum, T. J. Cade, and W. G. Hunt . 2002. Peregrine Falcon (Falco peregrinus). in A. Poole and F. Gill , editors. eds. The birds of North America, No. 660. The Academy of Natural Sciences. Philadelphia, PA and The American Ornithologists' Union. Washington DC U.S.A. Google Scholar
Casey A. Lott "A NEW RAPTOR MIGRATION MONITORING SITE IN THE FLORIDA KEYS: COUNTS FROM 1999–2004," Journal of Raptor Research 40(3), 200-209, (1 September 2006).[200:ANRMMS]2.0.CO;2
Received: 8 June 2005; Accepted: 1 June 2006; Published: 1 September 2006

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