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9 November 2016 Variation of Male Secondary Sexual Structures and the Taxonomy of Theritas lisus and Relatives (Lepidoptera, Lycaenidae, Eumaeini)
Ananda Regina P. Martins, Christophe Faynel, Robert K. Robbins
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Abstract

We assess variation of male secondary sexual structures, genitalia, and wing patterns in Theritas lisus (Stoll) and relatives to clarify the species level taxonomy, which is needed for a phylogenetic analysis. Variation in the male ventral hindwing scent pouch was especially useful taxonomically. This pouch may be absent, present without androconia, or present with androconia in T. lisus. Despite this variation, pouch size (when present) distinguishes T. lisus from its closest relatives. As an overview, we propose distinguishing traits for a phenetic T. lisus species group, re-examine and confirm the proposed synonymies of T. hisbon and T. orsina with T. lisus and of T. photeinos with T. viresco, confirm the association of the sexes of T. lisus and T. viresco, and describe Theritas silma Martins, Faynel, & Robbins, new species, from French Guiana. CO1 mitochondrial sequences are consistent with these taxonomic results, but the recently introduced barcode index numbers (BINs) recognized too many species. A nomenclatural list summarizes the taxonomic actions in the T. lisus species group.

Ananda Regina P. Martins, Christophe Faynel, and Robert K. Robbins "Variation of Male Secondary Sexual Structures and the Taxonomy of Theritas lisus and Relatives (Lepidoptera, Lycaenidae, Eumaeini)," Proceedings of the Entomological Society of Washington 118(4), 555-573, (9 November 2016). https://doi.org/10.4289/0013-8797.118.4.555
Published: 9 November 2016
KEYWORDS
Atlides Section
barcoding DNA
DNA barcoding
órgã odorífero dorsal
órgã odorífero ventral
scent pad
scent pouch
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