Notes on Early Land Plants Today. 91*. Types and original material of liverworts (Marchantiophyta) in Herbarium Haussknecht (JE) collected in Central America

Lars Söderström; Jörn Hentschel

doi:10.25227/linbg.25015

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29 June 2023 Notes on Early Land Plants Today. 91. Types and original material of liverworts (Marchantiophyta) in Herbarium Haussknecht (JE) collected in Central America

Lars Söderström, Jörn Hentschel

Author Affiliations +

Lindbergia, 2023(1): (2023). https://doi.org/10.25227/linbg.25015

Abstract

For a forthcoming catalogue of liverworts and hornworts of Central America, types and other original material from Central America deposited in the Herbarium Haussknecht (JE) are listed. We have tried to identify the earliest valid typification of all listed taxon names and lectotypifications for 36 names, as well as second-step typifications for five further names. One new synonym is proposed.

Introduction

Early authors of plant names often noted only the origin of the plants they described, without specifying any type (holotype). They very often described the taxon from several gatherings or made duplicates which were distributed to several herbaria. For all those names, unless it can be demonstrated that there exist only ‘a single specimen conserved in one herbarium’ (ICN2018 Art. 8.1; Turland et al. 2018), a lectotype must be selected. After 1990, a holotype must be specified for a name to be validly published. Thus, there is no need to select a lectotype for new names unless it can be demonstrated that the selected holotype consists of several taxa.

Before 1 January 2001, the mention of a ‘type’ or ‘holotype’ citing a single gathering can be accepted as a lectotypification, but after that date a statement as ‘lectotype here designated’ or equivalent is required (ICN2018 Art. 7.11; Turland et al. 2018).

It can be difficult to trace all possible type material among the world's herbaria, and if the original publication does not state what the holotype is, it is best to lectotypify the name. If only one possible type is known to the typifying author, but it can not be ruled out that there exist more material not traced, the ‘best practice’ recommended by McNeill (2014) should be followed and a statement like ‘lectotype here designated, if not a holotype’ would permanently attach the name to a single specimen, irrespective of if more possible type material will be found later or not. See Renner (2021) for further discussion on typification.

Many typifications, inadvertent or deliberate, are ‘hidden’ in publications not well known to the wide audience. Thus, later authors, unaware of the earlier typification, select a lectotype that may or may not be the same as the previous lectotypification. Unless it can be demonstrated that the earlier lectotypification is against the protologue, the later one must then be regarded as superfluous and disregarded.

In the first six volumes of Index Hepaticarum, Charles Edmond Bradlaugh Bonner identified type material, mostly from Conservatoire et Jardin botaniques de la Ville de Genève (G) where he was based. This has sometimes (Schuster 1980, Yuzawa 1991) been accepted as inadvertent lectotypification, but Engel and Smith Merrill (2019) argue that he did not intend to select any type. However, Renner (2021) did not accept Engel and Smith Merrill's arguments, and showed for Australasian Plagiochila (Dumort.) Dumort. that in most cases the same specimen would be used as lectotype also if Bonner's types were not accepted. Here, we accept Bonner's typification if it adds any information/restriction to the text in the protologue. However, if we are aware of any later lectotypification, supporting or disagreeing with Bonner, we make a note on it.

In many cases, material was separated from the main packet for microscopic examination and stored in some sort of microslide collection without a substantial reference from which collection it has been extracted. Often, when a new taxon is described, microcharacters are taken from the slides while macrocharacters must be taken from the larger sample. However, if the microscopically examined material is not returned to the original packet, and the connection is not clearly stated, the microslide creates a new sample and is in several herbaria, like JE, filed separately with different herbarium/barcode numbers. Our recommendation is that all examined material should be returned to the packet it was taken from instead of stored separately. In cases, where a microslide exists without a direct connection to where it was extracted, we recommend that the larger specimen is selected as lectotype whenever possible.

For a forthcoming catalogue of liverworts and hornworts of Central America, we here list all hepatic names with types or other original material from Central America that are known to occur in Herbarium Haussknecht (JE), both validly and invalidly published names. Many taxa were described by Theodor Herzog from material collected by Paul C. Standley and sent to him. One problem is that Herzog often gave a specimen a forma name without formally describing them, thus creating many nomina nuda. Most of the corresponding specimens are deposited in JE and/or US. Those names are effectively published and some of them cited in subsequent literature and thus we include also them. Here, we try to list all sorts of type or original material (for definitions of different type categories, Table 1).

As invalidly published names do not have types in the sense of the International Code of Nomenclature for Algae, Fungi, and Plants (ICN2018; Turland et al. 2018), we here instead use the term ‘original material’ for what could be potential type material if the name would have been validly published (i.e. a bit broader sense than the definition of ‘original material’ in ICN2018, Table 1). In case, there is no clear holotype mentioned in the original publication, we repeat the original citation of type material, and the lectotypification we are aware of, and cite lectotype material and its place of deposition. In case, the types are available in digital herbaria, we cite the barcode number and give the URL to the specimen.

We are taking a strict view on what a holotype is and accept only cases when it was definitely indicated in the protologue. In many cases, publications before 2001 cite material as ‘holotype’ or ‘type’ without being identified as such in the original publication. We accept this as inadvertent lectotypifications. However, if we are aware of only one possible type specimen in a single herbarium (c.f. ICN2018 Art. 8.1; Turland et al. 2018) we note that it may be a holotype as defined by the code unless more type material can be identified. In many cases in the past, a lectotype was selected from several syntypes without specifying a herbarium, or with several duplicates in the specified herbarium. In such cases a second-step lectotypification is needed to select a single specimen conserved in one herbarium (c.f. ICN2018 Art. 9.17; Turland et al. 2018).

We here lectotypify 36 names and make second-step lectotypification to another five names. We have not been able to find all types that should be in JE as many are on loan to other institutions and mostly lack barcode numbers. Also, the effort has been to mainly database types and thus original material of many non-valid names not databased and bar-coded yet. We have not very actively searched the herbarium for possible isotypes, syntypes, paratypes, etc. and there is certainly more type material of some kind in JE that we have not located yet. However, we think we have located all holotypes and lectotypes.

In this list, we mark illegitimate names with one asterisk (*) and invalid names with double asterisks (**). Unless a clear holotype is specified in the original publication, we here also cite the complete original material from which a type can be selected. We also cite such material in case of invalid names (Table 1). For all names that are not currently accepted, we provide the currently accepted name with a reference to the synonymisation.

Typification notes

Adelanthus parvus Herzog, Rev. Bryol. Lichénol. 11: 18, 1939 (Herzog 1938)

Original citation: Costa Rica, Cerro de Las Vueltas, 2700–3000 m., on tree ([Standley] n. 43.723/c).

Lectotype (Grolle 1972: 343 as ‘holotype’) if not holotype: Costa Rica, Prov. de San José, 2700–3000 m, Standley 43723c (JE04002323 https://je.jacq.org/JE04002323).

Currently accepted name: Adelanthus lindenbergianus (Lehm.) Mitt. (cf. Grolle 1972).

Aphanolejeunea angustissima Steph., Sp. Hepat. (Stephani) 5: 859, 1916 (Stephani 1916).

Original type citation: Hab. Guatemala.

Lectotype (Bonner 1962b: 235, also Pócs and Bernecker 2009 as ‘holotype’): Guatemala, Cobán, Türckheim ex hb. Levier 5826 (G00067015, www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=113712&lang=en, isolectotype JE04001106 https://je.jacq.org/JE04001106).

Note. Since the type material is poor and belongs to two species (Pócs and Bernecker 2009), the name should be rejected, as formally proposed by Söderström et al. (2022).

Table 1.

Definitions of type categories and related terms used in this publication (ICN2018; Turland et al. 2018).

Bazzania chimborazensis Spruce f. *cellulis-apicalibus*** Herzog, Rev. Bryol. Lichénol. 11(1): 24, 1939 (Herzog 1938), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa Rica, Cerro de La Carpintera, Prov. Cartago, 1500–1850 m., on tree ([Standley] n. 35.660, f. cellulis apicalibus 27 µ metientibus).

Original material: Costa Rica, Cerro de La Carpintera, Province of Cartago; altitude 1500–1850 m, February 1924, Standley 35660 (JE s.n., US04665880 http://n2t.net/ark:/65665/31a04163f-a345-4cfc-8a62-6b4586bd9 720).

Note. The original material in JE and US is named Mastigobryum chimborazense without forma and contains a later annotation ‘Probably Bazzania asperistipula. M. Fulford 1939’ but was not cited by Fulford (1946).

Currently accepted name: Bazzania hookeri (Lindenb.) Trevis. (following the synonymization of the species by Gradstein 2017).

Bazzania conchophylla Herzog, Rev. Bryol. Lichénol. 11(1): 20, 1939 (Herzog 1938)

Original citation: Costa Rica, Sta Maria de Dota, Prov. S. José, on tree, 1500–1800 m. ([Standley] n. 41.629). El Muñeco on the Rio Navarro, Prov. Cartago, 1400–1500 m. ([Standley] n. 50.884).

Lectotype (here designated): Costa Rica, Vicinity of Santa Maria de Dota, Provincia de San José, altitude 1500–1800 m, 14–26 Dec. 1925, Standley 41629 (JE04001113 https://je.jacq.org/JE04001113; synlectotypes Standley & Torres 50884 JE04001114 https://je.jacq.org/JE04001114, US04675710 http://n2t.net/ark:/65665/3ad7b7e89-9740-4cd0-b3aa-755feef75a65).

Note. Both specimens in JE cited in the protologue are named ‘Mastigobryum conchophyllum n. sp.’ in Herzog’s handwriting. The selected lectotype is richer.

Currently accepted name: Bazzania hookeri (Lindenb.) Trevis. var. hookeri (cf. Fulford 1963 as synonym of Bazzania teretiuscula (Lindenb. et Gottsche) Trevis. and Gradstein 2017).

Bazzania jamaicensis (Lehm. et Lindenb.) Trevis. var. chamaecardia Herzog, Rev. Bryol. Lichénol. 11(1/2): 19, 1938 (Herzog 1938)

Original type citation: Costa Rica, Cerros de Zurqui, Prov. de Heredia, 2000–2400 m. ([Standley] n 50.713. Cerro de Las Lajas, Prov. de Heredia, 2000–2400 m. ([Standley] n. 51.643/a, 51.645); Cerro de Las Caricias, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 52.117, 52.267).

Lectotype(Fulford1963:144):CostaRica,CerrodelasLajas,25 Mar 1926, Standley & Valerio 52267 (US04666041, http://n2t.net/ark:/65665/346b8e9d7-2f93-48b8-998b-6b62 9f3e78b5; isolectotype JE04006235 https://je.jacq.org/JE04006235 ;synlectotypes Standley 50713 JE04006236 https://je.jacq.org/JE04006236, Standley 51643a JE 04006237 https://je.jacq.org/JE04006237, Standley 52117 JE04006238 https://je.jacq.org/JE04006238).

Currently accepted name: Bazzania jamaicensis (Lehm. et Lindenb.) Trevis. (cf. Fulford 1963).

Bazzania portoricensis (Hampe et Gottsche) Steph. var. pycnodictyon Herzog, Rev. Bryol. Lichénol. 11(1/2): 19, 1938 (Herzog 1938)

Original type citation: Costa Rica, El Muñeco, on the Rio Navarro, Prov. Cartago, 1400–1500 m., on tree ([Standley] n. 51.338, 50.887); Alto de La Estrella, Prov. Cartago, on tree ([Standley] n. 39.106, 39.117); La Estrella, Prov. Cartago ([Standley] n. 39.403).

Lectotype (Fulford 1963: 133): El Muñeco, Standley 51338, 50887,US.Second-steplectotypification(Söderströmetal. 2023b: 150): Standley 50887 (US04665905 http://n2t.net/ark:/65665/395ca296e-afa0-40b3-89df-ad8acc4127e6; isolectotype JE04005948, https://je.jacq.org/JE04005948; synlectotype Standley 51338 JE04005949 https://je.jacq.org/JE04005949, US04665907 http://n2t.net/ark:/65665/m3951d11f7-b09f-4560-b6fd-a8b88ac1720b, Standley 39117 JE04005946 https://je.jacq.org/JE04005946, Standley 39106 JE04005947 https://je.jacq.org/JE04005947, Standley 39403 JE04005945 https://je.jacq.org/JE04005945).

Currently accepted name: Bazzania hookeri (Lindenb.) Trevis. (cf. Fulford 1963).

***Brachiolejeunea mamillata* Steph. f. brevifolia Herzog, Rev. Bryol. Lichénol. 20(1/2): 135, 1951 (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. [...] 54.372 und 55.428 (fo. brevifolia).

Original material: Honduras, Lancetilla Valley, near Tela, 20–600 m., Standley 54372 (JE04010821 https://je.jacq.org/JE04010821) and Standley 55428 (JE04010820 https://je.jacq.org/JE04010820).

Currently accepted name: Frullanoides mexicana van Slageren (van Slageren 1985).

Brachiolejeunea mamillata Steph. f. *obtusifolia*** Herzog, Rev. Bryol. Lichénol. 20(1/2): 135, 1951 (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. [...], 54.526.

Original material: Honduras, Lancetilla Valley, near Tela, 20–600 m., Standley 54526 (JE04010822 https://je.jacq.org/JE04010822).

Currently accepted name: Frullanoides mexicana van Slageren (van Slageren 1985).

Brachiolejeunea mamillata Steph. f. *typica*** Herzog, Rev. Bryol. Lichénol. 20(1/2): 135, 1951 (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. 55.249.

Original material: Honduras, Lancetilla Valley, near Tela, 20–600 m., Standley 55249 (JE04010819 https://je.jacq.org/JE04010819).

Currently accepted name: Frullanoides mexicana van Slageren (van Slageren 1985).

Brachiolejeunea parva Herzog, Rev. Bryol. Lichénol. 20: 125. 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Tela, Dep. of Atlántida, 20–600 m., [Standley] n. 56.075.

Lectotype (van Slageren 1985: 111 as ‘holotype’) if not holotype: Honduras, Lancetilla valley, near Tela, Dep. of Atlántida, 20–600 m, Standley 56075a (JE 04002172 https://je.jacq.org/JE04002172).

Currently accepted name: Frullanoides chinantlana (Gottsche) L. Söderstr. et A. Hagborg (cf. van Slageren 1985 as Frullanoides tristis (Steph.) van Slageren, Söderström and Hagborg 2023a).

Bryopteris tenuicaulis Taylor ex Gottsche, Lindenb. et Nees f. flagellifera Herzog, Rev. Bryol. Lichénol. 20(1/2): 132, 1951 (Herzog 1951)

Original type citation: Costa Rica, Prov. Limon, Guapiles, 300–500 m., [Standley] n. 37.049 und 37.317; Prov. Cartago, El Muneco, s. of Navarro, 1400 m., [Standley] 33.934 und 51.049; Prov. Limon, Hamburg Finca, rio Reventazon, below Cairo, 55 m., [Standley] n. 48.714; Prov. Guanacaste, El Arenal, on tree, 485–600 m., [Standley] n. 45.075 und 45.252; Prov. Guanacaste, Los Ayotes near Tilarán, on tree, 600–700 m., [Standley] n. 45.408 — Honduras: Lancetilla valley, near Tela, 20–600 m., [Standley] n. 53.219.

Lectotype (Stotler and Crandall-Stotler 1974: 142): Honduras, Dept. Atlantída: Lancetilla Valley, near Tela, 20–600 m, 1927–1927, Standley 53219 (F-C0000049F, https://fm-digital-assets.fieldmuseum.org/288/632/C0000049F.jpg, isolectotype JE04005919 https://je.jacq.org/JE04005919; synlectotypes Standley 33934 JE04005922 https://je.jacq.org/JE04005922, Standley 37049 JE04005923 https://je.jacq.org/JE04005923, Standley 37317 JE04005921 https://je.jacq.org/JE04005921, Standley 45075 JE04005925 https://je.jacq.org/JE04005925, Standley 45252 JE04005927 https://je.jacq.org/JE04005927, Standley 45408 JE04005924 https://je.jacq.org/JE04005924, Standley 48714 JE04005920 https://je.jacq.org/JE04005920, Standley 51049 JE04005926 https://je.jacq.org/JE04005926).

Currently accepted name: Bryopteris filicina (Sw.) Nees (cf. Stotler and Crandall-Stotler 1974).

Cephalozia subforficata Herzog, Rev. Bryol. Lichénol. 11: 17, 1938 (Herzog 1938)

Original citation: Costa Rica, Laguna de la Chonta, n. e. of Santa Maria de Dota, Prov. S. José, 2000–2100 m., on log ([Standley] n. 42.299).

Lectotype (Váňa 1988: 196 as ‘holotype’): Costa Rica, Laguna de la Chonta, northeast of Santa Maria de Dota, Provincia de San José, 2000–2100 m, 18.XII.1925, Standley 42299 (JE04001123 https://je.jacq.org/JE04001123; isolectotypes US00289479 http://n2t.net/ark:/65665/378eaac82-cd5e-4375-9f81-ba6fdec193a2).

Currently accepted name: Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L. Söderstr. (cf. Váňa 1988).

Ceratolejeunea deficiens Herzog, Rev. Bryol. Lichénol. 20: 148, 1951 (Herzog 1951)

Original citation: Costa-rica, Prov. Guanacaste, el Silencio, near Tilarán, 400–700 m., [Standley] n. 44.309 und 44.392p.; Prov. Heredia, Yerba Buena, n. e. of San Isidro, 2000 m., [Standley] n. 50.010 und 50.049. — Honduras: Lancetilla Valley, near Tela, Dep of Atlántida, 20–600 m., [Standley] n. 55.622p (parcissime).

Lectotype (Dauphin 2003: 63): Costa Rica, Guanacaste, vicinity of Tilarán, 500–650 m, 10–31 Jan. 1926, Standley & Valerio 44392p. (JE04001122 https://je.jacq.org/JE04001122, isolectotype US00070803 http://n2t.net/ark:/65665/321f74d78-ae3f-4927-8e55-2bb9cfe66bc6; synlectotypes Standley 44309 JE04005688 https://je.jacq.org/JE04005688, Standley 50010 JE04005661 h ttps://je.jacq.org/JE04005661, JE04005662 https://je.jacq.org/JE04005662, JE04005664 https://je.jacq.org/JE04005664,Standley5049JE04005663 https://je.jacq.org/JE04005663, JE04005665 https://je.jacq.org/JE04005665, JE04005666 https://je.jacq.org/JE04005666).

Currently accepted name: Ceratolejeunea laetefusca (Austin) R. M. Schust. (cf. Dauphin 2003).

Ceratolejeunea fusilobula Herzog, Rev. Bryol. Lichénol. 20: 149, 1951 (Herzog 1951)

Original citation: Costa Rica, Prov. Alajuela, Vienta Fresco, 1600–1900 m., [Standley] n. 47.806p.

Lectotype (here designated, cf. Dauphin 2003: 58 as ‘holotype’) if not holotype: Costa Rica, Viento Fresco, 1600–1900 m, 1908, Standley 47806, ‘Typus! Unicum’ (JE04001125 https://je.jacq.org/JE04001125).

Note. We have only been able to trace one possible type specimen, but it is possible that there exists more as Standley placed duplicates in several herbaria. Dauphin (2003: 58) named this specimen ‘holotype’ but as it was published after 1 January 2001 it cannot be accepted as an inadvertent lectotypification. The specimen in JE is marked as ‘unicum’ by Herzog. It may thus be a holotype.

Currently accepted name: Ceratolejeunea fallax (Lehm. et Lindenb.) Bonner (cf. Dauphin 2003).

Ceratolejeunea multiocellata Herzog, Rev. Bryol. Lichénol. 20: 150, 1951 (Herzog 1951)

Original citation: Costa-Rica, Prov. Cartago, vicinity of Pejivalle, ca 900 m., [Standley] n. 46.809.

Lectotype (here designated; c.f. Dauphin 2003: 59 as ‘holotype’): Costa Rica, Cartago: Pejivalle, 900 m, Standley 46809 (JE04002356 https://je.jacq.org/JE04002356; isolectotypes JE04007798 https://je.jacq.org/JE04007798, US00070808 http://n2t.net/ark:/65665/314bfd475-b8c1-4edc-8d0e-ea01f687a74c).

Note. Dauphin (2003) regarded this specimen as holotype, but as there are duplicates in several herbaria, a lectotype must be selected. Dauphin's designation as ‘holotype’ can not be accepted as an inadveretent lectotype as it was published after 1 January 2001 (c.f. ICN2018 Art. 7.11). We here formally designate the specimen as a lectotype.

Currently accepted name: Ceratolejeunea filaria (Taylor) Steph. (cf. Dauphin 2003).

Ceratolejeunea spinosa (Gottsche) Steph. var. costaricensis Zwickel, Beih. Bot. Centralbl. 49: 610, 1932 (Zwickel 1932).

Original citation: Costa Rica, Standley, n. 36.412, 37.691, 47.806. Herb. Herzog, Jena. Dominica: leg. Elliot, n. ?. Staatsherbar München, n. 7 als C. ceratantha. Costa Rica: leg. Tonduz. n.?. Staatsherbar München, n. 26 als C. involvens. Para: leg. Caripi, n. ?. Staatsherbar München, n. 29 als C. maritima.

Lectotype: (here designated): Costa Rica, La Hondura, Province of San José, altitude 1300–1700 m, 2–4 March 1924, Standley 36412 (JE04010817 https://je.jacq.org/04010817; isolectotype US00070810 http://n2t.net/ark:/65665/3a2ee40c9-7fb7-4922-8e0d-63bf13d92362).

Note. All syntypes in JE, including the lectotype, were reported as Ceratolejeunea filaria by Dauphin (2003). Syntypes from Costa Rica (leg. Tonduz), Dominica, and Pará are in M and their identities are unknown to us.

Currently accepted name: Ceratolejeunea filaria (Taylor) Steph. (cf. Dauphin 2003).

Cheilolejeunea pililoba (Spruce) A. Evans f. varians Herzog, Rev. Bryol. Lichénol. 20(1/2): 167, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. 55.577.

Lectotype (here designated) or possibly holotype: Honduras, Atlántida, Lancetilla Valley, near Tela, altitude 20–600 m, 6 Dec. 1927–20 Mar. 1928, Standley 55577 (JE04010835 https://je.jacq.org/JE04010835).

Currently accepted name: Lejeunea trinitensis Lindenb. var. varians (Herzog) B.Cañiza Cañiza et al. (2023).

Cheilolejeunea schaefer-verwimpii Gradst. et C. J. Bastos, Nova Hedwigia 113(1/2): 82, 2021 (Gradstein and Bastos 2021)

Holotype: Costa Rica, Cartago: Cordillera de Talamanca, Panamericano km 67, Reserva 3 de Junio, ‘in Quellmoor epiphytisch an Sträuchern,’ 2460 m, 3 Jan. 2000, A. Schäfer-Verwimp & I. Holz 0452/B as Cheilolejeunea choachina (Gottsche) Gradst. (JE s.n.; isotypes: GOET, CR).

Chiloscyphus nigrescens Lindenb. et Hampe, Linnaea 24(3): 640, 1851 (Hampe 1851)

Original citation: Hab. in Costa rica ad crateris Reventado aquas stagnates.

Lectotype (Grolle 1960: 84): Costarica, Reventado, 10.000' alt., im Kratersee Reventado, oja de Pascado, leg. Oersted ex hb. Lehmann in hb. S. Second-step lectotypification _(Söderström et al. 2023c: 151): S-B22184 ( https://herbarium.nrm.se/specimens/B22184; isolectotypes S, JE04002254 https://je.jacq.org/JE04002254).

Note. There exist three specimens in S fitting Grolle's lectotypification. The selected specimen is the richest of the three and also annotated as ‘type’ by J. J. Engel in 1978.

Currently accepted name: Clasmatocolea vermicularis (Lehm.) Grolle (cf. Grolle 1960).

*Chiloscyphus obliquefolius*** Herzog ex Fulford, Mem. New York Bot. Gard. 11: 448, 1976 (Fulford 1976), nom. inval. (ICN Art. 36.1(c); publ. in syn. [sub Lophocolea guadalupensis Steph. 1907])

Originalcitation:CostaRica,Cartago:CerrodelaCarpintera, 1500–1800 m, Standley 35724, type of Ch. obliquefolius (JE).

Original material: Costa Rica, Cartago: Cerro de la Carpintera, 1500–1800 m, Standley 35724 (JE04010814 https://je.jacq.org/JE04010814).

Currently accepted name: Cryptolophocolea guadalupensis (Steph.) L. Söderstr. et Váňa (cf. Fulford 1976).

Chiloscyphus pittieri Steph., Prim. Fl. Cost.: 112. 1893 (Stephani 1892)

Original citation: Bois humides entre la lagune du Barba et le Carrizal, 2800 m. (Pitt. 6014).

Lectotype (Grolle 1962: 57 as ‘holotypus’): Costa Rica. Entre la Lagune de Barba et le Carrisel, 2800 m., I 1889, Pittier 6014 (G00069439 www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=115868&lang=en; isolectotypes JE04002039 https://je.jacq.org/JE04002039, JE04002038 https://je.jacq.org/JE04002038).

Currently accepted name: Leptoscyphus amphibolius (Nees) Grolle (cf. Grolle 1962).

Cololejeunea ensifolia (Spruce) Steph. f. *angustifolia*** Herzog, Rev. Bryol. Lichenol. 20(1/2): 174, (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [...], [Standley] n. 54.846p.

Original material: Honduras, Lancetilla Valley, near Tela, 20–600 m., Dec. 1927–Mar. 1928, Standley 54846 (JE04010815 https://je.jacq.org/JE04010815).

Currently accepted name: Cololejeunea diaphana A. Evans (following var. Cololejeunea ensifolia var. pygmaea Herzog under which it was described, cf. Dauphin et al. 2008).

Cololejeunea ensifolia (Spruce) Steph. f. *genuina*** Herzog, Rev. Bryol. Lichenol. 20(1/2): 174, 1951 (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. 55 577.

Original material: Honduras, Lancetilla Valley, near Tela, 20–600 m., Standley 55577 (JE04010816 https://je.jacq.org/JE04010816).

Currently accepted name: Cololejeunea diaphana A. Evans (following var. Cololejeunea ensifolia var. pygmaea Herzog under which it was described, cf. Dauphin et al. 2008).

Cololejeunea minutilobula Herzog, Rev. Bryol. Lichénol. 20(1/2): 171, 1952 (Herzog 1951)

Original citation: Costa Rica, ohne Fundortsnotiz, [Standley] n. 26.176.

Lectotype (here designated): Panama, Río Tapia, 7 Dec. 1923–11 Jan. 1924, Standley 26176 (JE04004064 https://je.jacq.org/JE04004064, isolectotypes (JE04000973 https://je.jacq.org/JE04000973 (microslide), US00070983 http://n2t.net/ark:/65665/34d6b9a3a-a59d-455b-af8e-4a9e611388d8).

Note. Herzog mistakenly placed the locality in Costa Rica (‘ohne Fundortsnotiz’) although both the larger specimen in JE and the specimen in US clearly state they are from Panama. Although it is highly probable that Herzog did see the specimen in US, the specimen in JE comes from his personal herbarium. Thus, we select this as the lectotype.

Cololejeunea standleyi Herzog, Rev. Bryol. Lichénol. 20(1/2): 172, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., [Standley] n. 54.827.

Lectotype (here designated): Honduras, Atlántida, Lancetilla Valley near Tela, 20–600 m, 6 Dec. 1927, Standley 54827 (JE04003945 www.jacq.org/JE04003945, isolectotype US00070986, http://n2t.net/ark:/65665/324192e64-4c6a-48a3-8438-70b81447882b).

Note. Although, it is highly probable that Herzog did see the specimen in US, the specimen in JE comes from his personal herbarium. Thus, we select the larger specimen as the lectotype. The microslide was most probably extracted from it.

Cololejeunea tamasii Schäf.-Verw., Phytotaxa 60: 9, 2012 (Schäfer-Verwimp 2012)

Holotype: Panama, Chiriquí: Cordillera de Talamanca, Boquete, Ostseite des Vulkan Baru, Regenwald entlang der Erdstraße zum Sendero Los Quetzales, epiphyll, 1880 m, 8°51.0′N, 82°29.9′W, 7 Apr. 2010, A. Schäfer-Verwimp & I. Verwimp 30 992 (JE s.n.; isotype EGR).

Crossotolejeunea lindeniana Gottsche ex Steph. var. connivens Herzog, Rev. Bryol. Lichénol. 20: 141, 1951 (Herzog 1951)

Original citation: Costa-Rica, on tree, Cerro de las Caricias, n. of San Isidro, Prov. Heredia, 2000–2400 m., leg. P. C. Standley, n. 52.294.

Lectotype (Reiner-Drehwald and Goda 2000: 15 as ‘holotype’): Costa Rica, Heredia, C. de Las Caricias, N of San Isidro, Standley & Valerio 52294 (JE04002113 https://je.jacq.org/JE04002113; isolectotypes JE04002112 https://je.jacq.org/JE04002112, US00070849 http://n2t.net/ark:/65665/329a7ae97-31a4-4cf1-916e-800406f8cf12).

Note. There are two specimens in JE corresponding to this collection, but only JE04002113 is annotated as ‘holotype’ by Reiner-Drehwald. If a formal second lectotypification is required, this specimen is here designated as lectotype.

Currently accepted name: Lejeunea controversa Gottsche (cf. Reiner-Drehwald and Goda 2000).

Crossotolejeunea obtusiuscula Herzog, Rev. Bryol. Lichénol. 20: 142, 1951 (Herzog 1951)

Original citation: Costa Rica, on tree, vicinity of Pejivalle, Prov. Cartago, ca 900 m., leg. P. C. Standley, n. 47.128/b.

Lectotype (Reiner-Drehwald and Goda 2000: 15 as ‘holotype’) or possibly holotype: Costa Rica. Cartago: vicinity of Pejivalle, Standley 47128b (JE04002106 https://je.jacq.org/JE04002106).

Currently accepted name: Lejeunea controversa Gottsche (cf. Reiner-Drehwald and Goda 2000).

Cyclolejeunea schaefer-verwimpii M. L. M. Prado et Ilk.-Borg., Nova Hedwigia 113(1/2): 130, 2021 (Prado and Ilkiu-Borges 2021)

Holotype: Panama, Chiriquí, Reserva Forestal Fortuna, 8°44.3′N, 82°14.7′W, ‘epiphytisch, feuchtes Sekundärgebüsch am Ufer des Stausees’, 1100 m, 31 Mar. 2013, A. Schäfer-Verwimp & Verwimp 34520 (JE s.n.; isotype: MG).

Dicranolejeunea pulchella Herzog, Rev. Bryol. Lichénol. 20, 1951 (Herzog 1951)

Original type citation: Costa-Rica, on exposed rock, vicinity of Santa Maria de Dota, Prov. San José, 1500–1800 m., leg. P. C. Standley, n. 43 .153.

Lectotype (Gradstein 1994: 191 as ‘holotype’): Costa Rica, San José, Vicinity of Santa Maria de Dota. On exposed rock, 1500–1800 m., 26 Dec. 1925–3 Jan. 1926, Standley & Valerio 43153 (JE00401075 https://je.jacq.org/JE04001075; isolectotype US00070854 http://n2t.net/ark:/65665/3813e1b57-2f0d-445a-aaa8-ea1e8199a2ac).

Currently accepted name: Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt var. aberrans (cf. Gradstein 1994).

Diplasiolejeunea involuta S. Winkl., Rev. Bryol. Lichénol. 35(1/4): 320, 1967 (Winkler 1967)

Original type citation: SE-hang des Cerro Miramundo bei der Hazienda Montecristo, Metapan, Dep. Santa Ana, 2350 m A/17/3 (Typus) 23. 5. 62 leg. S. Winkler.

Isotype: JE04008786 ( https://je.jacq.org/JE04008786) ‘herb. R. Grolle ex TUB, Winkler 57a/8’. Collection number is not the same as in protologue but all other data seem to fit.

Note. A lectotype from TUB should be selected.

Diplasiolejeunea pluridentata Schäf.-Verw., Haussknechtia 8: 71, 2001 (Schäfer-Verwimp 2001)

Holotype:CostaRica,Prov.deSanJose,CordilleradeTalamanca, San Gerardo de Dota, Quercus-copeyensis-Primärwald ‘Los Rabies’, an freistehendem Strauch, 2400 m, 4 Jan. 2000, Schäfer-Verwimp & Holz Nr. SV/H-0486/B (JE04002281 https://je.jacq.org/JE04002281; isotypes in EGR, CR).

Drepanolejeunea bidens (Prantl) A. Evans f. *denticulata*** Herzog, Rev. Bryol. Lichénol. 20: 145, 1951 (Herzog 1951), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa-Rica, Prov. Heredia, Yerba Buena, on leaves, [Standley] n. 49.709p.

Original material (cf. also Bischler 1964 as ‘type’): Costa Rica, Heredia, Yerba Buena, on leaves, Standley 49 709 (JE04010818 https://je.jacq.org/JE04010818).

Currently accepted name: Drepanolejeunea infundibulata (Spruce) A. Evans (cf. Bischler 1964).

Drepanolejeunea integerrima Herzog, Rev. Bryol. Lichénol. 20(1/2): 146, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Tela, Dep. of Atlántida, 20–600 m., epiphyll mit Cyclolejeunea convexistipa, ausserst spärlich, [Standley] n. 56.755p.

Possible lectotype: Honduras, Lancetilla Valley, near Tela, Dep. of Atlántida, 20–600 m., Standley 56755 (US04659783 http://n2t.net/ark:/65665/38261d299-4865-46b1-afaf-488afb77377e).

Note. Bischler (1964) stated that the type is lost and that it is a Leptolejeunea. We have not been able to locate any type material in JE. US04659783 is filed under Drepanolejeunea mosenii after an annotation by H. Bischler.

Frullania armata Herzog et L. Clark, Bryologist 57(1): 36, 1954 (Clark 1954)

Original citation: Costa Rica, Provincia de San Jose, vicinity of Santa Maria de Dota, altitude 1500–1800 m (Paul Standley 42479) 14–26 Dec. 1925 (type).

Lectotype:?

Note. Clark (1954) said it was borrowed from US and is an isotype. This is illustrated in the original publication. UW holding Clark's herbarium is not databased. A microslide (JE04010811 https://je.jacq.org/JE04010811) was probably extracted by Herzog, but lectotypification should wait until it is known if more material exists in US and/or UW.

Frullania convoluta Lindenb. et Hampe var. ampliata Herzog, Rev. Bryol. Lichénol. 20(1/2): 129, 1952 (Herzog 1951)

Original citation: Costa-rica, Prov. San José, La Palma, 1600 m, [Standley] n. 33.014 und 33.039; Prov. San José, Zurqui 2000–2500 m, on tree, [Standley] n. 48.170; Prov. Heredia, Yerba Buena, on tree, 2000 m., [Standley] n. 49.844; Prov. Heredia, Cerros de Zurqui, 2000–2400 m., [Standley] n. 50.457; Prov. Heredia, Cerro de las Lajas, 2000–2400 m., [Standley] n. 51.458; Prov. Heredia, Cerro de las Caricias, 2000–2400 m., [Standley] n. 51.986.

Lectotype (Söderström et al. 2023a: 2): Costa Rica, La Palma, Province de San José, altitude about 1600 m, 3 Feb. 1924, Standley 33014 (JE04002308 https://je.jacq.org/JE04002308, synlectotypes Standley 33039 JE04010810 https://je.jacq.org/JE04010810, Standley 48170 JE04002307 https://je.jacq.org/JE04002307, Standley 49844 JE04002306 https://je.jacq.org/JE04002306, Standley 50457 JE04002305 https://je.jacq.org/JE04002305, Standley 51986 JE04002303 https://je.jacq.org/JE04002303, Standley 51458 JE04002304 https://je.jacq.org/JE04002304, Standley 33039 JE04010810 https://je.jacq.org/JE04010810).

Currently accepted name: Frullania convoluta Lindenb. et Hampe (cf. Söderström et al. 2023a).

Frullania hamiflora Herzog et L. Clark, Bryologist 56(3): 180, 1953 (Clark and Schultz 1953a)

Holotype: Costa Rica, Provincia de Alajuela, vicinity of Fraijanes, altitude 1500–1700 m, 12–13 Feb. 1926, Standley & Torres 47636, (US00070889 http://n2t.net/ark:/65665/335b4c841-0671-4c15-89b8-b88c8abac97d; isotypes G, JE04000986 https://je.jacq.org/JE04000986).

Frullania hondurae Herzog et L. Clark, Bryologist 56: 289, 1953 (Clark and Svihla 1953)

Original citation: Costa Rica, Province of San José, vicinity of La Hondura, altitude 1300–1700 m, 2–4 Mar. 1924 (Standley 36559).

Lectotype (Stotler 1969 as ‘type’): Costa Rica, San José: La Hondura, 1300–1700 m, Standley 36559 (US00070891 http://n2t.net/ark:/65665/3e774f1d9-cd2a-4f3b-b414 -7d170b75d4b2; isolectotype JE04002301 https://je.jacq.org/JE04002301).

Currently accepted name: Frullania brasiliensis Raddi (cf. Stotler 1969).

Frullania macrorhyncha Herzog et L. Clark, Bryologist 56(4): 283, 1953 (Clark and Schultz 1953b)

Holotype: Costa Rica, Provincia San José (Zurqui, altitude 2000–2500 m, 13 Feb. 1926, Standley & Valerio 48135 (US00070894 http://n2t.net/ark:/65665/30b865f22-a4b3-400b-96c2-99d75683c5e8, isotype JE04001005 https://je.jacq.org/JE04001005).

Currently accepted name: Frullania hamiflora Herzog et L. Clark (cf. Stotler 1969).

Frullania piliflora Steph. var. appendiculata Herzog, Rev. Bryol. Lichénol. 20(1/2): 129. 1951 (Herzog 1951)

Original citation: Costa Rica, south slope of volcan de Turrialba, near finca of V.d.T., 2000–2400 m., [Standley] n. 35.088; Prov. San José, Las Nubes, wet bank, 1500–1900 m., [Standley] n. 38.421.

Lectotype (Söderström et al. 2023a: 2): Costa Rica, Cartago, slope of Volcan de Turrialba, near Finca of V.d.T., 2000–2400 m., 2 Feb. 1924, Standley 35088 (JE04004146 https://je.jacq.org/JE04004146; isolectotypes G00265605 www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=271882&lang=en, US04646508 http://n2t.net/ark:/65665/30c6ff639-2fe7-4383-b919-296397863453).

Currently accepted name: Frullania peruviana Gottsche (cf. Söderström et al. 2023a).

Harpalejeunea exocellata Herzog, Rev. Bryol.

Lichénol. 20(1/2): 144, 1952 (Herzog 1951)

Original citation: Costa Rica, Prov. Heredia, Cerro de las Caricias, 2000–2400 m., leg. P. C. Standley, n. 51.988p., n. 47.157.

Lectotype (Grolle and Reiner-Drehwald 1999: 37): Costa Rica, Heredia, Cerro de las Caricias, 2000–2400 m., Standley 51998 (JE). Second-step lectotypification (here designated): JE04002123 ( https://je.jacq.org/JE04002123; isolectotype JE04002124 https://je.jacq.org/JE04002124).

Note. There exist two specimens in JE and Grolle and Reiner-Drehwald (1999) did not specify which of them they selected. The isolectotype is a microslide almost certainly extracted from the lectotype but as stated above we can not be sure of this and we prefer to designate the richer material as lectotype.

Herbertus angustifolius (Steph.) Herzog f. *minor*** Herzog, Rev. Bryol. Lichénol. 11(1): 24. 1939 (Herzog 1938), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa Rica, Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 50.292, f. minor).

Original material: Costa Rica, Cerros de Zurquí, Prov. de Heredia, 2000–2400 m., on tree, Standley & Valerio 50292 (JE04010823 https://je.jacq.org/JE04010823).

Currently accepted name: Herbertus pensilis (Taylor) Trevis. (cf. Morales 1991).

Herbertus bivittatus Spruce var. latilobus Herzog,

Rev. Bryol. Lichénol. 11(1): 24, 1939 (Herzog 1938)

Original citation: Costa Rica, Vicinity of Pejivalle, Prov. Cartago, 900 m., on tree ([Standley] n. 47.021, 47.173).

Lectotype (Fulford 1963: 98 as ‘type’): Costa Rica, Cartago: near Pejivalle, Standley & Valerio 47021 (JE04010828 https://je.jacq.org/JE04010828; isolectotype US00072894 http://n2t.net/ark:/65665/36cd9e300-ab36-4b54-b30f-f9e3530bd2f1).

Currently accepted name: Herbertus bivittatus Spruce (cf. Fulford 1963).

Herbertus cancerinus Herzog, Rev. Bryol. Lichénol. 11(1/2): 25, 1938 (Herzog 1938)

Original citation: Costa Rica, Cerro de Las Lajas, Prov. de Heredia, 2000–2400 m. on tree ([Standley] n. 51.629).

Lectotype (Fulford 1963: 100 as ‘type’): Costa Rica, Cerro de Las Lajas, 2000–2400 m, Standley & Valerio 51629 (JE04010836 https://je.jacq.org/JE04010836; isolectotype US00072895 http://n2t.net/ark:/65665/3e397f894-9316-4d72-b1e1-3548c1ddac53).

Currently accepted name: Herbertus acanthelius Spruce (cf. Fulford 1963 as Herbertus serratus Spruce, and Feldberg and Heinrichs 2006).

Herbertus cancerinus Herzog f. *foliis-minus-serratis*** Herzog, Rev. Bryol. Lichénol. 11(1/2): 27, 1938 (Herzog 1938), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original type citation: Costa Rica, Yerba Buena, Prov. de Heredia, 2000 m. on tree ([Standley] n. 50.011, f. foliis minus serratis).

Original material: Costa Rica, Heredia, Yerba Buena, NE San Isidro, 22 Feb. 1926, Standley & Valerio 50011 (JE04010804 https://je.jacq.org/JE04010804, US00072896, http://n2t.net/ark:/65665/3c32b5dad-1dc9-40a1-8661-a5b08bd9be68).

* **Herbertus durandii* (Steph.) Herzog var. dissectus Herzog, Rev. Bryol. Lichénol. 11(1/2): 25, 1939 (Herzog 1938), nom. inval. (ICN2018 Art. 39.1; no Latin description)

Original citation: Costa Rica, Cerros de Zurqui, Prov. de Heredia, 2000–2400 m. ([Standley] n. 50.338).

Original material: (Fulford 1963: 104 as ‘lectotype’): Costa Rica, Heredia: Cerros de Zurqui, 2000–2400 m, Standley & Valerio 50338 [...] (Hb. Herzog, JE04010809 https://je.jacq.org/JE04010809). Original material is also in US04667650 ( http://n2t.net/ark:/65665/3ed9e4d50-7465-4b82-8d46-2c605294398c).

Currently accepted name: Herbertus sendtneri (Nees) Lindb. (cf. Feldberg and Heinrichs 2005).

Herbertus pectinatus Herzog, Rev. Bryol. Lichénol. 11(1/2): 27, 1938 (Herzog 1938)

Original citation: Costa Rica, Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 50.380, 50.642).

Lectotype (Fulford 1963: 100 as ‘type’): Costa Rica, Cerros de Zurquí, Standley & Valerio 50642 (JE04010834 https://je.jacq.org/JE04010834; isolectotype US00072897 ( http://n2t.net/ark:/65665/366cd81be-dd7d-4644-98a0-dc0ecb803a53; synlectotype Standley & Valerio 50380 JE04010806 https://je.jacq.org/JE04010806).

Currently accepted name: Herbertus acanthelius Spruce (cf. Feldberg and Heinrichs 2006).

Herbertus penicillatus Herzog, Rev. Bryol. Lichénol. 11(1): 27, 1938 [1939] (Herzog 1938)

Original citation: Costa Rica, La Palma, Prov. S. José, 1600 m. ([Standley] n. 38.080).

Lectotype (Fulford 1963: 98 as ‘type’): Costa Rica, San José, La Palma, Standley 38080 (JE04010854 https://je.jacq.org/JE04010854; isolectotypes B-04270, US00072898 http://n2t.net/ark:/65665/31e26bb06-3bb1-4a25-9f0a-80f944fd77ce).

Currently accepted name: Herbertus bivittatus Spruce (cf. Fulford 1963).

Herbertus stenoschizon Herzog, Rev. Bryol. Lichénol. 11(1/2): 27, 1938 (Herzog 1938)

Original citation: Costa Rica, Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 50.392, 50.529).

Lectotype (Fulford 1963: 100 as ‘type’): Costa Rica, Heredia: Cerros de Zurquí 2000–2400 m, Standley & Valerio 50392 (JE04010833 https://je.jacq.org/JE04010833; isolectotype US00072900 http://n2t.net/ark:/65665/396f4181d-de9a-47ab-a37c-8b5c8fa53106; synlectotype Standley & Valerio 50529 JE04010807 https://je.jacq.org/JE04010807).

Currently accepted name: Herbertus acanthelius Spruce (cf. Feldberg and Heinrichs 2006).

Herbertus vulcanicola Herzog, Rev. Bryol. Lichénol. 11(1/2): 25, 1938 (Herzog 1938)

Original citation: Costa Rica, southern slope of the Volcán de Turrialba, near the Finca, 2000–2400 m., on tree ([Standley] n. 35.297).

Lectotype (Fulford 1963: 102 as ‘type’): Costa Rica, slope Volcán Turrialba, 2000–2400 m., Standley 35297, Hb. Herzog. Second-step lectotypification (here designated) (JE04010853 https://je.jacq.org/JE04010853; isolectotypes JE04010808 h ttps://je.jacq.org/JE04010808, US00072901 http://n2t.net/ark:/65665/372d95699-cfe6-4040-b231-f01ae6f6a9b7).

Note. The isolectotype in JE is a microslide almost certainly extracted from the lectotype.

Currently accepted name: Herbertus acanthelius Spruce (cf. Feldberg and Heinrichs 2006).

Hygrolejeunea costaricensis Steph., Hedwigia 35(3): 100, 1896 (Stephani 1896)

Original citation: Hab. Costarica, Carrisal 2800 m (Pittier).

Lectotype (Bonner 1966: 646; also Schuster 1994: 220 as ‘type’): Costa Rica, Carrisal, 2800 m., I 1889, Pittier; Pittier & Durand, Plantae Costaricenses Exsiccatae No 6013a (G00282422 www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=214857&lang=en; isolectotype JE04007655 https://je.jacq.org/JE04007655).

Currently accepted name: Lejeunea reflexistipula (Lehm. et Lindenb.) Lehm. et Lindenb. var. costaricensis (Steph.) M.E.Reiner (cf. Reiner-Drehwald 2005).

Hygrolejeunea punctata Herzog, Rev. Bryol. Lichénol. 20(1/2): 163. 1951 (Herzog 1951)

Original citation: Costa-Rica, on tree, Cerros de Zurqui, Prov. Heredia, 2000–2400 m., leg. P. C. Standley, n. 50.640/a (zwischen Hygrolejeunea rotundifolia).

Lectotype (Heinrichs et al. 2015: 224): Costa Rica, Heredia, Cerros de Zurqui, 2000–2400 m, 3 Mar. 1926, Standley 50640a (JE04002098 https://je.jacq.org/JE04002098 ‘handwritten label’; isolectotypes JE04002096 https://je.jacq.org/JE04002096, JE04002097 https://je.jacq/JE04002097).

Currently accepted name: Lepidolejeunea cuspidata (Gottsche) Heinrichs et Schäf.-Verw. (cf. Heinrichs et al. 2015).

Isotachis ripensis Spruce var. armata Herzog, Rev. Bryol. Lichénol. 11: 38, 1939 (Herzog 1938)

Original citation: Costa Rica, southern slope of Volcán de Turrialba, near the Finca, 2000–2400 m., on tree ([Standley] 35.279).

Lectotype (Grolle 1964: 240): Costa Rica, Volcán de Turrialba, 2000–2400 m, on tree, 1924, Standley 35279 (JE04004922 https://je.jacq.org/JE04004922, isolectotype US00289478 http://n2t.net/ark:/65665/3ed61fec0-b1ee-4e11-9dc2-448452c121a4).

Currently accepted taxon: Triandrophyllum subtrifidum (Hook.f. et Taylor) Fulford et Hatcher var. trifidum (Gottsche) Solari (cf. Grolle 1964).

Jubula hutchinsiae (Hook.) Dumort. var. *costaricensis*** Herzog, Rev. Bryol. Lichénol. 20: 128, 1951 (Herzog 1951), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa Rica, Prov. Cartago, Cerro de la Carpintera, 1500 m., [Standley] n. 35.674p.; Prov. Cartago, Vicinity of Pejivalle, 900 m., wet bank, [Standley] n. 47.017; Prov. Heredia, Yerba Buena, 2000 m., on wet rocks, [Standley] n. 49.678.

Originalmaterial(Guerke1978:92as‘holotype’):CostaRica, Cerro de La Carpintera, Standley 35674 (JE04010824 https://je.jacq.org/JE04010824, ‘isotype’ US04544769 http://n2t.net/ark:/65665/305f56692-050d-4ed4-a7d0-0502815b0baf).

Currently accepted name: Jubula hutchinsiae subsp. bogotensis (Steph.) Verd. (cf. Guerke 1978).

Lejeunea succulenta Herzog, Rev. Bryol. Lichénol. 20(1/2): 171, 1951 (Herzog 1951)

Original citation: Costa Rica, Prov. Guanacaste, vicinity of Tilarán, 500–600 m., [Standley] n. 46.647p. – Honduras: Lancetilla Valley, near Tela, Dep. of Atlántida, 20–600 m., [Standley] n. 56.369p.

Lectotype: (here designated): Costa-Rica, Prov. Guanacaste, vicinity of Tilarán, 500–600 m., Standley 46647p. (JE04010830 https://je.jacq.org/JE04010830, isolectotypes JE04009372 https://je.jacq.org/JE04009372, JE04009373 https://je.jacq.org/JE04009373, B300025227 https://herbarium.bgbm.org/object/B300025227, US 00070960 http://n2t.net/ark:/65665/34d0b12c7-8241-43b2-b288-10f63b277115.

Note. JE04009372 and JE04009373 are syntypes of Rectolejeunea defolians Herzog but annotated as also including Lejeunea succulenta ‘sehr spärlich’. The selected lectotype was most likely extracted from one of those packets.

Lejeunea talamancensis M. E. Reiner et Schäf.-Verw., Nova Hedwigia 87(3/4): 414, 2008 (Reiner-Drehwald and Schäfer-Verwimp 2008)

Holotype: Costa Rica, San José: Cordillera de Talamanca, San Gerardo de Dota, path from Chacón to ‘Las Cruces’ on river Savegre, epiphyte on Alnus sp., 1900 m, 25 Dec. 1999, Schäfer-Verwimp & Holz SV/H-0098/B (JE s.n.; isotypes GOET, CR).

Lepidozia karstenii Steph. f. *angustisecta*** Herzog, Rev. Bryol. Lichénol. 11(1/2): 22, 1939 (Herzog 1938), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa Rica, …; Zurqui, Prov. S. José, 2000–2500 m., on tree ([Standley] n. 48.173, 48.194/a, forma angustisecta); Cerro de Las Caricias, Prov de Heredia, 2000–2400 m., on tree ([Standley] n. 52.001, f. angustisecta; Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 50.512, 50.518, f. angustisecta); [...]; La Palma. Prov. S. José, 1600 m., wet bank, in large softe [sic!] cushions ([Standley] n. 32.968, f. angustisecta).

Original material: Standley 48173 (JE04002072 https://je.jacq.org/JE04002072), Standley 48194a (JE04002066 https://je.jacq.org/JE04002066), Standley 52001 (JE04002067 https://je.jacq.org/JE04002067), Standley 50512 (US04666846 http://n2t.net/ark:/65665/33f5090e7-f130-4285-9896-e5c90be44755), Standley 50518 (JE04002070 https://je.jacq.org/JE04002070), Standley 32968 (JE04002074 https://je.jacq.org/JE04002074, US04666849 http://n2t.net/ark:/65665/38b75a1e4-69ef-4944-bf2a-56db37824e97).

Note. All original material was cited under Lepidozia karstenii var. standleyi Herzog and are all syntypes of the variety although not considered ‘typical’ by Herzog.

Currently accepted name: Lepidozia armata Steph. (cf. Fulford 1966 following the synonymization of Lepidozia karstenii).

Lepidozia karstenii Steph. f. *brevifissa*** Herzog, Rev. Bryol. Lichénol. 11(1/2): 22, 1939 (Herzog 1938), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa Rica, [...]; Cerro de Las Lajas ([Standley] n. 51.540, f. brevifissa); Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree [Standley] n. 50.696; f. brevissima); Zurqui, Prov. S. José, 2000–2500 m. ([Standley] n. 48.214, f. brevissima).

Original material: Standley 50696 (JE04002069 https://je.jacq.org/JE04002069), Standley 48214 (JE04002071 https://je.jacq.org/JE04002071).

Note. All original material was cited under Lepidozia karstenii var. standleyi Herzog and are all syntypes of the variety although not considered ‘typical’ by Herzog.

Currently accepted name: Lepidozia armata Steph. (cf. Fulford 1966 following the synonymization of Lepidozia karstenii, but see Gradstein 2021).

Lepidozia karstenii var. standleyi Herzog, Rev. Bryol. Lichénol. 11(1/2): 22, 1939 (Herzog 1938)

Original type citation: Costa Rica, Las Nubes, Prov. S. José, 1500–1600 m., on tree ([Standley] n. 38.735, 38.466); Alto de La Estrella, Prov. Cartago, on tree ([Standley] n. 39.061); Zurqui, Prov. S. José, 2000–2500 m., on tree ([Standley] n. 48.173, 48.194/a, forma angustisecta); Cerro de Las Caricias, Prov de Heredia, 2000–2400 m., on tree ([Standley] n. 52.001, f. angustisecta; Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 50.512, 50.518, f. angustisecta); Cerro de Las Lajas, Prov. de Heredia, 2000–2400 m., on tree ([Standley] n. 51.432); La Palma. Prov. S. José, 1600 m., wet bank, in large softe [sic!] cushions ([Standley] n. 32.968, f. angustisecta); Cerro de Las Lajas ([Standley] n. 51.540, f. brevifissa); Cerros de Zurqui, Prov. de Heredia, 2000–2400 m., on tree [Standley] n. 50.696; f. brevifissa); Zurqui, Prov. S. José, 2000–2500 m. ([Standley] n. 48.214, f. brevifissa). – Cerro Galito, 2000 m. (Valerio n. 13); La Palma (Valerio n. 20).

Lectotype (Fulford 1966: 201 as ‘isotypes’): Costa Rica, San Jose: Las Nubes, 1500–1600 m, Standley 38735, Verdoorn, Hep. Select. & Crit. 375 (BR, C, F https://collections-botany.fieldmuseum.org/catalogue/2130884). Second-step lectotypification (Söderström et al. 2023b:152): BR5040110401156 ( www.botanicalcollections.be/specimen/BR5040110401156). Synlectotypes (excl. those cited as f. angustisecta or f. brevifissa): Valerio 20 JE04002073 https://je.jacq.org/JE04002073, Standley 38466 JE04002075 https://je.jacq.org/JE04002075, Standley 39061 JE04002076 https://je.jacq.org/JE04002076, Standley 51432 JE04002068 https://je.jacq.org/JE0400206).

Note. Original material cited under Lepidozia karstenii f. angustisecta and f. brevifissa above are technically synlectotypes also to var. standleyi although T. Herzog did not consider them ‘typical’.

Currently accepted name: Lepidozia armata Steph. (cf. Fulford 1966, but see Gradstein 2021).

Lepidozia macropatens Herzog, Rev. Bryol. Lichénol. 11(1/2): 20, 1939 (Herzog 1938)

Original citation: Costa Rica, Cerro de Las Vueltas, Prov. S. José, 2700–3000 m., on tree ([Standley] n. 43.723).

Lectotype (here designated): Costa Rica: Cerro de Las Vueltas, Prov. S. José, 2700–3000 m., on tree, Standley n. 43.723 (JE04002087 https://je.jacq.org/JE04002087; isolectotype US00289481 http://n2t.net/ark:/65665/3f0f8ebad-c8b4-4953-9a47-20a1f36bbbc9).

Note. Herzog did probably see the specimen in US but the description is most certainly based on the specimen in JE.

Currently accepted name: Lepidozia reptans (L.) Dumort. (cf. Fulford 1966).

Leptolejeunea stenophylla (Lindenb. et Gottsche) Schiffn. f. *latifolia*** Herzog, Rev. Bryol. Lichénol. 20: 146, 1951 (Herzog 1951), nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., on leaves, [Standley] n. [...] und 56.755.

Original material (cf. Bischler 1967): Honduras, Atlántida, Lancetilla Valley, near Tela, 20–600 m, Standley 56755 (JE, US).

Currently accepted name: Drepanolejeunea mosenii (Steph.) Bischl. (cf. Bischler 1967).

Note. No originally material is digitized in JE or US but should be there fide Bischler (1967).

Leptoscyphus lambinonii Vanderp., Schäf.-Verw. et D. G. Long, Taxon 59(1): 179, 2010 (Vanderpoorten et al. 2010)

Holotype: Costa Rica, Cartago: Orosi valley, Tapantí National Park, primary rain forest, Sendero La Oropendola, epiphytic, ca 600 m, 30 Dec. 1999, Schäfer-Verwimp & Holz SV/H-0342 (JE s.n.; isotype, CR).

Lophocolea aberrans Lindenb. et Gottsche var. costaricensis Herzog, Rev. Bryol. Lichénol. 11(1/2): 17, 1939 (Herzog 1938)

Original citation: Costa Rica, Vicinity os [sic!] Sta Maria de Dota, Prov. S. José, 1500–1800 m. ([Standley] n. 41.686, 41.758/a, 41.835); near Finca La cima, above Los Lotes, n. of El Copey, Prov. S. José, 2100–2400 m. ([Standley] . 42.711, 42.781. 42.800); Quebradillas, 1800 m. oak forest ([Standley] n. 42.933); El Muñeco, Rio Navarro, 1400–1800 m. ([Standley] 50.931); Cerro de Las Vueltas, 2700–3000 m., old log ([Standley] n. 43.758); Laguna de La Chonta, 2000–2100 m. ([Standley] n. 42.248).

Lectotype (here designated): Costa Rica, Cerro de las Vueltas, Provincia de San José, on old log, altitude 2700–3000 m, 29 Dec. 1925–1 Jan. 1926, Standley & Valerio 43758 (JE04002012 https://je.jacq.org/JE04002012; isolectotype JE04002011 https://je.jacq.org/JE04002011, US04628457 http://n2t.net/ark:/65665/331215e63-87b4-4a33-a9f3-8e3c14753ea1, synlectotypes Standley 41686 JE04002022 https://je.jacq.org/JE04002022, Standley 41758a JE04002021 https://je.jacq.org/JE04002021, Standley 41835 JE04002020 https://je.jacq.org/JE04002020, Standley 42248 JE04002019 https://je.jacq.org/JE04002019, Standley 42711 JE04002017 https://je.jacq.org/JE04002017, JE04002018 https://je.jacq.org/JE04002018, Standley 42781 JE04002016 https://je.jacq.org/JE04002016, Standley 42800 JE04002015 https://je.jacq.org/JE04002015, Standley 42933 JE04002014 https://je.jacq.org/JE04002014, Standley 50931 JE04002013 https://je.jacq.org/JE04002013).

Note. All syntypes in JE were studied and annotated by R. Grolle, and further also by M. Fulford. Material of all seems good and the selected lectotype seems appropriate.

Currently accepted name: Leptoscyphus amphibolius (Nees) Grolle (cf. Grolle 1962).

Lophocolea breutelii Gottsche var. *robusta*** Herzog ex Fulford, Mem. New York Bot. Gard. 11(4): 435. 1976 (Fulford 1976), nom. inval. (ICN Art. 36.1(c); publ. in syn.)

Original citation: Costa Rica, [...] Heredia: San Isidro, Standley & Valerio 51474 as L. breutelii var. robusta (JE).

Original material (Fulford 1976: 437): Costa Rica, Heredia, Cerro de Las Lajas, north of San Isidro Standley & Valerio 51474 (JE04010837 https://je.jacq.org/JE04010837; original material also in US04675709 http://n2t.net/ark:/65665/33a19c1a8-02ad-4d83-83b7-731f05dc6f02).

Currently accepted name: Leptoscyphus orbignianus (Nees et Mont.) L. Söderstr. et A. Hagborg (cf. Fulford 1976 and Söderström and Hagborg 2023b).

Lophocolea effusidens Steph., Bull. Herb. Boissier (sér. 2) 7(4): 301 [Sp. Hepat. (Stephani) 3: 165], 1907 (Stephani 1907)

Original type citation: Hab. Peruvia, St Gavan (Lechler) Costarica (Tonduz).

Lectotype (Fulford 1976: 431): Peru, St. Gavin, ex Hb Jack, Lechler (G00112460; www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=134928&lang=en; synlectotypes Tonduz JE04001875 https://je.jacq.org/JE04001875, JE04001876 https://je.jacq.org/JE04001876, M?).

Currently accepted name: Leptoscyphus orbignianus (Nees et Mont.) L. Söderstr. et A. Hagborg (cf. Gradstein 2021 and Söderström and Hagborg 2023b).

Note. The synlectotype from Costa Rica is Heteroscyphus marginatus (Steph.) Fulford (cf. Fulford 1976).

Lophocolea liebmanniana Gottsche var. *bisbifida*** Herzog ex Standl., Field Mus. Nat. Hist. Bot. Ser. 10: 62. 1931 (Standley 1931), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: [Honduras, Atlántida, Lancetilla valley].

Original material (cf. Fulford 1976: 427): Honduras, Lancetilla Valley, near Tela, on log, 20–600 m, 6 Dec. 1927–20 Mar. 1928, Paul C. Standley 54836, det Herzog Mar. 1929 (JE04010827 https://je.jacq.org/JE04010827).

Note. Annotated as no Lophocolea liebmanniana in packet by M. Fulford although she synonymized it with that species. If her annotation is correct, which we think it is, the synonymisation is wrong!.

Currently accepted name: Lophocolea liebmanniana Gottsche (cf. Fulford 1976).

Lophocolea tricuspidata Herzog, Rev. Bryol. Lichénol. 11(1): 17, 1939 (Herzog 1938)

Original citation: Costa Rica, Cerro de Las Vueltas, 2700–3000 m., on tree ([Standley] n. 43.723/c, mingled with Notoscyphus paramicola, Adelanthus parvus and Nowellia curvifolia).

Lectotype (here designated) if not holotype: Costa Rica, Cerro de Las Vueltas, 2700–3000 m, Standley 43723c (JE04001777 https://je.jacq.org/JE04001777).

Note. This specimen may have been taken from a richer specimen. Standley‘s collection number. 43723 is identified as Lepidozia macropatens in US00289481 and JE04002087. The type material was probably extracted from this gathering and, thus, an isotype probably exists among the US specimens. The type of Adelanthus parvus (JE04002323 https://je.jacq.org/JE04002323) is also marked with the same number. (43723c) and is poor and possibly extracted from the same specimen. Type material of Notoscyphus paramicola (JE04000934 https://je.jacq.org/JE04000934 and JE04000935 https://je.jacq.org/JE04000935) is also poor and numbered 43723a and may originate from the same gathering.

Macrolejeunea subsimplex (Mont. et Nees) Schiffn. f. *minor*** Herzog, Rev. Bryol. Lichénol. 20(1/2): 161. 1951 (Herzog 1951), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Costa-Rica, [...]; Prov. Guanacaste, Quebrada Serena, s. e., of Tilarán, 700 m., on tree, [Standley] n. 46.256.

Original material: Costa Rica, Quebrada Serena, southeast of Tilarán, Provincia de Guanacaste, altitude about 700 m, on tree, Standley & Valerio 46256 (JE04010825 https://je.jacq.org/JE04010825, US04658647 http://n2t.net/ark:/65665/33920f591-1c38-4054-9cf7-8ffce0d7e93a).

Currently accepted name: Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb. (cf. Grolle 1987).

Macrolejeunea subsimplex (Mont. et Nees) Schiffn. f. *parvistipula*** Herzog, Rev. Bryol. Lichénol. 20(1/2): 161. 1951 (Herzog 1951), nom. inval. (ICN2018 Art. 38.1(a); no description)

Original citation: Honduras, Dept of Atlántida, Lancetilla valley, near Tela, on Cyclopeltis, 20–600 m., [Standley] n. 56.865.

Original material: Honduras, Atlántida, Lancetilla Valley, Standley 56805 (JE04010826 https://je.jacq.org/JE04010826).

Note. The collection number in the ‘protologue’ and on the specimen differ but we assume it is an error in print.

Currently accepted name: Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb. (cf. Grolle 1987).

Madotheca maxonii Herzog, Beih. Bot. Centralbl. 61B (3): 569, 1942 (Herzog 1942)

Original citation: Costa Rica, Vara Blanca, between Poás and Barba-volcanoes, 1600–1700 m, tree trunk, leg. W. R. Maxon, 22. VII. 28 (no. 8277).

Lectotype (Swails 1970: 229 as ‘type’ hb Herzog) or possible holotype: Costa Rica, Vara Blanca, 1600–1700 m, Maxon & Harvey 8277 (JE04001321 https://je.jacq.org/JE04001321).

Currently accepted name: Porella leiboldii (Lehm.) Trevis. (cf. So 2005).

Madotheca subciliata Lehm. et Lindenb. var. endiviifolia Herzog, Rev. Bryol. Lichénol. 20: 127, 1951 (Herzog 1951)

Original citation: Costa Rica, Prov. San José, vicinity of Santa Mario de Dota, on log, [Standley] n. 42.374.

Lectotype (Swails 1970: 212 as ‘type’): Costa Rica, San José: nr Santa Maria de Dota, 1500–1800 m, 14–26 Dec. 1925, Standley 42374, Hb Herzog (JE04001320 https://je.jacq.org/JE04001320; isolectotype US00072952 http://n2t.net/ark:/65665/3f47db749-f76c-40e4-907c-67c176bf6ec8).

Currently accepted name: Porella crispata (Hook.) Trevis. (cf. Swails 1970).

Madotheca swartziana (F.Weber) Lindenb. var. decurrenti-spinulosa Herzog, Rev. Bryol. Lichénol. 20(1/2): 128, 1951 (Herzog 1951)

Original citation: Costa Rica, Prov. Cartago, La Estrella, on tree, [Standley] n. 39.553; Prov. Alajuela, Viento Fresco, on tree, [Standley] n. 47.921; Prov. Heredia, Cerros de Zurqui, on tree, 2000–2400 m., [Standley] n. 50.312.

Lectotype (Swails 1970: 229 as ‘type’): Costa Rica, Prov. Cartago, La Estrella, on tree, Standley 39553 (JE04010855 https://je.jacq.org/JE04010855; synlectotype Standley 50312 JE04010856 https://je.jacq.org/JE04010856).

Note. US00072948 ( http://n2t.net/ark:/65665/3f4968966-8fb1-4880-916e-5f04789947a4) is possibly an isolectotype (no image of label) although collector number differs a bit (error in registration?).

Currently accepted name: Porella leiboldii (Lehm.) Trevis. (cf. So 2005).

Madotheca swartziana (F.Weber) Lehm. var. guatemalensis Herzog, Beih. Bot. Centralbl. 61B(3): 570, 1942 (Herzog 1942)

Original citation: Guatemala, District of Petén, Fallabón–Yaxha Road, leg. C. L. Lundell, no 2066.

Lectotype (Swails 1970: 218 as ‘type’ in hb. Herzog), if not holotype: Guatemala, Petén: Fallabon–Yaxha Road, 23 Mar. 1933, Lundell 2052, Hb Herzog (JE04001294 https://je.jacq.org/JE04001294).

Note. Collection number in protologue is 2066 but only no. 2052 exists in JE and the number in the protologue is probably an error.

Currently accepted name: Porella swartziana (F.Weber) Trevis. (cf. Swails 1970).

Marchesinia longirostris Herzog, Rev. Bryol. Lichénol. 20: 138, 1951 (Herzog 1951)

Original citation: Costa-Rica, on tree, vicinity of Tilarán, Prov. Guanacaste, 500–600 m., leg. P. C. Standley, n. 44.309.

Lectotype (Gradstein 1994: 106 as ‘holotype’): Costa Rica, Guanacaste,Tilarán,Standley44309(JE04001750 https://je.jacq.org/JE04001750; isolectotypes B300025480 https://herbarium.bgbm.org/object/B300025480, US02482525 http://n2t.net/ark:/65665/3cda41101-edf0-4967-b917-5202c208db43).

Currently accepted name: Marchesinia bongardiana (Lehm. et Lindenb.) Schiffn. (cf. Gradstein 2021).

Marchesinia longirostris Herzog var. rivularis Herzog, Rev. Bryol. Lichénol. 20: 138, 1951 (Herzog 1951)

Original citation: Costa-Rica, on rock, along Rio Reventado, n. of Cartago, 1400–1600 m., leg. P. C. Standley, n. 49.561.

Lectotype (Gradstein 1994: 106 as ‘holotype’): Costa Rica, Cartago, Rio Reventado, 1460–1650 m, 26 Feb. 1926, Standley 49561 (JE04001751 https://je.jacq.org/JE04001751; isolectotypes B300025481 https://herbarium.bgbm.org/object/B300025481, US00070855 http://n2t.net/ark:/65665/3b03522e2-efa7-4473-bccc-7c4c122ebd95).

Currently accepted name: Marchesinia bongardiana (Lehm. et Lindenb.) Schiffn. (cf. Gradstein 2021).

Marchesinia minor Herzog, Rev. Bryol. Lichénol. 20: 137, 1951 (Herzog 1951)

Original citation: Costa Rica, on tree, vicinity of Tilarán, Prov. Guanacaste, 500–600 m., leg. P. C. Standley, n. 44.393.

Lectotype (Gradstein 1994: 106 as ‘holotype’): Costa Rica, Guanacaste,Tilarán,Standley44393(JE04001752 https://je.jacq.org/JE04001752; isolectotype US02482550 http://n2t.net/ark:/65665/39f497ced-0544-45b2-a053-1a360d8947aa).

Currently accepted name: Marchesinia brachiata (Sw.) Schiffn. (cf. Gradstein 1994).

Marchesinia trollii Herzog, Ann. Bryol.: 137, 1932 (Verdoorn 1932)

Original citation: Panama, Cerro Canajagua, Halbinsel Azuero, ea 950 m, leg. C. Troll, IV, 1929 [Hepaticae Selectae et Criticae 156].

Lectotype (Gradstein 1994: 106 as ‘holotype’): Panama, Peninsula Azuero, Cerro Canajagua, Troll s.n., Verdoorn (ed.), Hep. Sel. Crit. 156 (JE04001746 https://je.jacq.org/JE04001746; isolectotypes JE04001747 https://je.jacq.org/JE04001747, JE04001748 https://je.jacq.org/JE04001748, B, BR, FH, G, M, NY, S, U, US, YU).

Currently accepted name: Marchesinia brachiata (Sw.) Schiffn. var. brachiata (cf. Gradstein 1994).

Mastigobryum asperistipulum Steph., Sp. Hepat. (Stephani) 6: 453, 1924 (Stephani 1924)

Original type citation: Hab. Guatemala, Alta Verapaz (Türkheim legit.).

Lectotype (Fulford 1963: 156 as ‘type’): Guatemala, Alta Verapaz, Cobán, Tüerckheim s.n., Hb Levier 5817, Stephani herb. n. 22873 (G00067589 www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=123767&lang=en, isolectotypes JE04001013 https://je.jacq.org/JE04001013, NY253479 https://sweetgum.nybg.org/science/vh/specimen-details/?irn=610740, NY02697201 https://sweetgum.nybg.org/science/vh/specimen-details/?irn=2985283).

Currently accepted name: Bazzania hookeri (Lindenb.) Trevis. (cf. Fulford 1963).

Mastigolejeunea dentilobula Herzog, Rev. Bryol. Lichénol. 20(1/2):134, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley near Tela, 20–600 m. Costa-Rica: loco incerto, leg. P. C. Standley, n. 48.775p.

Lectotype (Gradstein 1994: 121): Honduras, Atlántida, Lancetilla valley, near Tela, Standley 48775a (JE04001662 https://je.jacq.org/JE04001662; synlectotype (Costa Rica) NY01059497 http://sweetgum.nybg.org/science/vh/specimen-details/?irn=1197081).

Currently accepted name: Caudalejeunea lehmanniana (Gottsche) A. Evans (cf. Gradstein (1994).

Mastigolejeunea innovans (Spruce) Steph. var. parciflora Herzog, Rev. Bryol. Lichénol. 20: 132, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Telas, Dep. of Atlántida, 20–600 m, [Standley] n. 55.342.

Lectotype (Gradstein 1994: 77 as ‘holotype’): Honduras, Atlántida, Lancetilla Valley, near Tela, Standley 55.342p. (JE04001656 https://je.jacq.org/JE04001656).

Note. The specimen in Jena is marked as ‘var. latistipula’ but it is almost certain that it was the base for Herzog’s name. He may have changed the epithet before publishing it without indicating it on the specimen. A specimen with the same collection number is present in US (US04660319, http://n2t.net/ark:/65665/3fbbe7d45-41fc-4212-a5a9-174150287093) originally identified as ‘Symbiezidium setosum?’ and re-identified as Archilejeunea parviflora by S. R. Gradstein and published as such by Gradstein (1994).

Currently accepted name: Thysananthus auriculatus (Wilson et Hook.) Sukkharak et Gradst. (cf. Gradstein 1994).

Mnioloma rhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930 (Herzog 1930)

Original citation: Costa Rica, leg. P. C. Standley n. 39.843 (on tree, vicinity of Orosi, Province of Cartago, 30 Mar. 1924).

Lectotype (here designated): Costa Rica, Vicinity of Orosí, Province of Cartago, 30 Mar. 1924, Standley 39843 (JE04001641 https://je.jacq.org/JE04001641, isolectotype JE04001642 https://je.jacq.org/JE04001642).

Note: The selected lectotype is in Herzog's handwriting and the isolectotype is probably a duplicate. There may also be isolectotypes in B and G.

Notoscyphus paramicola Herzog, Rev. Bryol. Lichénol. 11: 6, 1939 (Herzog 1938)

Original citation: Costarica, Cerro de las Vueltas, 2700–3000 m., on tree (n. 43.723/a).

Lectotype (Grolle 1971: 71 as ‘holotype’): Kostarika, Prov. S. José, C. de Las Vueltas, 2700–300 m, 1925, Standley 43723 (JE04000935 https://je.jacq.org/JE04000935 annotated as ‘holotype’ by R. Grolle in 1969, isolectotype JE04000934 https://je.jacq.org/JE04000934).

Currently accepted name: Syzygiella autumnalis (DC.) K. Feldberg, Váňa, Hentschel et Heinrichs (cf. Grolle 1971 as Jamesoniella autumnalis (DC.) Steph.).

Note. The lectotype may have been extracted from a larger gathering (see under Lophocolea tricuspidata above).

Oryzolejeunea grolleana Bernecker, Haussknechtia, Beih. 9: 37, 1999 (Bernecker-Lücking 1999)

Holotype: Costa Rica, Limón Province, Botarrama-Trail in the Braulio Carillo National Park, on leaves of Rhodospatha wendlandii Schott. ex Engl. (Araceae), Bernecker 92-147 (JE04001647 https://je.jacq.org/JE04001647).

Currently accepted name: Lejeunea grolleana (Bernecker) R. L. Zhu et W. Ye (cf. Ye et al. 2013).

Otigoniolejeunea quinqueumbonata (Spruce) Steph. var. rotundata Herzog, Rev. Bryol. Lichénol. 20(1/2): 154, 1951 (Herzog 1951)

Original citation: Honduras, Lancetilla Valley, near Tela, 20–600 m., on rotten log, [Standley] n. 55.379.

Lectotype (here designated): Honduras, Lancetilla Valley, near Tela, Departement of Atlántida; altitude 20–600 m, 6 Dec. 1927–20 Mar. 1928, Standley 55379 (JE04006773 ( https://je.jacq.org/JE04006773, isolectotype JE04001622 https://je.jacq.org/JE04001622).

Note. The selected type was annotated as ‘holotype’ by M. Reiner-Drehwald in 2007. The isolectotype is a microslide from R. Grolle’s herbarium that almost certainly was extracted from the specimen in T. Herzog’s herbarium. It is not known if there exists any duplicate material in US.

Currently accepted name: Lejeunea quinqueumbonata Spruce var. rotundata (Herzog) Sushil K. Singh.

Physocolea elliptica Steph., Sp. Hepat. (Stephani) 5: 876, 1916 (Stephani 1916).

Original citation: Hab. Guatemala.

Lectotype (here designated): Guatemala, Alta Verapaz, Cobán, 1600 m, Türckheim 5826 (G00069375, www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=115764&lang=en; isolectotype JE04002248 https://je.jacq.org/JE04002248 a microslide annotated ‘dupl. ex FI zwischen Diplasiol. brunnea’).

Note. The lectotype was annotated as ‘lectotypus’ by R. Grolle in 1976 and M. Reiner-Drehwald in 1994. It was mentioned as selected by R. Grolle in 1974 by Pócs et al. (2014), but this is probably only referring to Grolle’s annotation in G, and Pócs et al. (2014) did not say that they make a new lectotype and thus not validly selected one. A formal lectotypification seems not to have been made yet.

Currently accepted name: Cololejeunea camillii (Lehm. et Lindenb.) A. Evans (cf. Pócs et al. 2014).

Physocolea semidentata Steph., Sp. Hepat. (Stephani) 5: 883, 1916 (Stephani 1916)

Original citation: Hab. Costarica.

Lectotype (Pócs et al. 2015): Costa Rica, Bois de la vallée de Tuis, bassin du Reventarzón, September 1893, A. Tonduz, herb Cardot. n. 15591 (G00069367 http://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=115606&lang=en; isolectotype JE04004047 https://je.jacq.org/JE04004047).

Currently accepted name: Cololejeunea obliqua (Nees et Mont.) Schiffn. (cf. Pócs et al. 2015).

Plagiochila bursata Lindenb. f. *depauperata*** Herzog ex Carl, Ann. Bryol. Suppl. 2: 67, 1931 (Carl 1931). nom. inval. (ICN Art. 38.1(a); no description)

Original citation: Costarica, Standley …; n. 51.505, 50.709, 50.299 (var. minor f. depauperata Hzg.).

Original material: Costa Rica, Cerro de las Lajas, north of San Isidro, Provincia de Heredia, altitude 2000–2400 m, 7 Mar. 1926, Standley & Valerio 51505 (JE s.n, US04627055 http://n2t.net/ark:/65665/30cc6abf3-82de-494b-939a-0aedc2256b8b); Costa Rica, Cerros de Zurqui, northeast of San Isidro, Provincia de Heredia, altitude 2000–2400 m, 3 Mar. 1926, Standley & Valerio 50709 (JE s.n., M, US04627056 http://n2t.net/ark:/65665/3a52bef74-e37c-4d48-b3ab-8d41e26f82f8); ibid., Standley & Valerio 50299 (JE s.n., US04627054 http://n2t.net/ark:/65665/34e92e297-2c48-40cc-8b42-a0b79772318e).