Prionii Latreille, 1802: 212.

Acanthophorini Thomson, 1864: 289, 473. Type genus: Acanthophorus Audinet-Serville, 1832.

Anthracocentrus Quentin & Villiers, 1983: 82. Type species: Tithoes arabicus Thomson, 1877 (by original designation).

Tithoes arabicus Thomson, 1877: 266-267. Type locality: “Arabia: Djedda″.

Acanthophorus arabicus: Villiers, 1968: 846; Holzschuh, 1979: 293; Holzschuh, 1993: 111.

Material examined from Saudi Arabia: Hejaz, 10.XI.1945 (D.V. Fitzgerald); Asir, Wadi Lasaba, Tchama près Qunfidha, 9.XII.1945 (D.V. Fitzgerald); Hejaz, Bureiman Camp, près Jidda, 20.X.1945 (D. V. Fitzgerald) (Villiers, 1968); Asir Mts., Straße von Abha nach Gizan, km 53, Wadi Ad Dilla, 300 s.l.m., 21.IV.1976, Wittmer & Büttiker (Holzschuh, 1979); Marsa Zubeida, Al Wadj Dhuba, 18.IX.1983, W. Büttiker; Jeddah-Taif road, km 102, 10.XII.1982, W. Büttiker; Wadi Jizan, 10.XI.1974, P.D. Manser; Hakimah, 85 s.l.m., 12-15.X.1979, W. Büttiker; Usfan, 29.XI.1983, W. Büttiker (Holzschuh, 1993); Asir, Al Magardah, Wadi Yabah, 19°14.911′N 41°47.255′E, 402 s.l.m., 11.X.2013, at light; Asir, Nanira, 19°34.799′N 41°39.750′E, 440 s.l.m., 3.XI.2013 (KSMA); Bahah prov., 2 km E Nawan, 117 s.l.m., 19°32′48″N 41°11′34″E, 31.III.2017, several dead specimens under the bushes of Tamarix sp. [Tamaricaceae], P. Rapuzzi lgt; idem 12.IV.2019 (CPR).

Distribution: Algeria, Egypt, Israel, Jordan, Chad, Sudan, Ethiopia, Eritrea, Djibouti, Somalia (Adlbauer & Beck, 2015), Lybia (NHMP). Arabian Peninsula: Saudi Arabia, Yemen, Oman, Bahrain (Adlbauer & Beck, 2015).

Tithoes Thomson, 1864: 289, 473. Type species: Prionus maculatus Fabricius, 1793 (by original designation).

Tithoes confinis Laporte, 1840: 395. Type locality: “Sénégal″.

Acanthophorus confinis: Holzschuh, 1993: 111-112.

Material examined from Saudi Arabia: 81 km S of Biljurshi, 2000 s.l.m., VIII.1979, G. Vogel (Holzschuh, 1993); Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 2.IX.2015, at light (KSMA); Asir Mts., Khamis Mushayt env., 600 s.l.m., 5.V.1972, J. Lorenc (Jiri Lorenc collection, Chomutov, Czech Republic).

Distribution: Mauritania, Senegal, Gambia, Guinea, Mali, Chad, Sudan, Ethiopia, Eritrea, Djibouti, Somalia, Cameroon, Central African Republic, Gabon, Congo, Uganda, Kenya, Tanzania, Zambia, Malawi, Zimbabwe, Mozambique, Botswana, Angola, Namibia, Republic of South Africa. Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015), Yemen (Holzschuh, 1993).

Cantharocnemitae Thomson, 1861: 274. Type genus: Cantharocnemis Audinet-Serville, 1832.

Cantharocnemis Audinet-Serville, 1832: 132. Type species: C. spondyloides Audinet-Serville, 1832 (by monotypy).

Cantharocnemis spondyloides Audinet-Serville, 1832: 132. Type locality: “Sénégal″.

Cantharocnemis spondyloides: Villiers, 1968: 847; Holzschuh, 1993: 114-115.

Material examined from Saudi Arabia: Sharb Suwaidara, 21.V.1938, H.St.J.B. Philby; “Arabia″ W. Thesiger (Villiers, 1968); Fayfa, 29.IV.1982, collector not given; Jeddah, Munidatna, XI.1939, A.C. Trott; Shiara, I.1946, L.A. Tillin (Holzschuh, 1993); Srat Adibh, 25.VIII.2012; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 22.IV.2014, at light; idem, 2.IX.2015; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 2.IX.2015, at light (KSMA).

Distribution: Egypt, Mauritania, Senegal, Guinea, Mali, Burkina Faso, Niger, Sudan, Ethiopia, Eritrea, Djibouti, Somalia, Uganda, Kenya, Tanzania, Zimbabwe, Botswana, Namibia, Republic of South Africa. Arabian Peninsula: Saudi Arabia, Yemen, Oman (Adlbauer & Beck, 2015).

Cantharocnemis subg. Paracantharocnemis Plavilstshikov, 1933: 113-118. Type species: Cantharocnemis (Paracantharocnemis) strandi Plavilstshikov, 1933 (by original designation).

Cantharocnemis (Paracantharocnemis) strandi Plavilstshikov, 1933: 113-114. Type locality: “Arabia mer.: Yemen, Sanaa″ (San′a′).

Cantharocnemis strandi: Holzschuh, 1979: 293; Holzschuh, 1993: 115.

Material examined from Saudi Arabia: 81 km S of Biljurshi, 2000 s.l.m., VIII.1979, G. Vogel (Holzschuh, 1993); Asir Mts., Kamis-Mushayt, August 1971, leg. Di Lenardo (Holzschuh, 1979); Abha, Raydah, 18°11.618′N 42°24.420′E, 1772 s.l.m., 31.VII.2015, at light; Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 2.IX.2015, at light (KSMA).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen (Drumont, 2004).

Cantharocnemis subg. Cantharoctenus Westwood, 1866: 134. Type species: Cantharocnemis (Cantharoctenus) burchelli Westwood, 1866 (by monotypy).

Cantharocnemis filippovi Plavilstshikov, 1933: 263-264. Type locality: “Yemen, Mafchak″.

Material examined from Saudi Arabia: Bahah prov., Jabal al A′la, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III.2017, ex larva in Grewia sp. [Malvaceae], emerged 28.VIII-26.IX.2017, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, ex larva in Grewia sp., emerged 1.XI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality), Saudi Arabia (new record).

Description of the female: The female differs from the male mainly in the antennal shape. The antennae are shorter and thinner than in the male and show only one small tooth at the apex of each segment starting from the third. The antennae in the females are made of 22 segments instead of 46 in the males. The antennae reach the base of the elytra when extended posteriorly, while they reach the apical quarter of the elytra in the males. The legs are shorter and slightly slender than in the males and they possess only few teeth on the external side of the tibiae.

Macrotomini Thomson, 1860: 290, 312.

Macrotoma Audinet-Serville, 1832: 1: 137. Type species: Prionus serripes Fabricius, 1793 (designated by Thomson, 1860).

Prionus palmatus Fabricius, 1793: 249. Type locality: “Guinea″.

Macrotoma palmata Holzschuh, 1979: 293; Holzschuh, 1993: 111.

Material examined from Saudi Arabia: Asir Mts., über Bisha nach Kamis-Mushayt, Anfang August 1971, leg. Di Lenardo (Holzschuh, 1979; Holzschuh, 1993); Abha, 15.VII.1981 (KSMA); Asir Mts., 10.VIII.1992, J. Lorenc (Jiri Lorenc collection, Chomutov, Czech Republic); Asir Mts, 10.VII.1990 (CPR).

Distribution: Morocco, Algeria, Egypt, Mauritania, Senegal, Gambia, Mali, Chad, Sudan, Ethiopia, Eritrea, Djibouti, Somalia, Guinea, Sierra Leone, Ivory Coast, Ghana, Togo, Benin, Nigeria, Cameroon, Central African Republic, Equatorial Guinea, Gabon, Congo, Uganda, Rwanda, Burundi, Kenya, Tanzania, Zambia, Malawi, Zimbabwe, Mozambique, Botswana, Angola, Namibia, Republic of South Africa, Mauritius. Arabian Peninsula: Saudi Arabia, Yemen (Adlbauer & Beck, 2015).

Prioniens, Prionii Latreille, 1802: 212.

Polyarthron (subg.) Monocladum Pic, 1898: 27. Type species: Prionus aegyptiacus Guérin-Méneville, 1844 (designated by Sama, 2008: 220).

Monocladum aegyptiacum ssp. arabicum Villiers, 1961: 449. Type locality: “Hadramaut″ (Yemen).

Monocladum aegyptiacus ssp. arabicum: Holzschuh, 1979: 293.

Material examined from Saudi Arabia: Najran, 23-24.VII.1938, H.St.J.B. Philby (Holzschuh, 1993).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen (Holzschuh, 1993).

Monocladum aegyptiacus granulipenne Holzschuh, 1993: 112-113. Nom. nov. for Prionus unipectinatus ssp. arabicus Fuchs, 1969: 381 (nec Villiers, 1961). Type locality: “Sokna (Thima), 200 s.l.m., Yemen″ [junior homonym].

Monocladum aegyptiacus ssp. aegyptiacus: Villiers, 1968: 846; Holzschuh, 1979: 293.

Material examined from Saudi Arabia: Hejaz, 10.XI.1945 (D.V. Fitzgerald); Côte Ouest, près Al Lith, 6.XII.1945 (D.V. Fitzgerald); Nord Nedjed, 19.II.1946 (D. V. Fitzgerald) (VILLIERS, 1968); Umgeb. Riyadh, Bahara, 26.X.1976, W. Büttiker (Holzschuh, 1979); Makkah, 6.III.1981, W. Büttiker; Wadi Ilyab, 20°07′N 40°57′E, 10-11.XI.1983, W. Büttiker; Harithi, 21°18′N 10°18′E, 1910 s.l.m., W. Büttiker; Wadi Jizan, 16.XI.1976; idem 26.X.1978, Filipponi; Husayniyah, 23°50′N 38°53′E, 23.III.1936, H.St.J.B. Philby; Hadda′, 21°27′N 39°34′E, 25.XI.1938, H.St.J.B. Philby; Asir, Sabiya, 17°10′N 42°30′E, 1.XI.1946, G. Popov; idem, 8.VII.1945, A.R. Waterston; Ranya, 21°30′N 43°00′E, 23.V.1936, H.St.J.B. Philby (Holzschuh, 1993).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen (Holzshuh, 1993).

Polyarthron Audinet-Serville, 1832: 189. Type species: Prionus pectinicornis Fabricius, 1795 (by monotypy).

Polyarthron philbyi Villiers, 1968: 846-847. Type locality: “Arabie: Dalgan, à l′est de Quai′iya″.

Polyarthron philbyi: Holzschuh, 1993: 114.

Material examined from Saudi Arabia: Dalgan, à l′est de Quai′iya, 18.XI.1949, H.St.J.B. Philby (holotype); Wadi Ratimah, 14.X.1977, W. Büttiker; Ummal Jamajin, 26°55′N 45°22′E, 550 s.l.m., 26-27.X.1986, W. Büttiker (Holzschuh, 1993); Muzahimiyah, Al Khararah, 24°23′01″N 46°14′14″E, 3.X.2011, at light; idem 19.IX.2011; Rawdat Khorim, 25.IX.2012 (KSMA).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh, 1993).

Cerambycinae Latreille, 1802: 211.

Callichrominae Swainson & Shukard, 1840: 293-294. Type genus: Callichroma Latreille, 1817.

helymaeus Thomson, 1864: 179. Type species: helymaeus cyanipennis Thomson, 1864 (by original designation and monotypy).

helymaeus pedestris Pascoe, 1878: 370. Type locality: “Yemen″.

Material examined from Saudi Arabia: Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°22′01″E, 16.IV.2016, ex larva in Grewia sp. [Malvaceae], emerged 30.VI-27.VII.2016, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality), Saudi Arabia (new record).

helymaeus signaticollis Pascoe, 1878: 370. Type locality: “Yemen″.

Material examined from Saudi Arabia: Al Baha, Shada Al Ala, 19°51.762′N 41°18.089′E, 1225 s.l.m., 3.VI.2014 (KSMA); Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, 1 specimen on flowers of Acacia sp. [Fabaceae], P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality), Saudi Arabia (new record).

Callidiopsides Lacordaire, 1869: 340. Type genus: Callidiopis Thomson, 1864.

Prosype Thomson, 1864: 248. Type species: Oeme filiformis Buquet, 1859 (by original designation).

Prosype juniperi Holzschuh, 1993: 120-122. Type locality: “Saudi Arabia: Asir Mts., Jebel Sudan nr Abha″.

Material examined from Saudi Arabia: Asir Mts., Jebel Suda near Abha, 9.IV.1983, emerged from Juniperus procera Hochst. & Endl., 1847, 31.VIII.1983, C. Holzschuh (holotype); idem, emerged between 8.VII and 7.VIII.1983 (Holzschuh, 1993); Abha, Raydah, 18°11.679′N 42°23.691′E, 1851 s.l.m., 26.IV.2014, at light (KSMA); Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 16.IV.2016, ex larva in Rosa sp. [Rosaceae], emerged 21.IV, 29.IV, 9.VI, 21.VII, 12.X.2016, P. Rapuzzi lgt; idem, ex larva in Juniperus procera [Cupressaceae], emerged 5.VI.2016, P. Rapuzzi lgt; idem, emerged 1-19.VI.2017; Bahah prov., Bani Hassan, 2293 s.l.m., 20°03′02″N 41°26′47″E, 30.III.2017, ex larva in Juniperus procera, emerged 10.VII.2017, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (type locality), Yemen (Danilevsky, 2020).

Cerambycini Latreille, 1802: 211. Type genus: Cerambyx Linné, 1758.

Anarchambyx Sama, 2007: 101. Type species: A. pipposamai Sama, 2007 (by original designation).

Anarchambyx pipposamai Sama, 2007: 103. Type locality: “Yemen: Mabar″.

Material examined from Saudi Arabia: Al Bahah prov., Wadi Tourabah, 20°11′16″N 41°18′38″E, 1812 s.l.m., 6.IV.2019, ex larva in Acacia sp., emerged 25.V-25.VI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality); Saudi Arabia (new record).

Pachydissus (Derolus) Gahan, 1891: 26. Type species: D. arciferus Gahan (original designation).

Derolus asiricus Holzschuh, 1993: 117-118. Type locality: “Saudi Arabia: Asir Mts., Shaqiq Sharman″.

Material examined from Saudi Arabia: Asir Mts., Shaqiq Sharman, 8.IV.1983, fully developed in its pupal chamber in a dead lactiferous climber, C. Holzschuh (typ. loc.); idem, reared from larvae and emerged from 20.VII to 24.VIII.1983; Fayfa, at light, 27-31.III.1983, C. Holzschuh; Fayfa, 27.IV.1982, collector not given; Wadi Gaanah, 13-14.II.1980, W. Büttiker; Harithi, 21°18′N 40°18′E, 18-19.IV.1985, W. Büttiker (Holzschuh, 1993); Abha, Wadi Rida, 18°11′749″N 42°23′345″E, 1614 s.l.m., 24.II.2014, at light; Abha, Wadi Tourabah, 20°14.369′N 41°15.234′E, 1757 s.l.m., at light; Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 21.II.2014, at light; idem, 26.IV.2014; Abha, Raydah, 18°11.884′N 42°24.435′E, 2387 s.l.m., 26.IV.2014, at light; Abha, Raydah, 18°11.679′N 42°23.691′E, 1851 s.l.m., 6.VI.2014, at light; Abha, Raydah, 18°11.766′N 42°24.315′E, 2285 s.l.m., 26.IV.2014, at light; Abha, Raydah, 18°12.265′N 42°24.744′E, 2820 s.l.m., 26.IV.2014, at light; idem, 7.VI.2014; Abha, Raydah, 18°12.095′N 42°24.536′E, 2578 s.l.m., 6.VI.2014, at light; Abha, Raydah, 18°11.618′N 42°24.420′E, 1772 s.l.m., 8.VI.2014, at light; Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 27.VII.2015, at light; idem, 3.VI.2014; idem, 21.IV.2014; Abha, Raydah, 18°11.7749′N 42°23.345′E, 1614 s.l.m., 31.VII.2015, at light; idem, 4.III.2015; idem 26.IV.2014; Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m., 23.VIII.2015, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light; Al Baha, Shada Al Ala, 19°50.575′N 41°18.691′E, 1666 s.l.m., 3.VI.2015, at light; idem, 2.IX.2015; Al Baha, Shada Al Ala, 19°50.066′N 41°18.037′E, 1325 s.l.m., 5.V.2014, at light; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 2.IX.2014, at light; Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 21.IV.2014, at light (KSMA); Abha prov., Raedah res., 2809 s.l.m., 18°21′14″N 42°24′43″E, 16.IV.2016, ex larva in Euryops arabica Steud., 1837 [Asteraceae], emerged 15.VIII.2016, P. Rapuzzi lgt; idem, emerged 15.IV.2017; Asir prov., Jebel Soudah, Raydah res., 18°12′14″N 42°24′43″E, 2810 s.l.m., 11.IV.2019, in pupal cell of Dodonaea viscosa Jaq., 1760 [Sapindaceae], P. Rapuzzi lgt; Asir prov., wadi Mashwas, 18°10′06″N 42°22′01″E, 1289 s.l.m., 13.IV.2019, pupal cell of Cadia purpurea (G. Piccoli) Aiton (1789) [Fabaceae], P. Rapuzzi lgt; Al Baha prov., Wadi Tourabah, 1830 s.l.m., 20°11′36″N 41°17′50″E, 12.IV.2016, pupal cell on Prunus sp. [Rosaceae], P. Rapuzzi lgt; idem, emerged 4.XI.2016; Al Baha prov., 10 km NNW Al Makhwah, 630 s.l.m., 19°50′47″N 41°22′40″E, 31.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1965 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, adults under the bark of Ficus sp. [Moraceae], P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Bahah prov., Shada al Ala, 1692 s.l.m., 19°50′37″N 41°18′40″E, 8.IV.2019, ex larva from Ficus sp., P. Rapuzzi leg.; Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°22′01″E, 16.IV.2016, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 13.IV.2017, at light, P. Rapuzzi lgt; Asir prov., Jebel Soudah, 1632 s.l.m., 18°11′45″N 42°23′21″E, 2.IV.2017, sugar traps, P. Rapuzzi lgt; Asir prov., Jebel Soudah, Raydah vill. 18°11′42″N 42°23′04″E, 1572 s.l.m., 11.IV.2019, dead inside a spider net, P. Rapuzzi lgt; Bahah prov., Wadi Tourabah, E of An Na′amah, 1826 s.l.m., 20°11′01″N 41°18′42″E, 30.III.2017, in pupal cell on Kleinia anteuphorbium (L.) DC., 1838 [Asteraceae], P. Rapuzzi lgt; Bahah prov., Wadi Tourabah, E of An Na′amah, 1812 s.l.m., 20°11′16″N 41°18′38″E, 6.IV.2019, P. Rapuzzi lgt; idem, in pupal cell on Pistacia sp. [Anacardiaceae], 6.IV.2019, P. Rapuzzi lgt; idem ex larva from Pistacia sp., emerged 19.V.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh, 1993).

Derolus incultus ssp. yemenensis Villiers, 1977: 166-167. Type locality: “Nord-Yémen (Taizz)″.

Material examined from Saudi Arabia: Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 13.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Yemen (type locality.); Oman, Jebel al Akhdar, N of Birkat al Mouz, 2.III.2008, ex larva from Moringa peregrina (Forssk., 1911) (Moringaceae), emerged 22.VII.2008, D. Gianasso leg. (CGS) (new record); Saudi Arabia (new record).

Derolus martini ssp. hayekae Villiers, 1968: 847-848. Type locality: “Yémen: Taïz″.

Derolus martini ssp. hayekae Holzschuh, 1979: 293; Holzschuh, 1993: 117.

Material examined from Saudi Arabia: Taif, Ende Mai 1917, leg. Di Lenardo (Holzschuh, 1979); Fayfa, at light, 27-31.III.1983, C. Holzschuh; idem, emerged from Ficus (Holzschuh, 1993); Fifa, 27.IV.1982; Asir, Muhayil, Wadi Hali, 18°30.121′N 42°02.219′E, 440 s.l.m., 11.II.2016; Jazan, Fifa, 26.III.2014, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 3.VI.2014, at light; idem, 20.IV.2014; idem, 21.IV.2014; idem, 23.IV.2014; idem, 24-25. IV.2014; idem, 23.III.2015; idem, 1.III.2015; Abha, Raydah, 18°13.347′N 42°24.133′E, 2717 s.l.m., 25.IV.2014, at light; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 21.IV.2014, at light; Abha, Raydah, 18°12.265′N 42°24.744′E, 2820 s.l.m., 26.IV.2014, at light; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 3.VI.2014, at light; idem, 21.IV.2014; Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 3.VI.2014, at light; idem, 21.IV.2014; idem, 27.VII.2014; Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m., 23.VIII.2015, at light; idem, 3.VI.2014; idem, 21.IV.2014; Abha, Raydah, 18°12.095′N 42°24.536′E, 2578 s.l.m., 6.VI.2014, at light; Al Baha, Shada Al Ala, 19°50.575′N 41°18.691′E, 1666 s.l.m., 3.VI.2015, at light; Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 21.IV.2014, at light; Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 27.IV.2014, at light (KSMA); Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Ziziphus sp. [Rhamnaceae], emerged 14.IV-2.VI.2016, P. Rapuzzi lgt; Al Baha, Mekwah, Shada al Ala, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III-1.IV.2017, sugar traps, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1965 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, adults under the bark of Ficus salicifolia Vahl., 1790 [Moraceae], P. Rapuzzi lgt; Bahah prov., Shada al Ala, 1692 s.l.m., 19°50′37″N 41°18′40″E, 8.IV.2019, ex larva in Ziziphus spinachristi (L.) Desf., 1798 [Rhamnaceae], emerged 20.VII.2019, P. Rapuzzi leg.; idem, ex larva from Ficus sp., emerged 29.VII.2019, P. Rapuzzi lgt; Bahah prov., Wadi Tourabah, E of An Na′amah, 20°11′01″N 41°18′42″E, 1826 s.l.m., 30.III.2017, ex larva in Kleinia anteuphorbium, emerged 20.IV.2017, P. Rapuzzi lgt; Bahah prov., Thee Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh, 1979), Yemen (type locality).

Derolus thesigeri Villiers, 1968: 848. Type locality: “Jebel Quarra″ (Saudi Arabia).

Material examined from Saudi Arabia: Jebel Quarra, 2.II.1947, W. Thesiger, NHMUK (holotype); Bahah Wadi Tourabah, E of An Na′amah, 20°11′01″N 41°18′42″E, 30.III.2017, ex larva in Searsia retinorrhoea (Steud. ex Oliv.) Moffett, 2002 [Anacardiaceae], emerged 10.VI.2017, P. Rapuzzi lgt; idem, emerged 28.VIII.2017 (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Villiers, 1968).

Derolus vastus Holzschuh, 1993: 118-120. Type locality: “Saudi Arabia: Hofuf″.

Derolus mauritanicus ssp. iraniensis Holzschuh, 1979: 293.

Material examined from Saudi Arabia: Umgeb. Ryadh, Wadi Hofuf, 9.III.1977 W. Büttiker (under Derolus mauritanicus ssp. iraniensis Pic, 1956 in Holzschuh, 1979); Hofuf, 150 s.l.m., 12-13.IV.1983, dead in a spider′s web on the wall of a house, C. Holzschuh (holotype); Hofuf, 9.III.1977, W. Büttiker (Holzschuh, 1993).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh, 1993).

Neoplocaederus Sama, 1991: 123. Type species: Plocaederus cyanipennis Thomson, 1860 (original designation).

Cerambyx denticornis Fabricius, 1801: 271. Type locality: “Guinea″.

Plocaederus denticornis: Villiers, 1968: 847.

Plocaederus atlanticus: Holzschuh, 1993: 117.

Material examined from Saudi Arabia: Dhufar, Jebel Qara, Wadi Ghet, XI.1943, D.V. Fitzgerald; près Mecca, 12.VIII.1944, A.R. Waterston (Villiers, 1968); J. Lebaba, 13.II.1985, W. Büttiker; Ad Darb, 13.II.1985, W. Büttiker; Wadi Juwa, 8.II.1986, J. Grainger; Hesua, 610 s.l.m., 27.IX.1981, W. Büttiker; Jizan, 25-26.III.1983, fully developed in cocoons in a dead Commiphora opobalsamum (L.) Engl., 2007 [Burseraceae] tree, emerged from 9.VII to 18.IX.1983, C. Holzschuh (under Plocaederus atlanticus Rungs, 1952 in Holzschuh, 1993); Abha, Raydah, 18°11.766′N 42°24.315′E, 1897 s.l.m., 26.IV.2014, at light (KSMA).

Distribution: Senegambia, Guinea Bissau, Guinea, Sierra Leone, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Chad, Sudan, Djibouti, Somalia, Equatorial Guinea, Congo, Ethiopia, Kenya, Tanzania, Mozambique, Zimbabwe, Angola, Republic of South Africa (Adlbauer & Beck, 2015), Mali (Villiers, 1962), Morocco. Arabian Peninsula: Saudi Arabia, Yemen (Holzschuh, 1993), Oman (Strumia et al., 2007).

Plocaederus elongatulus Holzschuh, 1993: 117. Type locality: “Saudi Arabia: 90 km SE of Khamis Mushayt″.

Material examined from Saudi Arabia: 90 km SE Khamis Mushayt, 25.III.1983, C. Holzschuh (holotype) (Holzschuh, 1993); Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m., 21.IV.2014, at light; Al Baha, Shada Al Ala, 19°50.066′N 41°18.037′E, 1325 s.l.m., 03.VI.2014, at light; idem, 23.VIII.2014; idem, 14.XI.2015; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 3.VI.2014, at light; idem, 1.III.2015; Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 2.IX.2015, at light (KSMA).

Distribution: Arabian Peninsula: Saudi Arabia (type locality).

Xenoderolus Sama, 2007: 98. Type species: Microderolus arabicus Villiers, 1968 (by original designation).

Microderolus arabicus Villiers, 1968: 848. Type locality: “Arabie: Habarut ″ (Yemen).

Microderolus arabicus: Holzschuh, 1993: 120.

Xenoderolus arabicus: Sama, 2007: 98-101.

Material examined from Saudi Arabia: Shuqayq (NW of Jizan), 0 s.l.m., 2.IV.1983, emerged from Acacia 20-21.VIII.1983, C. Holzschuh; Fayfa, running very rapidly after dusk on a Acacia sp. branch, C. Holzschuh; Makkah (Holzschuh, 1993; Sama, 2007); Al Baha, Thee Ain, 19°55.774′N 41°28.574E, 754 s.l.m., 10.III.2012; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 24-25.IV.2014, at light; idem, 3.VI.2014; idem, 12.XI.2015; Jazan, Fifa, 26.III.2014, at light; Al Baha, Shada Al Ala, 19°51.762′N 41°18.089′E, 1225 s.l.m., 3.VI.2014, at light (KSMA); Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 28 km SSE Muhayl, 809 s.l.m., 18°17′55″N 42°07′41″E, 5.IV.2017, running on Acacia sp. twigs, P. Rapuzzi lgt; Asir prov., Abha city, 2248 s.l.m., 18°14′07″N 42°30′01″E, 11.IV.2019, at light, P. Rapuzzi lgt; Makkah prov., Wadi Shoqab, 20°40′26″N 41°13′23″E, 1426 s.l.m., 5.IV.2019, ex larva from Acacia sp., emerged 15.VI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality), Saudi Arabia, Oman (Holzschuh, 1993).

Sténopérites (Cartallites) Fairmaire, 1868: 149: Type genus: Cartallum Audinet-Serville, 1834.

Certallum Dejean, 1821: 111. Type species: Saperda ruficollis Fabricius, 1787 (= Cerambyx ebulinus Linnaeus, 1767) (by monotypy).

Certallum thoracicum Sharp, 1880: 247. Type locality: “Near Jeddah″ (Saudi Arabia).

Cartallum thoracicum: Holzschuh, 1993: 127.

Material examined from Saudi Arabia: near Jeddah (holotype).

Distribution: Israel, Syria, Iraq, Iran, Turkey (Löbl & Smetana, 2010), Lebanon (Coquempot et al., 2016). Arabian Peninsula: Saudi Arabia? Recorded in error because the species has not been found on the Arabian Peninsula since the original description.

Clytaires Mulsant, 1839: 70. Type genus: Clytus Laicharting, 1784.

Chlorophorus Chevrolat, 1863: 290. Type species: Callidium annulare Fabricius, 1787 (by original designation).

Clytus capensis Laporte & Gory, 1841: 95. Type locality: “Cap de Bonne-Espérance″ (Republic of South Africa).

Chlorophorus capensis: Holzschuh, 1993: 127.

Material examined from Saudi Arabia: 90 km SE Khamis Mushayt, 25.III.1983, ex larva from Ficus sp., emerged 16.VII and 30.VIII.1983, C. Holzschuh (Holzschuh, 1993); Baha, Aldawood, 18.VI.2003, ex larva in Acacia sp. (KSMA); Bahah prov., Bani Hassan, 2293 s.l.m., 20°03′02″N 41°26′47″E, 30.III.2017, ex larva in Acacia sp., emerged 28.VII.2017, P. Rapuzzi lgt (CPR).

Distribution: Republic of South Africa (type locality), Rwanda (Duffy, 1955). Arabian Peninsula: Saudi Arabia (Holzschuh, 1993).

Graciliaires Mulsant, 1839: 99. Type genus: Gracilia Audinet-Serville, 1834.

Type species: Arabogracilia saudita sp. nov.

Diagnosis: The new genus is very similar to the cosmopolitan genus Gracilia Audinet-Serville, 1834, with which it notably shares a small body size, the sharpness of the mandibles and the general aspect. However, these two taxa differ in the shape of the eyes, which consist in two lobes connected by a single row of ommatidia in Gracilia, while these lobes are connected by several rows of ommatidia in the new genus.

Etymology: The name of the new genus is composed of the words Arabo (from the Arabian peninsula) and Gracilia (in reference to the affinity with this genus).

Description: Male. Small, head and pronotum deeply punctate. Mandibles short, curved toward apex with small subbasal tooth on inner side. Frons large, flat. Eyes complete, not divided. Antennae long and slender, exceeding the apex of elytra when extended posteriorly; third segment shorter than fourth, and 3 times longer than second; segments third to sixth with few erect black apical bristles. Pronotum about twice longer than large, constricted at base and densely punctured; sides regularly rounded. Elytra parallel-sided, slightly constricted before middle; shoulders slightly rounded; disc shallowly depressed longitudinally beside slightly raised suture; pubescence short and dense, consisting of golden recumbent bristles, that pubescence denser at base; presence of several big and deep punctures from where very long erect black setae originate. Legs slender, with femora enlarged in apical half, notably on forelegs; tibiae thin and straight.

Female. Similar to male, but differing as follows: antennae shorter, reaching apex of elytra when extended posteriorly; elytra slightly enlarged toward apex, with very long and sparse erect black setae; golden elytral pubescence less dense, nearly absent toward apex; femora less clavate and legs slightly shorter.

Discussion: After thorough investigations in the field, we were not able to find the type species elsewhere than the type locality, where, however, it was quite abundant in an area limited to only a few square meters. It would be interesting to investigate other areas in SW Saudi Arabia to better assess the range of this taxon.

Material examined: Holotype 〖: Saudi Arabia: Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 16.IV.2016, P. Rapuzzi lgt, ex larva from Dodonaea viscosa, emerged 12.II.2017 (KSMA); Allotype ♀: Saudi Arabia: Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 16.IV.2016, P. Rapuzzi lgt, ex larva from Rosa cf abyssinica R. Br., 1814, emerged 04.XII.2016 (CPR); Paratypes 3♀: Saudi Arabia: Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 16.IV.2016, P. Rapuzzi lgt (CPR); 4♂ and 3♀: idem, ex larva from Acacia sp., emerged from 23.VI to 24.XI.2016 (CPR); 32♂ and 20♀: idem, ex larva in Rosa cf abyssinica, emerged from 29.XII.2016 to 1.V.2017 (CPR); 67♂ and 83♀: idem, ex larva in Dodonaea viscosa, emerged from 08.V.2016 to 17.IV.2017 (CPR, KSMA); 3〖: Saudi Arabia: Abha prov., Raedah reserve, 2810 s.l.m., 18°12′14″N 42°24′43″E, 02.IV.2017, P. Rapuzzi lgt (CPR); 33♂ and 22♀: idem, ex larva in Dodonaea viscosa, emerged 20.IV.2017 (CPR); 13♂ and 9♀: idem, ex larva in Rosa cf. abyssinica, emerged 27.IV.2017 (CPR); 45♂ and 60♀: idem, emerged 1.V-15.VI.2019; 1♀ Saudi Arabia: Jebel Soudah, Raydah reserve, 18°11′41″N 42°24′18″E, 2255 s.l.m., 11.IV.2019, in pupal cell of Abutilon bidentatum A. Ric., 1847 [Malvaceae], P. Rapuzzi lgt (CPR).

Diagnosis: The new species is similar to the related Gracilia minuta. It is easy to distinguish according to the characters used to separate the new genus. The type series shows a variability in terms of size and color. In fact, the length is from 2 to 7 mm and the transversal band in the middle of the elytra is variable in terms of width as sometimes it is very thin and sometimes very wide; only rarely it is totally missing. The ground color is from light brown to dark brown.

Etymology: The name saudita is derived from Saudi Arabia.

Description: Length 5.5 mm., width 1.0 mm (holotype); length 2.0-6.5 mm (males), 3.0-7.0 mm (females). Dark pitch coloured except for a lighter transverse elytral stripe; apex of femora and antennae yellowish. Head small, with punctation consisting of dense and small punctures; pubescence generally dense, consisting of short, recumbent golden bristles, becoming denser near mandibles and around eyes; frons transverse, with small medial depression between antennal tubercles. Antennae slender, consisting of eleven segments covered with short, coricate, golden hairs; first three segments entirely yellowish; basal portion of segments four to six dark brown; presence of several long erect black apical setae on each antennomere, those setae denser on first three segments. Pronotum long, twice longer than large, constricted near base, with sides curved, lacking any prominence; with two small flat callosities at basal quarter of disc; punctation fine and dense, with short but distinctly less punctate medial line; surface uniformly covered with dense coricate golden bristles hiding background color. Scutellum glabrous, thin and long, rounded toward apex. Elytra parallel-sided, slighty constricted at middle; apex rounded; disc dark brown, divided transversely by yellowish stripe; punctation fine, denser near base and sparser toward apex; disc covered with golden coricate short bristles; with several much larger and deeper punctures from which originate long and stout black erect setae. Legs long, covered with dense golden pubescence; femora swollen subapically, especially fore femora; base of femora lighter in colour, yellowish; tibiae and tarsi slender.

Female: Similar to male, but body golden pubescence sparser; antennae shorter, reaching only elytral apex when extended posteriorly; elytra enlarged toward apex; elytral sculpture sparser and shallower; legs slender with femura less swollen.

Distribution: The new species is known only from a few places in the Raydah reserve (Abha province, Saudi Arabia).

Remarks: In both sexes the background colouration can be more or less light and the yellow elytral transverse stripe can be interrupted near the suture and reduced to a thin yellow line, or even totally missing (only in a few specimens).

Biology: Arabogracilia saudita sp. nov. seems quite polyphagous as it was reared from different plants belonging to different groups: Rosa cf abyssinica (R. Br. ex Lindl., 1820) (Rosaceae), Dodonaea viscosa (L.) (Sapindaceae) and Acacia sp. (Fabaceae). A few specimens were collected running on small dry branches of their host. The larvae start feeding under the bark of small to medium sized twigs of (from 0.5 cm to 3 cm in diameter), and just before pupation they enter the wood where they prepare a pupal cell, which is oval. The adults run very fast on the dry twigs and suddenly stop for a while with the antennae oriented forward, as do Gracilia minuta (Fabricius, 1781).

Hesperophanaires Mulsant, 1839: 61. Type genus: hesperophanes Mulsant, 1839.

Jebusaea Reiche, 1878: CLIII-CLIV. Type species: Jebusaea hammerschmidti Reiche, 1878 (by monotypy).

Jebusaea hammerschmidti Reiche, 1878: CLIV. Type locality: “Palaestine: Jaffa″ (Israel).

Jebusaea hammerschmidti: Holzschuh, 1993: 116; Hamadttu & Maged, 2016: 55-65.

Material examined from Saudi Arabia: Hofuf, 23.V., 6, 10 and 26.VI.1978, W. Büttiker; Al Khobar, 5.VI.1982, E. Heiss; Eastern Province, Udhaitiyah Camp, 6.VI.1982, D.A. Pitcher; Qatif, 14-15.IV.1983, very many larvae in bark (frond pads) of living date palms [Phoenix dactylifera L., 1753, Arecaceae], but only a few successfully reared to pupae, C. Holzschuh (Holzschuh, 1993); King Faisal University Research Station, Date Palm Research Center of Excellence experimentation blocks, 25°16′24.37″N 49°42′27.82″E, 151 s.l.m. X.2011-XII.2015 (Hamadttu & Maged, 2016); Ryiadh, Dareb; Hofuf, 25.VI.1982; Al Waseel, 8.XII.2012; Wasiel, 45 km W of Riyadh, Tadamonia Co farm, 31.V.2008, at light; Al Jouf, Sakaka, Hamadan Al Faleh, 26.V.2018, 29°58.07′N 40°08.08′E, 597 m (KSMA); Al Khobar, 5.VI.1982, E. Heiss (CPR; CGS); Hofuf, 23.V., 6, 10 and 26.VI.1978, W. Büttiker (CGS).

Distribution: Iran, Irak, Israel. Arabian Peninsula: Saudi Arabia (Holzschuh, 1993), United Arab Emirates, Qatar, Oman (Danilevsky, 2020).

Zoodes Pascoe, 1867: 319. Type species: Stromatium maculatum White, 1855 (by original designation).

hesperophanes liturifer Walker, 1871: 18. Type locality: “Harkeko″ (Sudan, Red Sea).

Zoodes liturifer: Shalaby, 1961: 221; Holzschuh, 1993: 116.

Material examined from Saudi Arabia: Djedda, 19.IV.1956 (Shalaby, 1961); Jizan, at light, 25-26. III.1983, C. Holzschuh; Fayfa, at light, 27-31.III.1983, C. Holzschuh; Asir, Sabiya, 17°10′N 42°30′E, 1.XII.1946 (Holzschuh, 1993); “Saudi Arabia″; Al Baha, Shada Al Ala, 19°51.762′N 41°18.089′E, 1225 s.l.m., 24.VIII.2014, at light (KSMA).

Distribution: Egypt, Sudan, Ethiopia, Djibouti, Somalia, Kenya, Tanzania, Malawi, Mozambique, Zimbabwe, Namibia, Republic of South Africa. Arabian Peninsula: Saudi Arabia, Yemen (Adlbauer & Beck, 2015).

Hesperophanini (Daramina) Sama, 2008: 224. Type genus: Daramus Fairmaire, 1892.

Daramus Fairmaire, 1892: 121. Type species: Daramus serricornis Fairmaire (by monotypy).

Daramus serricornis Fairmaire, 1892, 11: 121. Type locality: “Obock″ (Djibouti).

Daramus serricornis: Lepesme, 1948: 256.

Material examined from Saudi Arabia: Najran (Lepesme, 1948); Al Baha prov., Wadi Thee Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, 1 dead specimen in pupal cell on Acacia sp., P. Rapuzzi lgt (CPR); Asir prov., 14 km N Khamis Mushait, Wadi Ibn Hashbal, 1926 s.l.m., 18°27′34″N 42°42′53″E, 2.IV.2017, ex larva in Acacia sp., emerged 31.XII.2018, P. Rapuzzi lgt (CPR); Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, P. Rapuzzi lgt, ex larva in Acacia sp., emerged 05.I.2019 (CPR).

Distribution: Egypt, Senegal, Niger, Chad, Sudan, Ethiopia, Djibouti, Somalia, Kenya, Tanzania. Arabian Peninsula: Saudi Arabia, Yemen, Oman (Adlbauer & Beck, 2015; Danilevsky, 2020).

Lygrini Sama, 2008: 222. Type genus: Lygrus Fåhroeus, 1872.

Note: The tribe name should be Lygrini and not Pelossini as stated by Tavakilian (2013) (Sama, 2023: 114-115).

Pelossus Thomson, 1864: 222. Type species: Corethrogaster ruber Thomson, 1858 (by original designation).

Digonium longicorne Pic, 1895: 77. Type locality: “Arabie″.

Lygrus longicorne: Holzschuh, 1993: 115-116.

Pelossus longicornis: Yokoi et al., 2016: 230.

Material examined from Saudi Arabia: Wadi Jizl, 26°42′N 37°15′E, 1000 s.l.m., 13-14.XI.1984, W. Büttiker; Harithi, 21°18′N 40°18′E, 1910 s.l.m., 10-11.V.1984, W. Büttiker; Makkah 21°06′N 40°24′E, 1730 s.l.m., 22-24.VIII.1985, W. Büttiker; Asir Mts., Wadi Qust (SE of Taif), 1400 s.l.m., 6.IV.1983, emerged ex Acacia sp., 4.IX.1983, C. Holzschuh; An Naamah, 20°15′N 41°16′E, 2100 s.l.m., 31.VIII.1984, W. Büttiker; Asir Mts., Shaqiq Shamran (between Taif and Abha), 8.IV.1983, emerged ex unknown host, 1.VI.1985, C. Holzschuh; Jebel Beles (Baha), 19°48′N 41°50′E, 2070 s.l.m., 19.X.1984, W. Büttiker; Asir Mts., An Namas (between Taif and Abha), 2400 s.l.m., 8.IV.1983, emerged ex Acacia sp. between 17.IV and 1.VI.1983, C. Holzschuh; Namas, 2380 s.l.m., 15.IV.1980, W. Büttiker; idem, G. Vogl; idem, 17.IV.1980, W. Büttiker; idem, 18-19.IX.1980, W. Büttiker; Asir Nat. Park, Abha, 7.VI.1982; idem, at light, 9.IV.1983, C. Holzschuh; Asir Mts, 85 km W of Najran, 10.IV.1983, emerged ex Acacia 4.IX.1983, C. Holzschuh; Asir Mts. Fayfa, at light, 27-31.III.1983, C. Holzschuh; idem, emerged ex Acacia sp. between 12.IV and 11.IX.1983, C. Holzschuh; idem, emerged ex Ficus salicifolia, C. Holzschuh; Jizan, 25-26. III.1983, emerged ex Acacia 7.VIII. and 11.IX.1983, C. Holzschuh; Bani Rizam, 12.IV.1980, W. Büttiker; Ash Sharayi, 42 km before Zahran, 24.IX.1978, W. Büttiker; Al Dalhan, near Ash Sharayi, 19-20.IX.1980, W. Büttiker; Riyadh, 23.V.1980, W. Büttiker; 28 km SSE of Athnen, 2390 s.l.m., 5-6.X.1980, Büttiker; An Nimas, 2450 s.l.m., 4.IV.1980, K.M. Guichard (Holzschuh, 1993); Asir Mts., An Namas (Yokoi et al., 2016); Taif, Al Wesam distr., 21°12′17″N 40°20′43″E; Taif, 9.IV.2003, ex larva in Acacia sp.; An Namar, 8.IV.1983; Asir, Al Magardah, W. Yabah, 402 s.l.m., 19°14.911′N 41°47.255′E 11.X.2013, at light; Baha, Station Shabreqah, 2267 s.l.m., 20°08.034′N 41°23.598′E; Mandaq, Wadhi Tourbah, 19°55.774′N 41°26.574′E, III.2012, at light; Fifa, 1.IV.1983; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 3.VI.2014, at light; idem, 15.II.2014; idem, 23.VIII.2014; idem, 12.XI.2015; idem, 14.XI.2015; Al Baha, Shada Al Ala, 19°50.575′N 41°18.891′E, 1666 s.l.m., 15.II.2014, at light; Al Baha, Shada Al Ala, Housing, 19°52.598′N 41°18.672′E, 892 s.l.m., 5.VI.2014, at light; Al Baha, Shada Al Ala, Housing, 18°13.347′N 42°24.133′E, 892 s.l.m., 5.VI.2014, at light; Abha, Reydah, 18°11.766′N 42°24.315′E, 2285 s.l.m., 18.XI.2015, at light; idem, 26.IV.2014; idem, 6.VI.2014; idem, 31.VI.2015; Abha, Raydah, 18°12.095′N 42°24.536′E, 2578 s.l.m., 27.IV.2014, at light; idem, 20.X.2014; idem, 26.VIII.2014; idem, 5.IX.2015; Abha, Raydah, 18°11.695′N 42°23.818′E, 1897 s.l.m., 6.VI.2014, at light; idem, 26.VIII.2014; Abha, Raydah, 18°11.884′N 42°24.435′E, 2387 s.l.m., 26.II.2014, at light; idem, 3.II.2014; idem, II.2014; idem, 7.VI.2014; idem, 20.X.2014; idem, 4.III.2015; idem, 8.V.2015; Abha, Reydah housing, 19°50.411′N 41°18.686′E, 2717 s.l.m., 8.V.2014, at light; Abha, Reydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 26.VIII.2014, at light; idem, 7.VI.2014; Abha, Reydah, 18°12.315′N 42°24.607′E, 2761 s.l.m., 26.VIII.2015, at light; Abha, Raydah, 18°12.265′N 42°24.744′E, 2820 s.l.m., 4.III.2015, at light; Abha, Raydah, 18°11.679′N 42°23.691′E, 1851 s.l.m., 6.VI.2014, at light (KSMA); Al Baha prov., Al Qissaimah, 2156 s.l.m., 20°18′30″N 41°20′02″E, 12.IV.2016, ex larva in Acacia sp., emerged 11-30.V.2016, P. Rapuzzi lgt; idem, emerged 1.VII.2016; idem, emerged 1-28. VIII.2016; 2-25.IX.2016; idem, emerged 2.XI.2016; idem, emerged 4.XII.2016; Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 15.IV-18.V.2016, ex larva in Acacia sp., emerged 17-28.V.2016, P. Rapuzzi lgt; idem, 2-23.VI.2016; idem, emerged 29.VII.2016; idem, emerged 12-22.VIII.2016; idem, emerged 11-25.IX.2016; idem, emerged 12-21.X.2016; idem, emerged 8.XI.2016; idem, emerged 4.XII.2016; idem, emerged, 18.II.2017; Asir prov., Jebel Soudah, Raydah res., 18°11′41″N 42°24′16″E, 2255 s.l.m., ex pupa Ficus sp., 11.IV.2019, P. Rapuzzi lgt; idem, ex larva from Acacia sp., emerged 21.III.2020, P. Rapuzzi lgt; Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Acacia sp., emerged 28.V.2016, P. Rapuzzi lgt; idem, emerged 20.IX.2016; idem, emerged 12.X.2016; Al Bahah prov., Jabal Shada al A′la, 1246 s.l.m., 19°51′42″N 41°18′15″E, 29.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1695 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, ex larva in Ficus palmata Forssk., 1775 [Moraceae], emerged 27.VI.2018, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Bahah prov., Shada al Ala, 1692 s.l.m., 19°50′37″N 41°18′40″E, 8.IV.2019, ex larva from Ficus sp., P. Rapuzzi leg.; Al Baha prov., Bani Hassan, 2258 s.l.m., 20°03′00′N 41°22′14″E, 12.IV.2016, ex larva in Acacia sp., emerged 14-16.V.2016, P. Rapuzzi lgt; idem, emerged 2-21.VI.2016; idem, emerged 13.VII.2016; idem, emerged 7.VIII.2016; idem, emerged 25.IX.2016; idem, emerged 12.X.2016; idem, emerged 14.XI.2016; Bahah prov., Bani Hassan, 2293 s.l.m., 20°03′02″N 41°26′47″E, 30.III.2017, ex larva in Acacia sp., emerged 3.V.2017, P. Rapuzzi lgt; idem, emerged 28.VII.2017; Al Baha prov., Raghadan forest, 2236 s.l.m., 20°01′07″N 41°26′53″E, 13.IV.2016, ex larva in Acacia sp., emerged 4-8.V.2016, P. Rapuzzi lgt; idem, emerged 19-29.VII.2016; idem, emerged 18.VIII-3.IX.2016; idem, emerged 12.X.2016; Al Baha prov., Wadi Tourabah, 1830 s.l.m., 20°11′36″N 41°17′50″E, ex larva in Acacia sp., emerged 2.IX.2016, P. Rapuzzi lgt; Asir prov., 14 km N Khamis Mushait, Wadi Ibn Hashbal, 1926 s.l.m., 18°27′34″N 42°42′53″E, 2.IV.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Bani Hassan, 2293 s.l.m., 20°03′02″N 41°26′47″E, 30.III.2017, in pupal cell on Acacia sp., P. Rapuzzi lgt; idem, ex larva in Acacia sp., emerged 3.V.2017; Bahah prov., Wadi Tourabah, E of An Na′amah, 20°11′01″N 41°18′42″E, 30.III.2017, ex larva in Pistacia sp., emerged 6.IV.2017, P. Rapuzzi lgt; Asir prov., Jebel Soudah, 1632 s.l.m., 18°11′45″N 42°23′21″E, 12.IV.2017, ex larva in Pistacia sp., emerged 6.IV.2017, P. Rapuzzi lgt; idem, emerged 1.IX.2017; Al Bahah prov., Jabal Shada al A′la, 1965 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, ex larva in Ficus palmata, emerged, 10.VII.2017, P. Rapuzzi lgt; idem, emerged 10.X.2017; idem, emerged 11-28.XI.2017; idem, emerged 15.XII.2017; Ryadh prov., Wadi Haysaha, 790 s.l.m., 24°55′22″N 46°12′15″E, 8.IV.2017, ex larva in Acacia sp., emerged 3.IX.2017, P. Rapuzzi lgt; Al Bahah prov., Wadi Tourabah, 20°11′16″N 41°18′38″E, 1812 s.l.m., 6.IV.2019, ex larva in Acacia sp., emerged 25.V-25.VI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen, Oman (Danilevsky, 2012).

Nathriini Linsley, 1963: 152. Type genus: Nathrius Brèthes, 1916.

Mourgliana Holzschuh, 1993: 125. Type species: Mourgliana conspicua Holzschuh, 1993 (by original designation and monotypy).

Mourgliana conspicua Holzschuh, 1993: 125-126. Type locality: “Saudi Arabia: Fayfa″.

Mourgliana conspicua: El-Hawagry et al., 2013: 28.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (holotype); Wadi Dhiyan, 19°50′N 41°28′E, 1050 s.l.m., 13.IX.1983, W. Büttiker; Wadi Dhiyan, 19°48′N 41°36′E, 830 s.l.m., W. Büttiker (Holzschuh, 1993); Al-Baha, Dhee Ain, May (El-Hawagry et al., 2013); Al Shefie road, Whadi Thee Ghazal, 21°05′28″N 40°21′04″E, 31.V.2011; Abha-Khamus road, Madina senaeva, 18°14′19″N 42°34′14.5″E, 2147 s.l.m., 28.IV.2011; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 23.VIII.2014, at light; Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 26.IV.2014, at light; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 12.IX.2014, at light; idem, 2.III.2015; idem, 12.XI.2015 (KSMA); Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Acacia sp., emerged 20.X.2016, P. Rapuzzi lgt; Al Baha prov., Wadi Tourabah, 1830 s.l.m., 20°11′36″N 41°17′50″E, 12.IV.2016, ex larva in Acacia sp., emerged 11.V.2016, P. Rapuzzi lgt; idem, emerged 15.V.2017; Asir prov., 14 km N Khamis Mushait, Wadi Ibn Hashbal, 1926 s.l.m., 18°27′34″N 42°42′53″E, 2.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4. IV.2017, at light, P. Rapuzzi lgt; Asir prov.: Wadi Ksan, 3 km N of Hubeil, 18°08′57″N 43°13′56″E, 462 s.l.m., at light, 12.IV.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (type locality).

Obriaires Mulsant, 1839: 95. Type genus: Obrium Dejean, 1821.

Iranobrium Villiers, 1967: 338. Type species: Iranobrium davatchii Villiers, 1967 (by monotypy and original designation).

Iranobrium brancuccii Holzschuh, 1993: 122-123. Type locality: “Saudi Arabia: Fayfa″.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (holotype) (Holzschuh, 1993).

Distribution: Arabian Peninsula: Saudi Arabia (type locality), Oman, Yemen (Danilevsky, 2020).

Iranobrium buettikeri Holzschuh, 1993: 123-124. Type locality: “Saudi Arabia: Shawaq (on the plain SE of Jeddah)″.

Material examined from Saudi Arabia: Shawaq (on the plain SE of Jeddah) at light, 5.IV.1983, C. Holzschuh (holotype); Wadi Qarma, 19°32′N 40°58′E, 14.II.1937, H.St.J.B. Philby (Holzschuh, 1993); Al Baha, Shada Al Ala, 19°51.762′N 41°18.089′E, 1225 s.l.m., 3.VI.2014, at light; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 3.VI.2014, at light (KSMA); Asir prov., 6 km S of Ainah, 200 s.l.m., 17°55′28″N 42°15′20″E, 3.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (type locality), United Arab Emirates (Danilevsky, 2012), Yemen (Danilevsky, 2020).

Ossibia Pascoe, 1867: 311. Type species: Obrium fuscatus Dejean, 1837 (nomen nudum = Obrium fuscatum Chevrolat, 1856) (by original designation).

Ossibia picata Holzschuh, 1993: 124-125. Type locality: “Saudi Arabia: Asir Mts.: Wadi Shuqub (SE Taif)″.

Material examined from Saudi Arabia: Asir Mts., Wadi Shuqub (SE Taif), 7.IV.1983, emerged from unidentified dry twigs 11.IX.1983, C. Holzschuh (holotype) (Holzschuh, 1993); Abha, Raydah, 18°11.766′N 42°24.315′E, 2285 s.l.m., 26.IV.2014, at light; idem, 31.VII.2014; Abha, Raydah, 18°11.695′N 42°23.818′E, 2285 s.l.m., 6.VI.2014, at light; Abha, Raydah, 18°12.095′N 42°24.536′E, 2578 s.l.m., 5.IX.2015, at light; Abha, Raydah, 18°11.766′N 42°24.315′E, 1897 s.l.m., 31.VII.2015, at light; Abha, Raydah, 18°11.679′N 42°23.691′E, 1851 s.l.m., 31.VII.2015, at light; Abha, Raydah, 18°11.618′N 42°24.420′E, 1772 s.l.m., 31.VII.2015, at light; Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 27.IV.2014, at light (KSMA); Al Bahah prov., Wadi Tourabah, 20°11′16″N 41°18′38″E, 1812 s.l.m., 6.IV.2019, ex larva in Pistacia sp., emerged 22.VI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (type locality).

yemenobrium Adlbauer, 2005: 1008. Type species: yemenobrium velutinum Adlbauer, 2005 (by original designation).

yemenobrium velutinum Adlbauer, 2005: 1008. Type locality: “Yemen, Sana′a – Haddah (15°17′24″N 44°09′59″E)″.

Material examined from Saudi Arabia: Asir prov., Jebel Soudah, Raydah reserve, 18°11′41″N 42°24′18″E, 2255 s.l.m., 11.IV.2019, in pupal cell ex Abutilon bidentatum, G. Magnani lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4. IV.2017, ex larva from Ficus sp., emerged 20.IV.2021, G. Magnani lgt (CPR).

Distribution: Arabian Peninsula: Yemen (type locality); Saudi Arabia (new record).

Phoracanthidae Newman, 1840: 2. Type genus: Phoracantha, Newman, 1840.

Phoracantha Newman, 1840: 19. Type species: Phoracantha synonyma Newman, 1840 (designated by Thomson, 1864).

Stenocorus semipunctatus Fabricius, 1775: 180. Type locality: “Nova Hollandia″ (Australia).

Material examined from Saudi Arabia: Al Baha (KSMA).

Distribution: Described from Australia. Introduced all over the World: New Zealand, Republic of South Africa, Algeria, Egypt, Mauritius, Malawi, Morocco, Mozambique, Tunisia, Zambia, Argentina, Uruguay, Brazil, Peru, Bolivia, Chile, USA, Mexico, Israel, Italy, Spain, Portugal, France, Turkey (Martins & Galileo, 2006), Cyprus (Martines M., 1983), Papua New Guinea (Wang, 1995), Zimbabwe, Lesotho (Adlbauer, 2001), Malta (Mifsud, 2002), Lybia (Sama et al., 2005), Greece (Berger, 2005), Cuba (Grillo & Fernandez, 2007), Argentina (Di Iorio, 2007), Syria (Ali & Rapuzzi, 2016), Lebanon (Coquempot et al., 2016), Reunion Island (Parnaudeau, 2016). Arabian Peninsula: Saudi Arabia (new record).

Xystroceritae Blanchard, 1845: 147. Type genus: Xystrocera Audinet-Serville, 1834.

Xystrocera Audinet-Serville, 1834: 69. Type species: Cerambyx globosus Olivier (designated by Thomson, 1864).

Xystrocera dispar Fähroeus, 1872: 49. Type locality: “Caffrariae″ (Republic of South Africa).

Material examined from Saudi Arabia: Fayfa, 3.X.1984, collector not given; Jizan, 21.XI.1981, W. Büttiker (Holzschuh & Téocchi, 1991); Jazan, Fifa 26.III.2011, at light.; Al Majardah, Talooth, Al Mandhar, Wadi Baqrah, 18°55.642′N 42°02.829′E 587 s.l.m., 31.V.2012, sweeping; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 14.XI.2015, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 3.XI.2015, at light; idem, 13.XI.2015; idem, 14.XI.2015; idem, 15.XI.2015; Abha, Raydah, 18°11.766′N 42°24.315′E, 1897 s.l.m., 20.X.2014, at light (KSMA).

Distribution: Burkina Faso, Chad, Sudan, Ethiopia, Djibouti, Somalia, Democratic Republic of the Congo, Republic of the Congo, Uganda, Kenya, Tanzania, Mozambique, Zimbabwe, Botswana, Namibia, Republic of South Africa. Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015), Yemen (Jabal Bura, NEE Al Hudaydah, 14°52′N 43°24′E, 261-600 s.l.m., 11.IV.2007, S. Kadlec lgt) (NHMP) (new record); United Arab Emirates (Fujairah, 20-27.VI.2006, CGS) (new record).

Lamiaires Latreille, 1825: 401.

Agapanthaires Mulsant, 1839: 165, 172. Type genus: Agapanthia Audinet-Serville, 1835.

hyllisia Pascoe, 1864: 285. Type species: hyllisia stenideoides Pascoe, 1864 (by monotypy).

Material examined: Holotype 〖: Saudi Arabia: Al Baha, Shada Al Ala, 19°50.575′N 41°18.691′E, 1666 s.l.m., 2.IX.2015, at light (KSMA); Paratypes. 1〖: Saudi Arabia: Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 23.VIII.2014, at light (KSMA); 1〖: Saudi Arabia: Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 28.IV.2014, at light (KSMA); 1♀: Saudi Arabia: Abha, Raydah, 18°11.749′N 42°23.345′E, 1614 s.l.m., 26.IV.2014, at light (KSMA); 1〖: Saudi Arabia: Al Baha, Shada Al Ala, 19°50.329′N 41°18.804′E, 1543 s.l.m., 23.VIII.2014, at light (KSMA); 1♀: Saudi Arabia: Abha prov., Raedah reserve, 26.IV.2014, at light (KSMA); 1♀: Saudi Arabia: Asir prov., Wadi Marabah, 18°10′18″N 42°22′12″E, 1197 s.l.m., 11.IV.2019, one dead specimen in pupal cell ex Euphorbia schimperi C. Presl, 1845 [Euphorbiaceae], P. Rapuzzi lgt; 3〖: Saudi Arabia: Asir prov., Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°02′22″E, 16.IV.2016, at light, P. Rapuzzi lgt; 1〖: Saudi Arabia: Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, ex larva from Euphorbia schimperi, emerged 15.V.2017, P. Rapuzzi lgt (CPR).

Diagnosis: hyllisia asirica sp. nov. most resembles h. niveovittata Aurivillius, 1910 from East Africa, from which it differs in having a rather homogeneous body pubescence instead of a pubescence that is irregular and scattered or forming more or less small dense rounded white spots. In addition, h. asirica sp. nov. has the elytral apex truncate instead of rounded as in h. niveovittata, and the apex of its tegmen have only very few setae, one long and a few very short instead of a tuft of setae of equal length in h. niveovittata.

Etymology: The name asirica derives from the Asir province (Saudi Arabia).

Description: Length 10.0 mm; width 2.3 mm (holotype); paratype males 9.0-11.9 mm, females 9.9-11.9 mm. Body brownish-black, dorsum covered with ochraceous pubescence. Head oriented downward, frons two times longer than wide, covered with ochraceous pubescence denser near inner ocular margin and on genae, and presence of few erect black setae near eyes; deeply punctured tegumen hidden by pubescence; temples straight; frons slightly convex; presence of deep medial furrow between antennal tubercles; occiput with distinct carina in middle. Antennae very long, 2 times longer that body when extended posteriorly, brownish-black and more or less densely covered with ochraceous pubescence; the latter pubescence denser on lower side and sparser on upper side of first antennomere; presence of several short, black, erect setae on inner side of antennomeres 2 to 4 only. Pronotum longer than wide, cylindrical, shallowly depressed subbasally, with sides fairly straight except middle section slightly arcuate; whitish pubescence forming three lines, one extended medially on entire length and the others along lateral margins (not visible from above); presence of few isolated black erect laterobasal setae; punctation deep, hidden by pubescence. Scutellum two times longer than wide, rounded toward apex; disc slighty swollen at middle, nearly glabrous except for a few ochraceous setae in middle. Elytra parallel-sided, slightly constricted subapically; disc convex, with suture slightly swollen; apex truncate; disc uniformly covered with dense and deep irregular punctures; pubescence dense, consisting of recumbent short ochraceous bristles, lacking erect setae. Legs short; tarsi long, expecially apical tarsomere. Apex of tegmina with only very few setae, one long and three or four very short on each tegmen.

Female: Similar to male except body stouter and antennae shorter, 1.5X body length.

Distribution: The new species is endemic to Southwest Saudi Arabia.

Ancylonotides Lacordaire: 299, 301. Type genus: Ancylonotus Laporte, 1840.

Idactus Pascoe, 1864: 273. Type species: Idactus tridens Pascoe, 1864 (by monotypy).

Idactus Bettoni Gahan, 1898: 46-47. Type locality: “Samburu and Voi, British East Africa″ (Kenya).

Idactus bettoni: Téocchi, 1988: 17; Holzschuh & Téocchi, 1991: 303.

Saudi Arabia: Fayfa, 23.X.1983 (Téocchi, 1988); Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Al Atif-Maccah road, 21°50′51″N 40°59′27″E, 8.XI.2011; Al Baha, Shada Al Ala, 19°52.717′N 41°18.712′E, 825 s.l.m., 2-3. III.2015, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 2-3.III.2015, at light (KSMA).

Distribution: Ethiopia, Somalia, Kenya, Tanzania, Malawi, Mozambique, Zimbabwe, Botswana, Namibia. Arabian Peninsula: Saudi Arabia (Téocchi, 1988).

Dichosthates coquereli Fairmaire, 1890: 551. Type locality: “Massouah″ (Massaoua, Erythrea).

Idactus coquereli: Adlbauer et al., 2008: 1174; Adlbauer & Beck, 2015: 114.

Material examined from Saudi Arabia: None (see Note below).

Distribution: Ethiopia, Djibouti, Somalia, Kenya. Arabian Peninsula: Yemen, Oman (Adlbauer & Beck, 2015).

Note: This record is due to a mistake. Adlbauer et al. (2008) and later Adlbauer & Beck (2015) reported Saudi Arabia as part of the range of this species based on Holzschuh & Téocchi (1991), but in this paper no record of I. coquereli is reported for Saudi Arabia. However, the presence of this species in the Kingdom is very probable.

Pogonocherus cristulatus Fairmaire, 1886: 458. Type locality: “Obok″ (Djibouti).

Idactus assimilis Breuning, 1938: 214-215. Type locality: “Arabien: Ktubu″ (Yemen).

Idactus cristulatus: Villiers, 1968: 849; Holzschuh & Téocchi, 1991: 303.

Material examined from Saudi Arabia: Jeddah, Buraiman, 16.III.1946, and Lith, I.1945; Dhufor, Jebel Qara, Wadi Geth, XI.1943; Lith, “10 m inland″, I.1945 (Villiers, 1968); Fayfa, at light, 27-31.III.1980, C. Holzschuh; Fayfa, reared from Calotropis procera (Aiton) W. T. Aiton, 1810 [Apocynaceae], emerged 12.IV.1983, C. Holzschuh; Fayfa, 3.X.1984, A. S. Talhouk; Gizan, at light, 25-26.III.1983, C. Holzschuh; Gizan, reared from branches of Cammiphora opobalsamum, emerged 17.IV.1983; NW Gizan, Shuqayq, at light, 2.IV.1983, C. Holzschuh (Holzshuh & Téocchi, 1991); El Kuber, (Yemen frontier); Shuqaiq, 7.2.27, leg. Philby; Anjaba, 6.I.37, Philby; Harabrab, 3.I.27, Philby; Idda, 31.XII.34; Lith, I.1945 (National Museum, London, United Kingdom); Fifa, 3.X.1984; Jizan, Al Aydabi, Haqu, Fayfa, 17°20.541′N 43°02.620′E, 237 s.l.m., 21.V.2012; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 24-25. IV.2014, at light; idem, 20.IV.2014; idem, 21.IV.2014; idem, 3.VI.2014; idem, 16.XI.2015; idem, 12.XI.2015; idem, 14.XI.2015; Al Baha, Shada Al Ala, 19°52.717′N 41°18.712′E, 825 s.l.m., 15.XI.2014, at light (KSMA); Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 28 km SSE Muhayl, 809 s.l.m., 18°17′55″N 42°07′41″E, 5.IV.2017, in pupal cell on Calotropis procera, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 13.IV.2019, ex larva in Calotropis procera, emerged 19.V.2019, P. Rapuzzi lgt (CPR).

Distribution: Mali, Ethiopia, Somalia, Ivory Coast, Cameroon, Central African Republic, Tanzania, Mozambique, Zimbabwe, Republic of South Africa. Arabian Peninsula: Saudi Arabia, Yemen, Oman (Adlbauer & Beck, 2015).

Pseudoparaphloeus Sama, 2009: 25. New name for Paraphloeus Breuning, 1968: 61. Type species: Paraphloeus scorteccii Breuning, 1968 (by original designation and monotypy).

Paraphloeus scorteccii Breuning, 1968: 92. Type locality: “Arabie: Taiz, 1350 s.l.m.″ (Yemen).

Paraphloeus scorteccii: Holzschuh & Téocchi, 1991: 303.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light (KSMA).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen (Adlbauer & Beck, 2015). A subspecies (erythreensis Téocchi, 1992) occurs in East Africa (Erythrea).

Apomecynitae Thomson, 1860: 66. Type genus: Apomecyna Dejean, 1821.

Apomecyna Dejan, 1821: 108. Type species: Saperda alboguttata Megerle, 1802 (= Lamia histrio Fabricius, 1793) (by monotypy).

Apomecyna binubila Pascoe, 1858: 253. Type locality: “Natal″ (Republic of South Africa).

Material examined from Saudi Arabia: Asir prov., Wadi Ksan, 3 km N of Hubeil, 18°08′57″N 43°13′56″E, 462 s.l.m., ex pupa from Citrullus colocynthis (L.) Schrad., 1838 [Cucurbitaceae], 12.IV.2019, P. Rapuzzi lgt (CPR).

Distribution: Senegal, Niger, Sudan, Ethiopia, Somalia, Guinea-Bissau, Guinea, Sierra Leone, Ivory Coast, Ghana, Togo, Benin, Nigeria, Congo, Rwanda, Tanzania, Angola, Zambia, Malawi, Zimbabwe, Mozambique, Namibia, Botswana, Republic of South Africa (Adlbauer & Beck, 2015). Arabian Peninsula: Saudi Arabia (new record).

Eurycotyle Lameerei Pic, 1895: 77. Type locality: “Arabia″.

Apomecyna arabica Breuning, 1938: 50. Type locality: “Arabie: La Mecque″.

Apomecyna lamereei: Villiers, 1968: 849; Holzschuh, 1979: 294; Amin et al., 1986: 312; Holzschuh & Téocchi, 1991: 301.

Material examined from Saudi Arabia: Mecca district, Bir Sanoeia, 12.VIII.1944, A.R. Waterston; Buraiman, près Jeddah, 18.XI.1944, à la lumière, A.R. Waterston; Musaina, 23.V.1938, H.St.J.B. Philby (Villiers, 1968); Umgeb. Ryadh, Araida, Selouly′s Farm, 9.X.1975, leg. W. Büttiker; Umgeb. Riyadh, Wadi Hanifa, 7.V.1976, W. Büttiker; Wadi Hanifa, 600 s.l.m., 25.IV.1976, W. Wittmer & W. Büttiker; Umgeb. Riyadh, Wadi Mizbil, 18.IV.1977, W. Büttiker; Umgeb. Riyadh, Wadi Shaib Luha, 30.IX.1976, W. Büttiker (Holzschuh, 1979); Jeddah, trap light (Amin et al., 1986); Wadi Majarish, 1020 s.l.m., 7.II.1980, W. Büttiker; 80 km N Ryadh, Thumama park, 570 s.l.m., 24.III.1985, H. Kessler; Bahara, 9.XII.1980, W. Büttiker; Wadi Huraymala, 25°09′N 46°08′E, 770 s.l.m., ex Citrullus colocynthis, emerged 11-25.VIII.1988, C.W. Mills; 30 km W of Medina, 500 m, 27.IV.1979, Exp. N. Hedjaz (Holzschuh & Téocchi, 1991); Rhodet Khorim, 100 km E of Riyadh, 25°22′58″N 47°16′44″E, 1.X.2011, at light; idem, 16.X.2011; Al Aflag, Al Naifiyah, Farshet Sheaal, 22.42496N 46.57556E, 606 s.l.m., 12.IV.2015, at light; idem, 19.V.2015; Riyadh, Az Zulfi, Rawdat Al Sabalah, 26°22.040′N 44°59.137′E, 670 s.l.m.,25.X.2015, at light; Riyadh, Jobilah, Salam Farm, 7.VII.2007, at light; Riyadh, Jobilah, 28.IV.2007, at light; Riyadh, Al Obaiteh, Obikan Farm, 6.VIII.2007, at light; idem, 7.IV.2008; idem, 30.IV.2007; idem, 21.V.2007; idem, 23.VI.2007; idem, 2.VII.2007; idem, 6.VIII.2007; idem, 20.VIII.2007; idem, 27.VI.1985; idem, 27.VI.1988; Derabh, 17.XI.1989; Dimiyah, VIII.1988, at light; Dairab, 4.XI.1985, at light; Riyadh, Derab; Dairab, 2.XII.1985, at light; Riyadh, IV.1982; Riyadh, Wasiel, Tadamonia Co. Farm, 12.VII.2008, at light; Aljabilah, Bander Prince Farm, 12.IV.2008, at light; Riyadh, Rodet Khoreem, 25°22′58″N 47°16′44″E, 1.X.2011, at light; idem, 30.X.2011; idem, 16.X.2011; idem, 13.XI.2011; idem, 26.IV.2012; idem, 28.IV.2012; idem, 15.V.2012; Tumayr-Sudayr, 25°42′36″N 45°52′11″E, 5.V.2010; Riyadh, Al Amariyah, 24°81′84″N 46°44′70″E, 696 s.l.m., 15.XII.2013; Muzahimiyah, Al Khararah, 24°24′21″N 46°14′40″E, 17.IV.2012, at light; idem, 30.III.2011; idem, 26.IV.2011; idem, 13.IV.2011; idem, 30.X.2013; Al Amariyah, Animal Production Dept. Farm KSU, 20.IV.2011, at light; idem, 28.IV.2011; Muzahimiyah, Al Khararah, 24°24′21″N 46°14′40″E, 8.III.2014, at light (KSMA); Riyadh umg. (Coll. National Museum Prague, Cezch Republic); Riyadh prov., 36 km NW Ryadh, Al Uyaynah, 761 s.l.m., 24°53′33″N 46°17′40″E, 10.IV.2016, ex larva in Citrullus colocynthis, emerged 1.V.2016, P. Rapuzzi lgt; Riyadh prov., 30 km NW Ryadh, Al Uyaynah, 761 s.l.m., 24°53′33″N 46°17′40″E, 8.IV.2017, P. Rapuzzi lgt; idem, ex larva in Citrullus colocynthis, emerged 3.VI.2017; Ryadh prov., Rawdat Khuraim, 559 s.l.m., 25°25′57″N 47°13′52″E, 9.IV.2017, P. Rapuzzi lgt; Riyadh prov., 30 km NW Ryadh, Al Uyaynah, 756 s.l.m., 24°53′36″N 46°17′31″E, 2.IV.2019, many specimens under twigs of Citrullus colocynthis, P. Rapuzzi lgt; idem, many specimens ex larva from Citrullus colocynthis, emerged 01.V-20.V.2019, P. Rapuzzi lgt; Ryadh prov., W of Al Hulwah, Ibex reserve, 23°26′56″N 46°39′59″E, 635 s.l.m., 3.IV.2019, under twigs of Citrullus colocynthis, P. Rapuzzi lgt; Ryadh prov., Wadi Nesah, 24°13′35″N 46°51′14″ E, 570 s.l.m., 4.IV.2019, under twigs of Citrullus colocynthis, P. Rapuzzi lgt; Al Qassim prov., NE Abu Marwah, 22°09′33″N 42°12′02″E, 1001 s.l.m., 5.IV.2019, P. Rapuzzi lgt (CPR).

Distribution: Mauritania (Rungs, 1952), Western Sahara (Breuning, 1953), Niger, Chad, Senegal, Djibouti (Lepesme & Breuning, 1955), Mali (Villiers, 1962), Sierra Leone (Mourglia & Téocchi, 1994), Egypt, Morocco, Iran, Iraq, Israel, Pakistan (Löbl & Smetana, 2010). Arabian Peninsula: Saudi Arabia, Yemen (Löbl & Smetana, 2010), Oman (Ambrus & Grosser, 2012).

Note: In Al Uyaynah (756 m, 24°53′36″N 46°17′31″E) on 2.IV.2019, we collected a pair of specimens under twigs of Cucumis prophetarum L., 1758 (Cucurbitaceae). These specimens are different from A. lameerei in the pubescence which is almost totally missing (it is only represented by two whitish incomplete transverse bands on the elytra), and the elytral punctures are sparser and larger. It is possible that these specimens belong to a new species, but it would be necessary to study a larger number of specimens before a decision can be reached on the real status of this population. Moreover, all of the specimens of A. lameerei that we have studied (several hundreds of specimens from Mauritania to Iran) strictly live on Citrullus colocynthis (Cucurbitaceae) and the use of a different species of plant is really suspicious.

Apomecyna scorteccii Breuning, 1968: 90. Type locality: “Yemen: El Haurat, 1550 s.l.m.″.

Apomecyna scorteccii: Holzschuh & Téocchi, 1991: 301.

Material examined from Saudi Arabia: Wadi Daykah, 3-4.IV.1980, W. Büttiker; Wadi Shuqub/ Turabah, 1250 s.l.m., 21.IV.1980, W. Büttiker; Muhayel, 4.III.1986; Fayfa, 1240 s.l.m., 23.IX.1981, W. Büttiker (Holzschuh & Téocchi, 1991); Al Huabil, Wadi Reem, 18°06.981′N 42°13.939′E, 451 m, 20.X.2014, at light; Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m.., 2.IX.2015, at light; Al Baha, Shada Al Ala, 19°52.685′N 41°18.663′E, 851 s.l.m., 14.XI.2014, at light; Asir, Wadi Yabah, 19°16.484′N 41°48.557′E, 445 s.l.m., 15.IV.2016 (KSMA); Al Bahah prov., Jabal Shada al A′la, 1248 m, 19°51′40″N 41°18′16″E, 29.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 m, 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 m, 18°10′18″N 42°22′12″E, 1-4. IV.2017, at light, P. Rapuzzi lgt; Asir prov., 2 km N of Habeel, 489 m, 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991), Yemen (type locality).

Enaretta Thomson, 1864: 325. Type species: Enaretta castelnaudi Thomson, 1864 (by monotypy).

Enaretta caudata: Löbl & Smetana, 2010: 231; Ambrus & Grosser, 2012: 454.

Material examined: Holotype 〖: Saudi Arabia: Al Bahah prov., Jabal Shada al A′la, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III.2017, at light, P. Rapuzzi lgt (KSMA); Paratypes. 2♂ and 4♀: idem (CPR); 1〖: Saudi Arabia: Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 26.I.2015, at light; 1♀: Saudi Arabia: Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 3.VI.2014, at light (KSMA); 1〖: Saudi Arabia: Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, 1 dead specimen inside the pupal cell on Acacia sp., P. Rapuzzi lgt; 1♀: Saudi Arabia: Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt (CPR); 1〖: Oman, Dhofar prov., 5 km NE Dhalqut, 16°43′22.48″N 53°16′27.26″E, 22.IX.2011, R. Ambrus lgt (under Enaretta caudata Fåhraeus, 1872 in Ambrus & Grosser, 2012) (CPR); 1〖: Oman: Dhofar, Jabal al Qamar, 5 km NE Dhalqut, 16°43′22.48″N 53°16′27.26″E, 22.IX.2011, W. Grosser leg. (Walter Grosser private collection, Czech Republic). 2〖: Oman: Dhofar, Jabal al Qamar, 15 km NW Rakhyut, 16°46′7.54″N 53°20′13.92″E, 24.iX.2011, W. Grosser leg. (Walter Grosser private collection, Czech Republic); 1〖: Oman: Dhofar, Jabal Samhan, 15 km NW Jufa, 17°11′10.14″N 54°56′34.26″E, 27.IX.2011, W. Grosser leg. (Walter Grosser private collection, Czech Republic); 1♀: Oman: Dhofar, Jabal al Qamar, 15 km W Al Mughsayl, 16°51′44.56″N 53°43′18.38″E, 26.IX.2011; W. Grosser leg. (Walter Grosser private collection, Opava, Czech Republic).

Diagnosis: All of the specimens of E. samai sp. nov. from the Arabian Peninsula (Saudi Arabia and Oman) that we checked differ from E. caudata in several characters: elytral apex truncate and not flat in E. samai instead of extended to a flat apex in E. caudata; tooth at the external side of the elytral apex of A. caudata lacking in E. samai; elytra entirely covered with light pubescence in E. samai, while only partly covered in E. caudata; pronotal punctation denser and larger in E. samai than in E. caudata. The new species is much more like Enaretta somaliensis Breuning, 1938 from Somalia, from which it is easily separated by the pubescence pattern: in E. somaliensis the orange pubescence is darker and more developed, and the light pubescence is white instead of whitish, resulting in a more contrasting colour pattern than in E. samai. In addition, the pubescence is arranged in longitudinal stripes made by alternate denser and sparser areas in the new species whereas the pubescence is more homogeneous in density in E. somaliensis.

Etymology: We dedicate this new species to Gianfranco Sama (Cesena, Italy), who recently passed away, in gratitude for transmitting his knowledge to one of us (Pierpaolo Rapuzzi) as one of the greatest connaisseurs of the longhorn beetle fauna of the desert areas of the Arabian countries.

Description: Length 6.2 mm., width 3.0 mm. Body dark brown, entirely covered with light pubescence. Head small, densely covered with yellowish pubescence entirely hiding tegumen, that pubescence darker on vertex; frons square with deep medial furrow; eyes large, largely faceted; lower eye lobe very close to mandibles. Antennae short, reaching half of elytral length when extended posteriorly; scape and pedicel covered with orange pubescence, that on remaining antennomeres whitish. Pronotum bell-shaped, strongly constricted apically and larger at base; sides evenly curved; pubescence dense, predominantly yellowish, consisting of recumbent, strong bristles; presence of few thin, erect, black laterobasal setae; disc with two dark yellow/orange lines forming an inverted Y reaching apical margin; tegumen with sculpture coarse, consisting of big dense punctures, except apex densely covered with very fine punctures. Scutellum wider than long, covered with orange pubescence. Elytra short, oval, with apex truncate; pubescence orange on disk beside suture up until first quarter, then thinner and reaching apex in small thin line covered with dark yellow/orange pubescence; rest of elytra covered with whitish pubescence; both kinds of pubescences (whitish and yellow) arranged in denser and sparser longitudinal stripes giving a striped appearance; each elytron less pubescent at middle, giving the appearance of a darker median line interrupted near suture; with two small tufts of orange pubescence at base of elytra as well as two near apex, the latter forming a sort of arch. Legs short and robust, covered with dense whitish pubescence intermixed with black recumbent hairs; femora short, swollen in middle; hind and median tibiae strongly arched and bearing long fringe of orange erect setae; front tibiae straight and swollen subapically, lacking orange fringe.

Female: Very similar to the male, but with antennae shorter, reaching just before half of the elytral length when extended posteriorly.

Distribution: The new species is endemic to the Arabian Peninsula (Oman, Saudi Arabia, and probably Yemen).

Biology: We collected a dead specimen in a pupal cell inside a branch of Acacia sp.

Notes: The known specimens are very similar in color except that those from Oman are darker with sparser light pubescence and a wider and larger dark median stripe. In the absence of further distinctive characters, we consider this variation as intraspecific. Genus Enaretta was reported from the Arabian Peninsula based on specimens collected in Yemen and identified by K. Adlbauer as Enaretta caudata (Fåhroeus, 1872). This record was reported in the Catalogue of the Palaeartic Cerambycidae (Löbl & Smetana, 2010: 231) and in its subsequent revision by M. Danilevsky (2020: 324), where a record from Oman made by Grosser & Ambrus (2012: 454) was also reported. In our opinion this record of E. caudata from Yemen is strongly suspicious and most likely refers to the new species. However, we couldn′t examine these specimens, nor other from the same area to confirm our hypothesis.

Ogmodera Aurivillius, 1908: 149. Type species: Ogmodera sulcata Aurivillius, 1908 (by monotypy).

Ogmodera forticornis Breuning, 1942: 113-175. Type locality: ″Uganda (Turkana prov.)″.

Material examined from Saudi Arabia: Wadi Jiza, 28.X.1978, Filipponi (CGS); Al Bahah prov., Wadi Reyem, 19°50′28″N 41°22′34″E, 473 s.l.m., 7.IV.2019, at light, P. Rapuzzi lgt (CPR).

Distribution: Uganda (type locality); Ethiopia, Somalia, Kenya (Adlbauer & Beck, 2015). Arabian Peninsula: Saudi Arabia; Yemen (Danilevsky, 2020).

Ogmodera sulcata Aurivillius, 1908: 150. Type locality: “Kilimandjaro: Kibonoro in der Steppe″ (Tanzania).

Ogmodera sulcata: Holzschuh & Téocchi, 1991: 301; Adlbauer & Beck, 2015: 151.

Material examined from Saudi Arabia: Wadi Jizan (Gizan), 9.XI.1978, Filipponi; Hakimah, 14.IV.1971, collector not given (Holzschuh & Téocchi, 1991).

Distribution: Ethiopia, Somalia, Kenya, Tanzania. Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991; Adlbauer & Beck, 2015), Yemen (Danilevsky, 2020).

Ceroplesitae Thomson, 1860: 93. Type genus: Ceroplesis Audinet-Serville, 1835.

Ceroplesis Audinet-Serville, 1835: 93. Type species: Lamia aethiops Fabricius, 1775 (designated by Breuning, 1937).

Ceroplesis millingeni Pic, 1895: 77. Type locality: “Arabie″.

Ceroplesis atropos: Holzschuh, 1979: 294.

Ceroplesis millingeni: Holzschuh & Téocchi, 1991: 305.

Material examined from Saudi Arabia: Asir Mts., Qaraah Village, 2000 s.l.m., 18.IV.1976 (Holzschuh, 1979); Asir Mts., An Namas, 2400 s.l.m., on dead Acacia sp. branches, 8.IV.1983, C. Holzschuh; idem ex larva from Acacia sp., emerged 10.V-18.IX.1983; Abha, 2260 s.l.m., 4.X.1984, W Büttiker; Biljurshi, 2350 s.l.m., 13.IX.1983, W. Büttiker; Wadi Aziza, 18°13′N 42°28′E, 2400 s.l.m., 18.IX.1983, W. Büttiker; Lahifa, 7.IX.1984, W. Büttiker (Holzschuh & Téocchi, 1991); Al Baha, Wadi Turubah, 918 s.l.m., 20°12′610″N 41°17′056″E, 14.V.2011; Abha, 14.VII.1981; Al Baha, IX.2009; Al Baha, Rahwan, 10 km out Baha, 2272 s.l.m., 20°04.2272′N 41°27.013′E, 24.IV.2013; Al Baha, Wadi Rurubah, 1856 s.l.m., 20°10.749′N 41°19.396′E, 24.IV.2013; Taif, Mesan Balhareth, 18.VI.2002; Asir, Abha, Taminal, 2301 s.l.m., 18°01.353′N 42°45.814′E, 14.XI.2012; Al Baha, Al Mandaq, 8.VI.2003; Al Baha, Wadi Al Mandaq, 20°14.369′N 41°15.234′E, 1757 s.l.m., 3.VI.2012; Al Baha, Buljurashi, 1.VI.2012; Al Baha, El-Baher, 25.V.2011; Al Baha, Al-Zafeer, 14.V.2012; Al Baha, Ghabet Shobba, 19.VI.2011; Al Baha, Raghadan 20°00′575″N 41°26′974″E, 2222 s.l.m., 16.V.2010; Al Baha, Al Biljurashi, 19°48′407″N 41°41′437″E, 1931 s.l.m., 17.V.2010; Central prov., Riyadh area (doubtful data) (KSMA); Al Baha prov., Bani Hasan, 2258 s.l.m., 20°03′00′N 41°22′14″E, 12.IV.2016, ex larva in Acacia sp., emerged 30.IV.2016, P. Rapuzzi lgt; Al Baha prov., Al Qissaimah, 2156 s.l.m., 20°18′30″N 41°20′02″E, 12.IV.2016, ex larva in Acacia sp., emerged 11-16.V.2016, P. Rapuzzi lgt; Abha prov., Raedah reserve, 2809 s.l.m., 18°12′14″N 42°24′43″E, 16.IV.2016, ex larva in Acacia sp., emerged on 9.VI.2016, P. Rapuzzi lgt; Al Bahah prov., Wadi Tourabah, 20°11′16″N 41°18′38″E, 1812 s.l.m., 6.IV.2019, P. Rapuzzi lgt; idem, ex larva from Acacia sp., emerged 25.V.-25.VI.2019 (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991), Yemen (Danilevsky, 2020).

Ceroplesis elegans Gestro, 1889: 70-71. Type locality: “Yemen meridionale: Tes″.

Ceroplesis elegans: Holzschuh & Téocchi, 1991: 305.

Material examined from Saudi Arabia: Makkah, Al Uqdah, 21°07′N 40°25′E, 1780 s.l.m., 21-22. VIII.1984, W. Büttiker (Holzschuh & Téocchi, 1991); Al Baha, Wadi Turubah, 20°14′24″N 41°15′12″E, 1.VI.2011; Taif, Wadi Thi-Ghazal, 13.V.2012; Taif, Thaquif, 19.VI.2002; Al Baha, Wadi Turubah, 918 s.l.m., 20°12′610″N 41°17′056″E, 14.V.2011; Al Baha, Ghabat Amathan, 20°12′162″N 44°13′902″E, 1881 s.l.m., 19.V.2010; Shafa off Shafa Hwy., 21°08′21″N 40°21′05″E, 12.X.2010; Taif, Al Wesam district, 21°12′17″N 40°20′43″E, 11.X.2010; Al Baha, Al Mandaq, Wadi Turubah, 20°14.369′N 41°15.234′E, 1757 s.l.m., 3.VI.2012; Al Baha, Shuaib, Wadi Turubah, 20°10.749′N 41°19.396′E, 1856 s.l.m., 24.IV.2013; Al Taif, Wadi Thee Gazal, 21°04′29″N 40°21′41″E, 25.VIII.2008; Khamis Meshid, 24.VII.2002 (KSMA); Al Baha prov., Wadi Tourabah, 1830 s.l.m., 20°11′36″N 41°17′50″E, 12.IV.2016, ex larva in Acacia sp., emerged 11.V.2016, P. Rapuzzi lgt; Abha prov., Khamis Mushayt, 1944 s.l.m., 18°25′25″N 42°42′05″E, 17.IV.2016, ex larva in Acacia sp., emerged 1.VIII.-25.IX.2016, P. Rapuzzi lgt; Al Bahah prov., Wadi Tourabah, 20°11′16″N 41°18′38″E, 1812 s.l.m., 6.IV.2019, ex larva in Acacia sp., emerged 25.V.-25. VI.2019, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991), Yemen (type locality).

Titoceres Thomson, 1868: 201. New name for Ceratites Audinet-Serville 1835: 34. Type species: Ceratites jaspidea Audinet-Serville, 1835.

Ceratites Audinet-Serville, 1835: 34 [nec Ceratites De Haan, 1825 (Mollusca, Cephalopoda)]. Type species: Ceratites jaspidea Audinet-Serville, 1835 (monotypy).

Paratitoceres arabicus Breuning, 1962: 235. Type locality:″ N-Yemen: Sana Distr.″.

Titoceres arabicus: Holzschuh & Téocchi, 1991: 305.

Material examined from Saudi Arabia: Taif, 26.VI.1983, collector not given; Maraba, 30.IX.1978, W. Büttiker; Asir Mts., NW of Shuqayq, Muhail, 500 s.l.m., 4.IV.1983, emerged ex Acacia sp. 1.IX.1983, C. Holzschuh; Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 20.IV.2014, at light; 16.II.2014; idem, 3.VI.2014; idem, 12.XI.2015; idem, 14.XI.2015; idem, 15.XI.2015; Al Baha, Shada Al Ala, 19°51.066′N 41°18.037′E, 1325 s.l.m., 27.VII.2014, at light (KSMA).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991), Yemen (type locality).

Crossotitae Thomson, 1864: 64. Type genus: Crossotus Audinet-Serville, 1835.

Biobessa Gahan, 1898: 56. Type species: Biobessa beatrix Gahan, 1898 (by monotypy).

Biobessa holzschuhi Téocchi, 1991: 304. Type locality: “Saudi Arabia: Fayfa″.

Material examined from Saudi Arabia: Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light; idem, 16.II.2014; idem, 12.XI.2014; idem, 13.XI.2014; idem, 14.XI.2014; idem, 15.XI.2014; idem, 8.XII.2014; idem, 26.I.2015; Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m., 21.IV.2015, at light; Abha, Wadi Rida, 18°11′749″N 42°23′345″E, 1614 s.l.m., 26.IV.2014, at light; idem, 26.VIII.2014; Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 21.IV.2014, at light; Al Baha, Shada Al Ala, 19°50.575′N 41°18.691′E, 1666 s.l.m., 3.VI.2015, at light; idem, 27.VII.2014; idem, 2.IX.2014; Abha, Raydah, 18°11.679′N 42°23.691′E, 1851 s.l.m., 31.VII.2014, at light; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 12.XI.2014, at light (KSMA); Bahah prov., Jabal Shada al A′la, 1559 s.l.m., 19°50′16″N 41°18′38″E, 29.III.2017, ex larva in Carduus cf. pychnocephalus L., 1763 [Asteraceae], emerged 20.IV.2017, P. Rapuzzi lgt; idem, emerged 7.V.2017; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir prov., Wadi Mashwas, 18°10′06″N 42°22′01″E, 1289 s.l.m., 13.IV.2019, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Téocchi, 1991), Yemen (Danilevsky, 2020).

Crossotus Audinet-Serville, 1835: 52. Type species: Lamia plumicornis Audinet-Serville, 1835 (by monotypy).

Crossotus arabicus Gahan, 1896: 458. Type locality: “Lahej″ (Yemen).

Crossotus arabicus: Breuning, 1942: 16, 69; Villiers, 1968: 850; Holzschuh & Téocchi, 1991: 304.

Material examined from Saudi Arabia: Jidda, La Mecque (Breuning, 1942); Jiddat el Harash, XII.1946-I.1947, W. Thesiger; Haband, 16.III.1947, W. Thesiger; Dhufar, Jebel Qara, Wadi Geth, XI.1943, D. V. Fitzgerald; West Herkot, 12.II.1947, W. Thesiger (Villiers, 1968); S of Ryadh, Al Hair, 550 s.l.m., 17.IV.1983, ex larva in Acacia sp., emerged 12.V.1983, C. Holzschuh; SW of Ryadh, Sulayyil, 600 s.l.m., 11.IV.1983, ex larva in Acacia sp., emerged 28.VIII.1983, C. Holzschuh; SE of Jeddah, Shawag, 5.IV.1983, C. Holzschuh; Fayfa, 27-31. III.1983, C. Holzschuh; Gizan, at light, 25-26.III.1983, C. Holzschuh; Gizan, 25-26.III.1983, ex larva in Acacia sp., emerged 8.IV-31.VII.1983, C. Holzschuh (Holzschuh & Téocchi, 1991).

Distribution: Arabian Peninsula: Saudi Arabia (?), Oman, Yemen (Sudre et al., 2007).

Note: The records from Saudi Arabia are doubtful. We haven′t seen any specimens of this species collected in the Kingdom and very likely the old records belong to another species unknown at the time (Crossotus katbeh Sama, 2000), which is common and widespread in a large part of Saudi Arabia. We prefer to delete this species from the list of the Saudi Arabian species until confirmed records are provided.

Crossotus albicollis saudicola Téocchi, 1991. In Holzschuh & Téocchi, 1991: 304. Type locality: “Saudi Arabia: Fyfa″.

Crossotus albicollis Holzschuh, 1979: 294.

Crossotus albicollis ssp. saudicola: Holzschuh & Téocchi, 1991: 304.

Crossotus erlangeri ssp. saudicola: Sudre et al., 2007: 40 (comb. nov.).

Crossotus vagepictus: Danilevsky, 2010: 336 (part.).

Saudi Arabia: Asir Mts., Straße von Abha nach Gizan, km 28, Wadi ad Dilla, 700 s.l.m., 22.IV.1976, Wittmer & Büttiker (Holzschuh, 1979, under Crossotus albicollis Guérin, 1844); Fayfa, 27-31.III.1983, emerged from Acacia, 21.VIII.1983, C. Holzschuh; idem, emerged 12.IV.-11.IX.1983; NW of Shuqayq, Muhail, 500 s.l.m., 4.IV.1983, emerged ex Acacia, 12.IV.1983, C. Holzschuh; Muhayel, 4.III.1986; road from Abha to Gizan, km 28, Wadi Ad Dilla, 700 s.l.m., 22.IV.974, Wittmer & Büttiker (Holzschuh & Téocchi, 1991); Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Acacia sp., emerged 14.IV.-2.VI.2016, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Sudre et al., 2007), Yemen (Jabal Bura, NEE Al Hudaydah, 14°52′N 43°24′E, 240-350 s.l.m., 4.XI.2010, S. Kadlec lgt) (NMHP); Yemen, Wadi Zabid, W of Zabid, 14°09′N 43°32′E, 325 s.l.m., e.l., 22.III.2007, M. Rejzek leg. (CGS) (new record).

Note: Danilevsky (2020: 336) synonymized Crossotus erlangeri ssp. saudicola with C. vagepictus Fairmaire, 1886. In our opinion this synonymy is wrong. In fact, C. vagepictus has a dark line in the middle of the pronotum which is missing in C. e. saudicola for which the pronotum is entirely covered with whitish pubescence. Moreover, the tufts of pubescence on the elytral disc consists of golden bristles in C. vagepictus whereas it consists of black bristles in C. e. saudicola.

Crossotus kadleci Sama & Sudre, 2010: 953. Type locality: “Yemen, Lawdar NE Aden (13°53′N 45°48′E)″.

Material examined from Saudi Arabia: Abha, Raydah, 18°13.347′N 42°24.133′E, 2717 s.l.m., 25.IV.2014, at light (KSMA); Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°22′01″E, 16.IV.2016, ex larva in Acacia sp., emerged 11.IX.2016, P. Rapuzzi lgt; Al Baha prov., Wadi Tourabah, 1830 s.l.m., 20°11′36″N 41°17′50″E, 12.IV.2016, ex larva in Acacia sp., emerged 30.IV.2016, P. Rapuzzi lgt; Bahah prov., Wadi Tourabah, E of An Na′amah, 1826 s.l.m., 20°11′01″N 41°18′42″E, 30.III.2017, ex larva in Pistacia sp., emerged 1.IX.2017, P. Rapuzzi lgt; Bahah prov., Shada al Ala, 1692 s.l.m., 19°50′37″N 41°18′40″E, 8.IV.2019, P. Rapuzzi leg.; Asir prov., Wadi Mashwas, 18°10′06″N 42°22′01″E, 1289 s.l.m., 13.IV.2019, ex larva from Cadia purpurea, emerged 15.X.2019, P. Rapuzzi lgt; idem, emerged 21.III.2020 (CPR).

Distribution: Arabian Peninsula: Yemen (Sama & Sudre, 2010), Saudi Arabia (new record).

Note: The specimens collected in Saudi Arabia differ from C. kadleci in several characters, notably the color of the pubescence, which is dark grey instead of yellowish, and the missing longitudinal elytral stripes that are well marked in C. kadleci. However, more specimens of the Saudi Arabian population are needed to assess its real status.

Crossotus katbeh Sama, 2000. Type locality: “Arava valley: Israel″.

Crossotus subocellatus Holzschuh, 1979: 294. (Misidentification).

Crossotus strigifrons: Holzschuh & Téocchi, 1991: 304. (Misidentification).

Material examined from Saudi Arabia: Asir Mts., Straße von Abha nach Gizan, km 28, Wadi ad Dilla, 700 s.l.m., 22.IV.1976, W. Wittmer & W. Büttiker [under Crossotus subocellatus (Fairmaire, 1886); identification according to picture of specimen in Holzschuh′ (1979); the same specimens were identified as Crossotus strigifrons by Holzschuh & Téocchi (1991)]; Riyadh area; Muzahimiyah, Al Khararah, 24°24′24″N 46°14′40″E, 26.IV.2011, at light; Jizan-Aiban Sabia road, Shahdan, 17°28.266′N 42°51.191′E, 433 s.l.m., 13.XI.2012; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light; idem, 22.IV.2014; idem, 3.VI.2014; idem, 8.XII.2014; idem, 13.XI.2015 (KSMA); Ryadh prov., Farheat Sheaal, 591 s.l.m., 22°24′34″N 46°35′34″E, 17.IV.2016, ex larva in Acacia sp., emerged 1.V.2016, P. Rapuzzi lgt; idem, emerged 19.VI.2016; idem, emerged 12-21.VII.2016; idem, emerged 9.IX.2016; Riyadh prov., Ibex Reserve, W of Hotat Bani Tamim, 23°27′26″N 46°33′37″E, 11.IV.2017, ex larva in Acacia sp., emerged 15.V.2017, P. Rapuzzi lgt; Riyadh prov., Wadi Nessah, 24°13′54″N 46°53′37″E, 531 s.l.m., P. Rapuzzi lgt; Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 11.IV.2016, ex larva in Acacia sp., emerged 1-19.VI.2017, P. Rapuzzi lgt; idem, emerged 21.VII.2016; idem, emerged 28.VIII.2017; Al Baha prov., Wadi Yabah, 439 s.l.m., 19°16′32″N 41°48′33″E, 15.IV.2016, ex larva in Acacia sp., emerged 23.VI.2017, P. Rapuzzi lgt; Al Baha prov., Wadi The Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, ex larva in Acacia sp., 23.VI.2017, P. Rapuzzi lgt (CPR).

Distribution: Israel, Jordan. Arabian Peninsula: Yemen (Sudre et al., 2007), Saudi Arabia (new record).

Crossotus sublineatus Gestro, 1892: 788. Type locality: “Gubbet″ (Somalia).

Material examined from Saudi Arabia: Jazan prov., 12 km WSW of Marabah Dam, Wadi Reem, 139 s.l.m., 17°51′56″N 42°16′21″E, 3.IV.2017, ex larva in Acacia sp., emerged IX.2017, P. Rapuzzi lgt; Al Baha prov., Wadi Yabah, 439 s.l.m., 19°16′32″N 41°48′33″E, 15.IV.2016, ex larva in Acacia sp., emerged 10.X.2017, P. Rapuzzi lgt (CPR).

Distribution: Algeria, Djibouti, Ethiopia, Kenya, Mali, Morocco, Mauritania, Niger, Western Sahara, Somalia Sudan, Chad (Sudre et al., 2007). Arabian Peninsula: Saudi Arabia (new record).

Dichostates subocellatus Fairmaire, 1886: 435-462. Type locality: “Obok″ (Djibouti).

Crossotus subocellatus: Villiers, 1968: 850.

Material examined from Saudi Arabia: Buraiman près Jidda, 13.I.1946, E.S. Brown; Hejaz, 10.XI.1945, D.V. Fitzgerald; Mecca, 17.X-5.XI.1949, H.St.J. Philby (Villiers, 1968); Al Baha prov., Wadi Yabah, 439 s.l.m., 19°16′32″N 41°48′33″E, 15.IV.2016, ex larva in Acacia sp., emerged 23.VI.2017, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Asir prov., 6 km S Aineh, Wadi Reem, 17°52′34″N 42°16′39″E, 143 s.l.m., pupal cell in Acacia sp., 12.IV.2019, P. Rapuzzi lgt; idem, ex larva in Acacia sp., emerged 30.IV.2019 (CPR).

Distribution: Algeria, Egypt, Erythrea, Ethiopia, Lybia, Mali, Morocco, Mauritania, Western Sahara, Senegal, Djibouti, Ethiopia, Kenya, Niger, Nigeria, Somalia, Sudan, Chad, Tanzania. Doubtfulyl recorded from Syria. Arabian Peninsula: Saudi Arabia, Oman (Sudre et al., 2007), Yemen (Jabal Bura, NEE Al Hudaydah, 14°52′N 43°24′E, 261-600 s.l.m., 11.IV.2007, S. Kadlec lgt) (NHMP) (new record).

Dichostates strigifrons Fairmaire, 1886: 457. Type locality: “Soudan″.

Crossotus arabicus: Holzschuh, 1979: 294 (misidentification).

Crossotus strigifrons: Villiers, 1968: 850; Holzschuh & Téocchi, 1991: 303-304.

Material examined from Saudi Arabia: Jiddat el Harash, XII.1946-I.1947, A.R. Waterston; Asir, El Mahala, 19.IX.1944, W. Thesiger; K. Wir, 8.II.1947; K. Wir, 8.II.1947, W. Thesiger; West Shibun, 12.II.1947, W. Thesiger; Buraiman, près Jeddah, 24.I.1946, E.S. Brown; West Boi, 4.II.1947, W. Thesiger (Villiers, 1968); Asir Mts., Kamis-Mushayt, April 1971, leg. Di Lenardo (under Crossotus arabicus) (Holzschuh, 1979); Asir Mts., SE of Taif, Wadi Qust, 1400 s.l.m., 6.IV.1983, ex larva in Acacia sp. emerged from 16.VII. to 21.VIII.1983, C. Holzschuh; SE of Taif, Wadi Shuqub, 1300 s.l.m., 7.IV.1983, ex larva in Acacia sp., emerged 27.IV.1983, 22.VII.1983, 4.IX.1983, C. Holzschuh; Asir Mts., NW of Shuqayab, 500 s.l.m., 4.IV.1983, ex larva in Acacia sp., emerged 12.IV.-28.VIII.1983, C. Holzschuh; Gizan, 25-26.III.1983, ex larva in Acacia sp., emerged 7-28.VIII.1983, C. Holzschuh; Fayfa, 27-31.III.1983, ex larva in Acacia sp., emerged 16.VII.1983, C. Holzschuh; 85 km W of Najran, 10.IV.1983, ex larva in Acacia sp., emerged 25.VI-16.VII.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Asir, Salos Al Manzar, Wadi Baqrah, 424 s.l.m., 18°47.529′N 42°01.121′E, 12.X.2013; Asir, Al Huhall, Wadi Reem, 18°06.981′N 42°13′939′E, 451 s.l.m., 20.X.2014, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 20.IV.2014, at light; idem, 30.IV.2014; idem, 2-3.III.2015; idem, 27.VII.2015; idem, 12.XI.2015; idem, 14.XI.2015; idem, 16.XI.2015; idem, 18.XI.2015; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 3.VI.2014, at light; idem, 12.XI.2015; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light; idem, 20.IV.2014; idem, 3.VI.2014 (KSMA); Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Acacia sp., emerged, 20.IX.2016, P. Rapuzzi lgt; idem, emerged 9.VI.2016; Al Baha prov., Raghadan forest, 20°01′07″N 41°26′53″E, 2226 s.l.m., 13.IV.2016, ex larva in Acacia sp., emerged 28.VIII.2018, P. Rapuzzi lgt; Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 14 km N Khamis Musayht, Wadi Hashbal, 1926 s.l.m., 18°27′34″N 42°42′53″E, 2.IV.2017, at light, P. Rapuzzi lgt; idem, ex larva in Acacia sp., emerged 2.VI.2017, P. Rapuzzi lgt; idem emerged 28.VIII.2017; idem, emerged 4.XI.2017; idem, emerged 6.II.2018; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir, 28 km SSE Muhayl, 809 s.l.m., 18°17′55″N 42°07′41″E, 5.IV.2017, ex larva in Acacia sp., emerged 1.IX.2017, P. Rapuzzi lgt; Asir, 14 km N Khamis Mushait, Wadi Ibn Hashbal, 1936 s.l.m., 18°27′34″N 42°42′53″E, 2.IV.2017, ex larva from Acacia sp., emerged 26.VII.2018, P. Rapuzzi lgt; Asir, WSW of Abha, Wadi Marabah, 18°10′18″N 42°22′12″E, 1197 s.l.m., 13.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Djibouti, Ethiopia, Egypt, Israel, Jordan, Somalia, Sudan. Arabian Peninsula: Saudi Arabia, Yemen, Oman (Sudre et al., 2007).

Desmiphoritae THOMSON, 1860: 74. Type genus: Desmiphora Audinet-Serville, 1835.

Sophronica Blanchard, 1845: 160. Type species: Sophronica calceata Chevrolat, 1855 (designated by Chevrolat, 1855).

Sophronica talhouki Holzschuh, 1991: 306-307. Type locality: “Saudi Arabia: Gizan″.

Material examined from Saudi Arabia: Gizan, at light, 25-26.III.1983, C. Holzschuh (holotype); idem, 22.II.1979, A.S. Talhouk; Fayfa, at light, 27-31. III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Wadi Qounonah, 19°24.673′N 41°36.388′E, 348 s.l.m., 11.III.2012, at light (KSMA); Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°02′22″E, 16.IV.2016, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991); Yemen (about 12 km NW Manakhah, 24.VI-4.VIII.2003; Manakhah, Bani Mansu, 1500 s.l.m., 15°06′N 43°50′E, 15.X.2001) (new record).

Sophronica wittmeri Holzschuh, 1991: 308-309. Type locality: “Saudi Arabia: Gizan″.

Material examined from Saudi Arabia: Gizan, at light, 25-26.III.1983, C. Holzschuh (holotype); idem, ex dry twigs of Euphorbia triaculeata Forssk., 1913 [Euphorbiaceae], emerged between 22.V & 7.VIII.1983; Gizan, Sabiyah, 16.II.1985, W. Büttiker; Fayfa, 1240 s.l.m., 23.IX.1981, W. Büttiker; Fayfa, at light, 27-31.III.1983, C. Holzschuh; Asir Mts., NW of Shuqayq, Muhail, 500 s.l.m., 4.IV.1983, ex dry Euphorbia triaculeata, emerged 17.VI.1983, C. Holzschuh; Wadi Juwa, 8.II.1986, J. Grainger (Holzschuh & Téocchi, 1991).

Distribution: Arabian Peninsula: Saudi Arabia (Holzschuh & Téocchi, 1991), Yemen (Danilevsky, 2020).

Sybrinus Gahan, 1900: 12. Type species: Sybrinus commixtus Gahan, 1900 (by monotypy).

Sybrinus (Arabosybrinus) albosignatus Breuning, 1948: 16. Type locality: “Arabie: Aden″ (Yemen).

Sophroniella albosignata: Holzschuh & Téocchi, 1991: 306.

Material examined from Saudi Arabia: Fayfa, 27-31. III.1983, ex larva in Calotropis procera, emerged 6.V., 9.VII.1983; Hofuf, 28.VI.1981, collector not given (Holzschuh & Téocchi, 1991); Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, ex larva in Calotropis procera, emerged, 14.VI.2017, P. Rapuzzi lgt; idem, emerged 30.XI.2017; idem, emerged 28.VIII.2017; Asir prov., 3 km N of Habeel, Wadi Ksan, 462 s.l.m., 18°06′57″N 42°13′56″E, 12.IV.2019, in pupal cell in Calotropis procera, P. Rapuzzi lgt; idem, ex larva from Calotropis procera, emerged 1-28.II.2020, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen, Socotra Island (Hájek & Kabátek, 2012).

Sybrinus (Arabosybrinus) flavescens Breuning, 1948: 16. Type locality: “Arabie: Aden″ (Yemen).

Sophroniella flavescens: Holzschuh & Téocchi, 1991: 306.

Material examined from Saudi Arabia: Najran, 24.III.1983, ex larva in Calotropis procera, emerged 4.IV., 17.IV., 31.VII., 4.IX.1983 (Holzschuh & Téocchi, 1991); Wadi Targ, 19°37.385′N 42°18.020′E, 1317 s.l.m., 14.III.2012, at light (KSMA); Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia, Yemen (Hájek & Kabátek, 2012).

Eunidiini Téocchi, Sudre & Jiroux, 2010: 13. Type genus: Eunidia Erichson, 1843.

Eunidia Erichson, 1843: 262. Type species: Eunidia nebulosa Erichson, 1843 (monotypy).

Eunidia arabica Breuning, 1968: 89. Type locality: “Arabie: U. Ezone; U. El Kasaba″ (Yemen).

Eunidia arabica: Holzschuh & Téocchi, 1991: 298.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991).

Distribution: Yemen (type locality); Saudi Arabia (Danilevsky, 2020).

Eunidia breuningae Villiers, 1951: 201, fig. 1. Type locality: “Aïr: Agadez, Niger″.

Eunidia submarmorata Breuning, 1968: 91. Type locality: “Yemen, Est di Radà, Rocce Nere, 1880 s.l.m.″. Synonymy in Holzschuh & Téocchi, 1991.

Eunidia naviauxi Villiers, 1977: 168. Type locality: “Nord-Yémen, Achali″. Synonymy in Holzschuh & Téocchi, 1991.

Eunidia breuningae: Holzschuh, 1979: 293; Holzschuh & Téocchi, 1991: 298.

Material examined from Saudi Arabia: Asir Mts., Straße Abha nach Gizan, km 28, Wadi ad Dilla, 700 s.l.m., 22.IV.1976, Wittmer & Büttiker (Holzschuh, 1979); Asir Mts., Khamis Mushait, 15.VII.1981, A.S. Talhouk; Wadi Gaanah, 13-14.II.1980, W. Büttiker; Wadia Majarish, 1050 s.l.m., 6.IV.1980, W. Büttiker; AashSharaiy, 23.IX.1978, W. Büttiker; road from Abha to Gizan, km 28, Wadi Ad Dilla, 700 s.l.m., 2.X.1979, W. Büttiker; Gizan, 25-26.III.1983, emerged from Acacia sp., C. Holzschuh; Asir Mts., NW of Shuqayq, Muhaul, 500 s.l.m., at light, 4.IV.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Asir, Al Magardah, Wadi Yabah, 19°14.911′N 41°47.255′E, 402 s.l.m., 11.X.2013, at light; Wadi Qounonah, 19°24.673′N 41°36.388′E, 348 s.l.m., 11.III.2012; Abha, Taminah Dam, 18°01.353′N 42°45.814′E, 2301 s.l.m., 14.XI.2012; Al Huabil, Wadi Reem, 18°06.981′N 42°13.939′E, 451 s.l.m., 20.X.2014, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 23.IV.2014, at light; idem, 27.VII.2014; idem, 23.VIII.2014; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 21.IV.2014, at light; idem, 3.VI.2014; idem, 27.VII.2014; idem, 26.I.2015; Al Baha, Shada Al Ala, 19°52.717′N 41°18.712′E, 825 s.l.m., 15.IX.2014, at light; Al Baha, Shada Al Ala, 19°52.685′N 41°18.663′E, 851 s.l.m., 14.XI.2014, at light (KSMA); Al Baha prov., Wadi Yabah, 439 s.l.m., 19°16′32″N 41°48′33″E, 15.IV.2016, ex larva in Acacia sp., emerged 9.VI.-12.VIII.2016, P. Rapuzzi lgt; Al Baha, Mekwah, Shada al Ala, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III.-1.IV.2017, ex larva from Acacia sp., emerged X.2019, G. Magnani lgt; Al Baha prov., Wadi Thee Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, ex larva in Acacia sp., emerged 12.X.2016, P. Rapuzzi lgt; idem, at light, 13.IV.2016; Al Baha prov., 10 km NNW Al Makhwah, 630 s.l.m., 19°50′47″N 41°22′40″E, 31.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada al A′la, 1248 s.l.m., 19°51′40″N 41°18′16″E, 29.III.2017, at light, P. Rapuzzi lgt; Al Bahah prov., 5 km N Al Makhwah, 464 s.l.m., 19°43′47″N 41°26′47″E, 30.III.2017, at light, P. Rapuzzi lgt; Asir prov., 6 km S of Ainah, 200 s.l.m., 17°55′28″N 42°15′20″E, 3.IV.2017, at light, P. Rapuzzi lgt; Jazan prov., 12 km WSW of Marabah Dam, Wadi Reem, 139 s.l.m., 17°51′56″N 42°16′21″E, 3.IV.2017, ex larva in Acacia sp., emerged IX.2017, P. Rapuzzi lgt; Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 6 km S of Ainah, 200 s.l.m., 17°55′28″N 42°15′27″E, 3.IV.2017, ex larva in Acacia sp., emerged 4.VIII.2017, P. Rapuzzi lgt (CPR).

Distribution: Senegal, Niger, Chad, Egypt, Ethiopia. Arabian Peninsula: Saudi Arabia, Yemen, Oman (Adlbauer & Beck, 2015).

Eunidia haplotrita Aurivillius, 1911: 32. Type locality: “Abyssinien, Harrar″ (Ethiopia).

Eunidia haplotrita: Holzschuh & Téocchi, 1991: 298.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Al Huabil, Wadi Reem, 18°06.981′N 42°13.939′E, 451 s.l.m., 20.X.2014, at light; Abha, Wadi Rida, 18°11′749″N 42°23′345″E, 1614 s.l.m., 31.VII.2015, at light; Al Baha, Shada Al Ala, 19°50.710′N 41°18.267′E, 1474 s.l.m., 2.IX.2015, at light (KSMA); Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir prov., N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt; Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt; Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°02′22″E, 16.IV.2016, at light, P. Rapuzzi lgt (CPR).

Distribution: Niger, Sudan, Ethiopia, Erythrea, Djibouti, Somalia, Kenya, Tanzania. Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015), Yemen (Danilevsky, 2020); Oman (Dhofar, 19 km WSW Al Mughsayl, 7-12.VIII.2014, Prepsl leg.) (new record).

Eunidia nebulosa Erichson, 1843: 262. Type locality: ″Angola″.

Eunidia nebulosa: Holzschuh & Téocchi, 1991: 298.

Material examined from Saudi Arabia: Fayfa, 27-31. III.1983, at light, C. Holzschuh (Holzschuh & Téocchi, 1991); Asir, Wadi Marabah, 18°10′18″N 42°22′12″E, 1197 s.l.m., 1-4.IV.2017, at light, P. Rapuzzi leg.; Asir, 2 km N Habeel, 18°07′12″N 42°13′55″E, 489 s.l.m., 5.IV.2017, at light, P. Rapuzzi leg.; Asir, Wadi Marabah, WSW of Abha, 18°10′18″N 42°22′12″E, 1197 s.l.m., at light, P. Rapuzzi leg.; Asir, Jebel Soudah, 18°11′37″N 42°23′50″E, 1900 s.l.m., 13.IV.2019, P. Rapuzzi lgt (CPR).

Distribution: Senegal, Niger, Ethiopia, Somalia, Ivory Coast, Cameroon, Gabon, Republic of the Congo, Kenya, Tanzania, Mozambique, Angola, Namibia, Botswana, Republic of South Africa (Adlbauer & Beck, 2015); Chad: Salamat, 60 km SW of Am Timan, 420 s.l.m., 25.VI.2016 (CPR) (new record). Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015), Oman (Ambrus & Grosser, 2012), Yemen (Danilevsky, 2020).

Eunidia Kristenseni Aurivillius, 1911: 29-30. Type locality: “Abyssinien: Harrar; British Ostafrika: Ikutha″ (Ethiopia and Kenya).

Eunidia kristenseni: Villiers, 1968: 849; Holzschuh & Téocchi, 1991: 300.

Material examined from Saudi Arabia: Buraiman, près Jedda, 7.I.1946, E. S. Brown (Villiers, 1968); Jizan, Fayfa, 21.III.2014, at light; Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 2.IX.2015, at light; Al Baha, Shada Al Ala, 19°52.685′N 41°18.663′E, 851 s.l.m., 15.XI.2014, at light; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 13.XI.2015, at light (KSMA); Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4. IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Senegal, Niger, Chad, Sudan, Ethiopia, Erythrea, Djibouti, Somalia, Kenya, Tanzania, Namibia, Botswana. Arabian Peninsula: Saudi Arabia, Yemen (Adlbauer & Beck, 2015).

Eunidia Thomseni Distant, 1898: 377. Type locality: “Transvaal, Pinnaars River, Sta Lucia Lake, South Africa″.

Eunidia thomseni: Holzschuh & Téocchi, 1991: 300.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Al Baha prov., Wadi Thee Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, ex larva in Ziziphus sp., emerged 12.VIII.2016, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°02′22″E, 16.IV.2016, at light, P. Rapuzzi lgt; Asir prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°02′01″E, 1-4.IV.2017, ex larva in Acacia sp., emerged 9-15.V.2017, P. Rapuzzi lgt; idem, emerged 1-16.VI.2017; idem, emerged 14.IX.2017 (CPR).

Distribution: Senegal, Niger, Chad, Ethiopia, Somalia, Cameroon, Central African Republic, Congo, Uganda, Kenya, Tanzania, Malawi, Mozambique, Zimbabwe, Botswana, Namibia, Republic of South Africa (Adlbauer & Beck, 2015), Benin (Téocchi et al., 2016). Arabian Peninsula: Saudi Arabia, Yemen (Adlbauer & Beck, 2015).

Mesosini Mulsant, 1839: 165. Type genus: Mesosa Audinet-Serville, 1835.

Lachnia (Coptops) Audinet-Serville, 1835: 64. Type species: Lachnia parallela Audinet-Serville, 1835 (= Lamia ædificator Fabricius, 1793) (by monotypy).

Lamia aedificator Fabricius, 1793: 275. Type locality: “India orientalis″.

Coptops aedificator: Villiers, 1968: 849; Holzschuh & Téocchi, 1991: 296; Dawah et al., 2013: 1-9.

Material examined from Saudi Arabia: Mukalla (Villiers, 1968); An Namas, 10.IV.1983, A.S. Talhouk et al.; Fayfa, 30.VII.1982, A.S. Talhouk; Fayfa, 27-31. III.1983, ex larva from Acacia sp., C. Holzschuh (Holzschuh & Téocchi, 1991); Jazan Agriculture Research Station at Abu Aresh, Almahdage village, 17°00′N 42°45′E, 50 s.l.m., from Mangifera indica L., 1753 [Anacardiaceae]; Aseer, Mraaba, 17°54′N 42°23′E, 80 s.l.m., from Mangifera indica (Dawah et al., 2013); Riyadh VIII.1988; Jizan, Fayfa, 21.III.2014, at light; Jizan, Aiban, Wadi Gowra, 17°17.569′N 43°04.211′E, 451 s.l.m., 11.XI.2012; Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 21.IV.2014, at light; idem, 23.IV.2014; idem, 16.II.2014; idem, 17.X.2014; idem, 2.IX.2015; idem, 15.XI.2015; Al Baha, Shada Al Ala, 19°51.762′N 41°13.089′E, 1225 s.l.m., 3.VI.2014, at light; idem, 1.IV.2014; Al Baha, Shada Al Ala, 19°50.411′N 41°18.686′E, 1611 s.l.m., 3.VI.2014, at light; idem, 15.XI.2015; Al Baha, Shada Al Ala, 19°50.329′N 41°18.604′E, 1563 s.l.m., 14.XI.2015, at light; Jizan, Abu Arish, All Abu Ashwarl, 10.V.2018, 16°59.923′N 42°50.059′E, 73 s.l.m., at light (KSMA); Al Bahah prov., Jabal Shada Al A′la, 1695 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, adult in pupal cell on Ficus palmata, P. Rapuzzi lgt; idem, ex larva in Ficus palmata, emerged, 12.VII.2017; idem, emerged 11-30.XI.2017; Al Bahah prov., Jabal Shada Al A′la, 1695 s.l.m., 19°50′32″N 41°18′42″E, 29.III.2017, ex larva in Ficus palmata, emerged 01.VIII.2018, P. Rapuzzi lgt; Al Bahah prov., Jabal Shada Al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, at light, P. Rapuzzi lgt; Bahah prov., Thee Ain, 759 s.l.m., 19°55′47″N 41°26′38″E, 13.IV.2016, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, at light, P. Rapuzzi lgt; Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, in pupal cell on Ficus sp., P. Rapuzzi lgt; Asir prov., Jebel Soudah, Raydah reserve, 1632 s.l.m., 18°11′45″N 42°23′21″E, 2.IV.2017, ex larva in Ricinus communis L., 1753 [Anacardiaceae], emerged 14.VI.2017, P. Rapuzzi lgt; idem, emerged 21.IX.2017 (CPR).

Distribution: India, Cameroon, Cape Verde, Central African Republic, China, Comoroes, Congo, Djibouti, Ethiopia, Gabon, Ivory Coast, Mauritius, Reunion Island, Kenya, Liberia, Madagascar, Malawi, Namibia, Nigeria, Philippines, Principe Islands, Senegal, Seychelles, Somalia, Republic of South Africa, Sri Lanka, Taiwan, Tanzania, Uganda, Zambia (Saha et al., 2013), Sierra Leone (Téocchi & Mourglia, 1987), Togo (Adlbauer & Mourglia, 1999). Arabian Peninsula: Saudi Arabia, Yemen (Saha et al., 2013), Oman (Ambrus & Grosser, 2012).

Monochammidae Gistel, 1848: 9. Type genus: Monochamus Dejean, 1821.

Anoplophora Hope, 1839: 43. Type species: A. stanleyana Hope, 1839 (by monotypy).

Cerosterna glabripennis Motschulsky, 1853: 48. Type locality: “China bor., env. de Pékin“.

Material examined from Saudi Arabia: Al Saleh Al Taweel Farm; Al Afif, W Riyadh, KSA, 8.IV.2005; Baha (KSMA).

Distribution: Austria, Czech Republic, Croatia, Finland, France, Great Britain, Germany, Italy, The Netherlands, Slovakia, China, North Korea, South Korea, Turkey, Australia, North America (Danilevsky, 2020); Arabian Peninsula: Saudi Arabia (new record).

Phytoeciares Mulsant, 1839: 165. Type genus: Phytoecia Dejean, 1835.

Phytoecia Dejean, 1835. Type species: Saperda cylindrica Fabricius, 1775 (= Cerambyx cylindricus Linnaeus, 1758), designated by Breuning (1952).

Phytoecia (Pseudoblepisanis) Breuning, 1950: 31. Type species: Saperda analis Fabricius, 1781 (by original designation).

Material examined: Holotype 〖: Saudi Arabia: Baha, Wadhi Gala, 20°08′079″N 41°20′561″E, 1900 s.l.m., 16.V.2011 (KSMA); Paratypes. 1〖: idem (KSMA); 1♀: Al Bahah prov., Jabal Shada Al A′la, 1559 s.l.m., 19°50′16″N 41°18′38″E, 29.III.2017, flying, P. Rapuzzi lgt (CPR).

Diagnosis: The new species mostly resembles Phytoecia (Pseudoblepisanis) cylindricollis (Kolbe, 1893) from Western Africa. However, it is easily separated by the larger black portion of the elytra, the antennae totally black instead of partly reddish in the central portion, and by the ventral body colour predominantly black instead of orange.

Etymology: The name indicates the origin of the species from the Arabian Peninsula.

Description: Length 12.0 mm., width 3.5 mm (holotype). Body orange, except antennae and apical part of elytra black, ventral surface (except part of abdominal segments) black with several short silvery erect setae. Head wide, with frons transverse and large; punctation dense and very deep, consisting of granules umbilicate and occasionally fused with adjoining ones; with very thin carina between eyes; presence of few erect dark setae on occiput and just behind eyes, and few light setae close to mandibles, the latter long and darkened toward apex. Antennae black, longer than body when extended posteriorly with last three segments exceeding the apex, slightly enlarged toward apex; antennomeres 1 to 3 bearing several short black setae on inner side, and antennomeres 4 to 6 bearing few short black erect setae on inner side of apex. Pronotum cylindrical, slightly longer than wide; lateral sides fairly straight, except middle portion slightly sinuate; apical and basal margins very thin, impunctate; disc deeply and densely punctate, except small swollen and shiny area in middle; pubescence denser laterally on margin and subbasally, consisting of very short and very sparse erect brownish bristles. Scutellum apically rounded, nearly impunctate, bearing few golden erect setae denser near apical margin. Elytra parallel-sided, slightly constricted toward apex, the latter rounded; punctation consisting of several longitudinal rows of punctures, the latter becoming gradually smaller toward apex; tegumen densely microsculptured, making elytra appearing dull; presence of few short and erect dark setae along lateral margins, these setae on orange parts and nearly black on black portion of elytra. Legs stumpy, orange except for slightly darkened tarsi and subapical portion of hind tibiae; middle tibiae on external side bearing short fringe of dense golden hairs as well as small tooth medially.

Female: Similar to male except body slightly stouter and antennae shorter, exceeding the elytral apex with the last two segments when extended posteriorly.

Distribution: The species is apparently endemic to Southwest Saudi Arabia.

Biology: The female collected in Jabal Shada A′la was flying around a plant of Carduus cf. pychnocephalus, possibly the host of the species.

Note: This species is very interesting because it is the first record for the subgenus Pseudoblepisanis for the Arabian Peninsula, the Asian continent, and the Palaeartic Region.

Prosopoceritae Thomson, 1864: 72. Type genus: Prosopocera Blanchard, 1845.

Prosopocera Blanchard, 1845: 160. Type species: Lamia fronticornis Fabricius, 1781 (= Cerambyx bipunctatus Drury, 1773) (by monotypy).

Alphitopola Thomson, 1857: 299. Type species: Alphitopola lactea Thomson, 1857 (by monotypy).

Alphitopola unicolor (Gahan, 1898): 40-59. Type locality: “Samburu, British East Africa″ (Kenya).

Prosopocera albescens: Holzschuh & Téocchi, 1991: 297.

Saudi Arabia: Fayfa, 27-31.III.1983, ex larva from Fabaceae (not Acacia), C. Holzschuh; 81 km S of Biljurshi, 2000 s.l.m., VIII.1979, G. Vogel (Holzschuh & Téocchi, 1991); Al Baha, Shada Al Ala, 19°52.598′N 41°18.672′E, 892 s.l.m., 14.XI.2015, at light; idem, 2.IX.2015 (KSMA).

Distribution: Ethiopia, Somalia, Kenya. Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015).

Dalterus Fairmaire, 1892: 125-126. Type species: Dalterus auberti Fairmaire, 1892 (by monotypy).

Prosopocera dejeani Gahan, 1890: 297-328, pl. IX. Type locality: “Natal″ (Republic of South Africa).

Material examined from Saudi Arabia: Abha prov., Wadi Mashwas, 1289 s.l.m., 18°10′06″N 42°22′01″E, 16.IV.2016, ex larva in Acacia sp., emerged 1.VIII.2016, P. Rapuzzi lgt; idem, 1-4. IV.2017, ex larva in Acacia sp., emerged 3.VII.2017; Al Baha prov., Shada al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Acacia sp., emerged 28.VII.-7.VIII.2016, P. Rapuzzi lgt; Asir prov., 28 km SSE of Muhayl, 809 s.l.m., 18°17′55″N 42°07′41″E, 5.IV.2017, ex larva in Acacia sp., emerged 20.VIII.2017, P. Rapuzzi lgt (CPR).

Distribution: Somalia, Democratic Republic of the Congo, Tanzania, Mozambique, Zimbabwe, Botswana, Republic of South Africa (Adlbauer, 1997). Arabian Peninsula: Saudi Arabia (new record).

Pteroplitae Thomson, 1860: 73. Type genus: Pteroplius Lacordaire, 1830.

Niphona Mulsant, 1839: 169. Type species: N. picticornis Mulsant, 1839 (by monotypy).

Niphona appendiculata Gerstaecker, 1871: 78. Type locality: “Vom See Jipe″ (Tanzania).

Niphona appendiculata: Holzschuh & Téocchi, 1991: 302.

Material examined from Saudi Arabia: Fayfa, 27-31.III.1983, ex larva in Ficus salicifolia, emerged 17.IV.1983, 7.VIII.1983, 4.IX.1983, C. Holzschuh (Holzschuh & Téocchi, 1991); Asir prov., Jebel Soudah, 1632 s.l.m., 18°11′45″N 42°23′21″E, 2.IV.2017, malaise traps, P. Rapuzzi lgt (CPR); Al Baha, Shada Al Ala, 1358 s.l.m., 19°50′51″N 41°18′06″E, 14.IV.2016, ex larva in Ephedra pachyclada Boiss., 1884 [Ephedraceae], emerged 29.VIII.-25.IX.2016, P. Rapuzzi lgt (CPR).

Distribution: Ethiopia, Somalia, Senegal, Sierra Leone, Ivory Coast, Cameroon, Central African Republic, Congo, Kenya, Tanzania, Angola, Malawi, Mozambique, Zimbabwe, Botswana, Republic of South Africa. Arabian Peninsula: Saudi Arabia (Adlbauer & Beck, 2015).

Pterolophia Newman, 1842: 370. Type species: Mesosa (?) bigibbera Newman, 1842 (by original designation).

Pterolophia (Arabopraonetha) Téocchi, 1991: 302. Type species Pterolophia arabica Téocchi, 1991 (by monotypy).

Pterolophia (Arabopraonetha) arabica Téocchi, 1991: 302. Type locality: “Saudi Arabia: Gizan″.

Material examined from Saudi Arabia: Gizan, at light, 25-26.III.1983, C. Holzschuh (holotype); Al Ammariyah, 24°82.313′N 46°45.469′E, 26.XI.2013; Asir, Wadi Quonunah, 19°26.099′N 41°36.486′E, 405 s.l.m., 11.X.2013; Wadi Baqrah, 18°47.476′N 41°56.310′E, 331 s.l.m., 13.III.2012, at light; Wadi Qounonah, 19°24.673′N 41°36′388′E, 348 s.l.m., 11.III.2012, at light (KSMA); Asir prov., 2 km N of Habeel, 489 s.l.m., 18°07′12″N 42°13′55″E, 3.IV.2017, at light, P. Rapuzzi lgt (CPR).

Distribution: Arabian Peninsula: Saudi Arabia (type locality), Yemen (Danilevsky, 2020).

Tragocephalites Thomson, 1857: 25. Type genus: Tragocephala Laporte, 1840.

Dinocephalus Péringuey, 1899, p. 323. Type species: Dinocephalus ornatus Péringuey, 1899 (monotypy).

Dinocephalus heissi Holzschuh, 1991: 296-297. Type locality: “Saudi Arabia: Asir National Park″.

Material examined from Saudi Arabia: Asir, Asir National Park, Abha distr., 2200 s.l.m., 7.VI.1982, E. Heiss (holotype); Asir prov., Wadi Marabah, 1197 s.l.m., 18°10′18″N 42°22′12″E, 1-4.IV.2017, on Acacia sp., P. Rapuzzi lgt; Al Bahah prov., Jabal Shada Al A′la, 1286 s.l.m., 19°51′42″N 41°18′15″E, 08.IV.2019, on Acacia sp., P. Rapuzzi lgt; idem, 9.IV.2019 (CPR).

Distribution: Ethiopia (Adlbauer et al., 2008). Arabian Peninsula: Saudi Arabia (type locality).

Kerochariesthes Téocchi, 1990: 19. Type species: Pseudo-chariesthes variegata Breuning, 1939 (by monotypy).

Freapomecyna holzschuhi Téocchi, 1991: 300-301. Type locality: “Saudi Arabia: Fayfa″.

Kerochariesthes holzschuhi: Adlbauer, 2002: 19.

Material examined from Saudi Arabia: Fayfa, at light, 27-31.III.1983, C. Holzschuh (Holzschuh & Téocchi, 1991).

Distribution: Arabian Peninsula: Saudi Arabia (type locality), Yemen (Löbl & Smetana, 2010).